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Volume 106
No. 1 January 2002
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2001
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AMERICAN JOURNAL OF ARCHAEOLOGY
Volume 106
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38
HG CH PO RH
Key: HG = hanging garden, CH = channels, PO = garden terrace portico, RH = refuse heap.
Table 3. Summary of Ancient Mollusk Remains Showing Relative Number of Individuals (RNI)
Fig. 12. Helix aspersa found in the channels (regio III,
context 186)
Fig. 11. Cepaea nemoralis found in the hanging garden area
(regio III, context 104)
27
Pliny HN 9.130; see also Andr 1961, 106.
28
Pers. comm., D.S. Reese, September 2000.
EZEQUIEL M. PINTO-GUILLAUME 46 [AJA 106
today around the Mediterranean, at most archaeo-
logical sites the specimens of this shell appear to
have been collected dead from beaches.
29
Apart
from the archaeological evidence, ancient sourc-
es mention an occasion where G. glycymeris was
served as a first course at a banquet.
30
The name
Glycymeris means, in fact, sweet-sour.
31
The gar-
den excavations of the Villa of Livia have yielded
five valves, of which one is water-worn. It is proba-
ble that this last mentioned specimen was not used
as food but rather was already dead when collect-
ed. Comparative examples of valves found at
Pompeii, Boscoreale, and Herculaneum are not
considered to be food remains.
32
A chronological
comparanda from the Imperial period is found at
Settefinestre (table 2). There are also 10 valves
from the first century B.C. to the first century A.D.
at Paestum and one valve from the first century
B.C. from Albintimilium.
33
Other specimens were
encountered at the Villa dei Quintilli, Monteporiz-
io (table 2).
A very attractive bivalve found is the Thorny or
Spiny oyster, Spondylus gaederopus (Linn 1758),
family Spondylidae (fig. 15). It is distributed
throughout Eurasia. While its shell resembles an
oyster with thorns or spines, it is in fact relat-
ed to scallops. In the same manner as oysters, it
attaches its lower valve to the substrate. S. gaedero-
pus is edible; it was a significant food item on the
East Palatine hill in Rome from the first to the
fifth centuries A.D.
34
Even though S. gaederopus is
common at some Mediterranean archaeological
sites,
35
it is interesting to note that this species is
not easy to obtain. It usually lives hidden beneath
and behind rocks, in crevices.
36
This is an inter-
esting aspect to which I shall return. Relevant
comparative examples are valves from Pompeii,
Herculaneum, the Schola Praeconum, and Set-
tefinestre (table 2). Other specimens were found
at Carthage.
37
Another bivalve encountered is the Common
or flat oyster, Ostrea (= Peloris) edulis (Linn 1758),
family Ostreidae (fig. 15). The O. edulis lives in
warm and temperate seas. This species is found
from the low water mark down to about 80 m on a
firm substrate, to which the oyster attaches its low-
er valve. The Common or flat oyster has a thick
irregular shell with a scale-like surface. It usually
reaches between 70 and 100 mm long. It is com-
monly eaten throughout the Mediterranean; in
addition, ancient sources describe their use as
Fig. 13. Unio elongatus found in the hanging garden area
(regio III, context 184)
Fig. 14. Murex brandaris (left) and Glycymeris glycymeris (right),
found in the channels (regio III, context 186)
Fig. 15. Ostrea edulis (above left), Spondylus gaederopus (above
right), and Pisania striata (below), found in the channels
(regio III, context 185)
29
Reese (forthcoming b), 3; (forthcoming c), 28.
30
Macrinus 3.13.12.
31
Pers. comm., A. Warn, September 1998.
32
Reese (forthcoming c), 267.
33
Paestum: Reese (forthcoming d), 335. Albintimilium:
Lamboglia 1950, 87, fig. 39, no. 105.
34
Reese (forthcoming c), 37.
35
Pers. comm., D.S. Reese, November 1998.
36
Pers. comm., A. Warn, September 1998.
37
Reese 1982, 93.
MOLLUSKS FROM THE VILLA OF LIVIA AT PRIMA PORTA 47 2002]
food and in medicine.
38
Evidence suggests that
the Romans began cultivating oysters by the first
century B.C.
39
Moreover, oyster farms are well
known from depictions on glass vessels.
40
At the
Villa of Livia a very peculiar O. edulis was found
attached to an amphora sherd. In spite of its sin-
gularity, this oyster was possibly brought to the vil-
la together with other bivalves as a food source.
41
This bivalve is common at classical farmsteads in
Molise and at Carthage.
42
There are also valves from
Pompeii, Herculaneum, the Schola Praeconum,
Settefinestre, and the Late Roman villa at S. Gio-
vanni di Ruoti (Potenza) (table 2). Other speci-
mens were found at Paestum and the Roman Villa
of San Potito-Ovindoli (LAquila).
43
Another bivalve yielded by the excavations is the
European, Edible, or Common cockle, Cerastoderma
edule (Linn 1758), family Cardiidae (fig. 16). It is
the most common bivalve in sandy shores and estu-
aries in Europe. It lives buried immediately below
the sand surface. The C. edule is eaten today both
raw and cooked throughout Europe. Comparative
examples particularly found in gardens are those
specimens from Pompeii (I.xiv.2).
44
A chronologi-
cal comparanda can be found in specimens from
the Republican period at Ficana and from the Au-
gustan cemetery (Phase I) at Otranto (table 2). It is
also found at Settefinestre and San Giovanni di
Ruoti (Potenza) (table 2). Other C. edule have been
collected at Knossos and Paestum.
45
In two places within the gardens fragments of
Wedge-shells were encountered. The Wedge-shell,
Donax (=Cuneus) trunculus (Linn 1758), family
Donacidae (fig. 17), is usually found living on the
shore, buried 1015 cm deep in the sand. The shell
has a triangular or elongated shape, posteriorly
shorter than anteriorly. They seem to have been a
common food item during antiquity at Morgantina
in Sicily and on the East Palatine of Rome,
46
and
they are eaten today around the Mediterranean.
47
Comparative examples from gardens include three
specimens from Pompeii (one valve in I.xiv.2; one in
the Garden of the Fugitives, I.xxi.2, and one valve in
the triclinium in the small north garden at I.xxi.3).
48
There are also similar specimens from Oplontis, the
Villa dei Quintilli at Monteporizio (table 2). Other
examples were found at Paestum and Jerusalem.
49
The Warty venus, Venus verrucosa (Linn 1758),
family Veneridae (fig. 18) is another bivalve fre-
Fig. 16. Cerastoderma edule found in the garden portico area
(regio III, context 427)
Fig. 17. Donax trunculus found in the hanging garden area
(regio III, context 104)
Fig. 18. Venus verrucosa found in the garden portico area
(regio III, context 427)
38
Pliny HN 32.5964.
39
Pliny HN 9.168; Columella Rust. 8.16.58; Verg. G. 1.204;
see also Andrews 1948; Andr 1961, 108; Eyton 1858.
40
Gnther 1897.
41
Pinto-Guillaume (forthcoming).
42
Molise and Carthage: Reese 1982, 93.
43
Paestum: Reese (forthcoming d), 335. San Potito-Ovin-
doli: Bknyi 1986, 90, table 1.
44
Jashemski 1979, 96 (as Cardium edule); see also Reese
(forthcoming c), 22.
45
Knossos: Reese 1992a, 4946. Paestum: Reese (forthcom-
ing d), 26.
46
Reese (forthcoming c), 25.
47
Personal observation, Naples, October 2000.
48
Jashemski 1979, 96, 247; see also Reese (forthcoming c),
25.
49
Paestum: Reese (forthcoming d), 334. Jerusalem: Reese
1995, 2738.
EZEQUIEL M. PINTO-GUILLAUME 48 [AJA 106
quently found throughout the Mediterranean Sea.
The shell is strong and has a round triangular
shape with irregular radial ribs. The V. verrucosa is
an edible species. Comparative examples are three
valves in fountains at Pompeii and one valve dating
to the Neronian period from Knossos.
50
distribution and occurrence of
mollusks in the gardens
The small garden is a peristyle garden included
within the private quarters of the villa complex it-
self (fig. 1).
51
Twenty-six land snails were found in
this area. Two small deposits of H. aperta, which
proved to be modern, were unearthed at a depth of
ca. 15 cm. It is difficult to establish if this particular
species could have been present in this area dur-
ing the Roman period because of the fact that the
H. aperta presents a very soft shell with a low calci-
um content that decomposes and does not preserve
well. The other land snails yielded by this area are
a few specimens of R. decollata and P. elegans. Beside
this, the small garden produced only one marine
mollusk: a broken valve of G. glycymeris (table 3).
Regio I consisted of a set of trenches opened in
the garden terrace. These trenches extended from
the northeastern limit of the villa toward the center
of the garden terrace (fig. 1). Three land snails of
the H. aspersa and the C. nemoralis species were en-
countered in this area, and a fair concentration of
Oxychilus sp. also was found. Regarding the marine
shells produced in regio I, there were two M. bran-
daris, one fairly large water-worn valve of G. glycymer-
is, and an O. edulis fragment (table 3).
Regio II, located in the very center of the garden
terrace (fig. 1), yielded only three land snails. I
have noticed modern living C. nemoralis (July 1999)
and H. aperta (May 1997 and June 1998) wander-
ing in the central area of the garden terrace under
the sun, probably completing their daily reconnais-
sance.
52
The low occurrence of both land snail and
marine shell remains corresponds with the scarcity
of archaeological material in this area (table 3).
53
Located in the far northeastern corner of the
garden terrace is regio III (fig. 1). Here we found
the largest concentrations of ancient shell remains
with 1,674 land snails and 32 marine shells. Based
on the preliminary archaeological results, four spe-
cialized areas with differentiated characteristics are
defined within regio III (table 1). First, located in
the excavated northeastern section of regio III is
the hanging garden area, a series of apses and rect-
angular planting cassettes that were distributed in
a terraced fashion and provided with drainage fea-
tures (fig. 2).
54
Among the 192 land snails retrieved
in this area, Oxychilus sp., C. planospira, and P. elegans
predominate. The high concentrations of P. elegans
in the hanging garden and in the garden portico
(fig. 19) seem to indicate that these two areas not
only are highly carbonated, but also present a cer-
tain degree of disturbance in the soil. This evidence
could be indicative of clearance, for example. This
particular area, which has been identified as a plant-
ing terrace,
55
yielded the largest variation of ma-
rine shells: five M. brandaris, one O. edulis, one S.
gaederopus, one D. trunculus, one G. glycymeris, and
one freshwater bivalve, one U. elongatus (table 3).
Second, the channels, which are located imme-
diately to the southwest and below the above de-
scribed area, consist of a set of parallel channels
that apparently run along the entire length of the
garden terrace (fig. 2). Within the channels a total
of 1,060 land snails were encountered, of which
642 were C. planospira. The distribution of this spe-
cies throughout regio III shows that its concentra-
tion is almost exclusive to the rear channel (fig.
20). This high density of C. planospira is about four
times higher than in the rest of regio III. The great
quantities of ceramic material found within the rear
channel have shown that this channel was filled at
the same time and not gradually.
56
It is possible to
imply that the ceramic material was brought to the
channel from a kitchen refuse heap. If this theory
is correct, two scenarios are possible. The first hy-
pothesis is that the C. planospira lived in a pottery
dump, feeding off the organic waste from the kitch-
en, while the second hypothesis is that C. planospira
could have been eaten by the villas inhabitants.
Even though there is neither literary nor archaeo-
logical evidence for C. planospira being consumed
in ancient times, the question raised is an interest-
ing one. Did the Romans make an exception? It is
noteworthy that this species is not eaten today. In
addition to this species, a high number of Oxychilus
sp. was also found in the channels. Also present
50
Pompeii: Reese (forthcoming c) 389. Knossos: Reese
1992a, 4946.
51
Calci and Messineo 1984, 48; see also Messineo 1991, 230.
52
Pers. comm., A. Warn, October 1999.
53
Pinto-Guillaume 1998, 189; see also Liljenstolpe and
Klynne 19971998, 137; and Klynne and Liljenstolpe 2000,
226.
54
Klynne and Liljenstolpe 2000, 227 and figs. 89.
55
Khler 1959, 10; see also Klynne and Liljenstolpe 2000,
22731.
56
Klynne and Liljenstolpe 2000, 230.
MOLLUSKS FROM THE VILLA OF LIVIA AT PRIMA PORTA 49 2002]
were 11 marine shells of the following species: M.
brandaris, G. glycymeris, D. trunculus, O. edulis, S. gaed-
eropus, and P. striata (table 3).
Third, descending toward the center of the gar-
den terrace, the next distinguishable area was oc-
cupied by a garden portico, which ran along all four
sides of the garden terrace (fig. 2). As mentioned
above, the P. elegans is one of the most frequent land
snails in this area. The garden portico area pro-
duced a total of 356 land snails. A high concentra-
tion of R. decollata was observed (fig. 21), as was a
low number of H. aspersa individuals. This possible
scenario has clear parallels in gardens of today: the
decollate snail feeding on young brown garden
snails. It is difficult to prove if R. decollata could
have been introduced by the Romans with the par-
ticular purpose of fighting back the mollusk spe-
cies that caused damage to the garden. We do know,
however, that the Romans possessed the knowledge
of how to control those animals that were damaging
to the household and the gardens.
57
The Romans
used ritual and magic practices as well as mechan-
ical, chemical, and biological methods to control
pests. The practice of fighting back pests by means
of using other life forms was not unknown to the
Romans.
58
Moreover, Pliny suggests how certain veg-
etables could be protected from damaging insects
by sowing other plant species among the vegeta-
bles.
59
Finally, in the same area, regarding the ma-
rine species, there are one S. gaederopus, one O. edu-
lis, one V. verrucosa, and one C. edule (table 3).
The last of these specialized areas in regio III is
the refuse heap, located immediately extra muros to
the east of the garden terrace (fig. 2). Here 66 land
snails were found, of which C. nemoralis and C. plano-
spira are the dominant land snails. Five marine shells
retrieved from this area belong to the M. brandaris
and S. gaederopus species. The shell remains encoun-
tered in the refuse heap are fewer in abundance
when compared to the channels or to the rest of
regio III. In addition to the scarcity of remains here,
the shells in this particular area also were very frag-
mented (table 3).
Only three land snails were found in regio V,
which is a long strip running north to south across
the garden terrace (fig. 1). The scarcity of individ-
uals seems to parallel the situation in regio II dis-
cussed above, where very few remains of ancient
land snails and no marine shells have been found
(table 3).
Fig. 19. Plan of regio III showing the distribution of the land snail Pomatias elegans. Each dot
represents one snail.
57
Varro Rust. 3.9.14-15 (lice, snakes, and hawks); Cato Agr.
Orig. 92, 95, 98 (weevils, caterpillars, and moths); Columella
De Arboribus, 145 (ants and mice); Verg. G. 1.17886 (weeds,
rodents, and other pests); 4.24250 (snakes), and Pliny HN
17.21863 (several different pests) are all important examples.
58
Ordish 1976, 34.
59
Pliny HN 19.1789.
EZEQUIEL M. PINTO-GUILLAUME 50 [AJA 106
Finally, regio VI consisted of two trenches, which
were opened on the eastern limit of the garden
terrace among a grove of poplar trees (fig. 1). This
area produced exclusively land snails, with a total
of 166 individuals. The most frequent species en-
countered here was Oxychilus sp. (table 3).
reconstructing ecological niches
Considering the complex activity of snails, it has
been difficult for scientists to define a specific niche
to a particular species.
60
In spite of this difficulty,
some observations can be made based on inherent
characteristics of the various species, such as habitat
preferences (shaded areas, open country, woodland,
etc.) and preferred levels of humidity or dryness.
The actual site of the Villa of Livia is located about
50 m above sea level on a hill. The immediate sur-
rounding area to the south and to the east is the
Tiber River valley. The geomorphology of this area
is characterized by a number of low hills, which are
RNI/Species Present
Small garden
3 Pomatias elegans
1 Rumina decollata
Regio I
52 Oxychilus sp.
2 Cepaea nemoralis
1 Helix aspersa
Regio III: hanging garden
60 Oxychilus sp.
49 Campylaea planospira
40 Pomatias elegans
22 Cepaea nemoralis
6 Helix aspersa
6 Retinella pura
6 Rumina decollata
2 Cochlicella barbara
Regio III: channels
642 Campylaea planospira
233 Oxychilus sp.
76 Cepaea nemoralis
60 Helix aspersa
28 Cernuella virgata
11 Retinella pura
4 Chondrula tridens
3 Rumina decollata
3 Pomatias elegans
Regio III: garden portico
103 Oxychilus sp.
82 Rumina decollata
65 Pomatias elegans
62 Cepaea nemoralis
27 Campylaea planospira
9 Retinella pura
7 Helix aspersa
1 Chondrula tridens
Regio III: refuse heap
29 Campylaea planospira
27 Cepaea nemoralis
4 Cernuella virgata
3 Pomatias elegans
2 Rumina decollata
1 Helix aspersa
Partial shade, reduced woodland
Open areas/shrub, red. woodland
Open areas, dry walls, gardens
Indifferent, bushes, gardens
Shaded areas, shrub, red. woodland
Open areas, dry walls, gardens
Slopes, dry walls and gardens
Partial shade, reduced woodland
Indifferent, bushes, gardens
Shaded areas, shrub, red. woodland
Shaded areas near rocks and walls
Open areas/shrub, red. woodland
Slopes, dry walls and gardens
Slopes, dry walls and gardens
Open areas, dry walls, gardens
Indifferent, bushes, gardens
Shaded areas, shrub, red. woodland
Open areas, shrub, red. woodland
Shaded areas near rocks and walls
Open areas, under leaves, bushes
Open areas/shrub, red. woodland
Partial shade, reduced woodland
Open areas, dry walls, gardens
Open areas/shrub, red. woodland
Partial shade, reduced woodland
Indifferent, bushes, gardens
Slopes, dry walls and gardens
Shaded areas near rocks and walls
Shaded areas, shrub, red. woodland
Open areas, under leaves, bushes
Slopes, dry walls and gardens
Indifferent, bushes, gardens
Open areas, shrub, red. woodland
Partial shade, reduced woodland
Open areas/shrub, red. woodland
Shaded areas, shrub, red. woodland
Preferred Environs/Vegetation
Br, High CaCo
Ni, CaCo
Or, CaCo
High CaCo
High CaCo
Or, CaCo
CaCo
Br, High CaCo
High CaCo
High CaCo
CaCo
Ni, CaCo
CaCo
CaCo
Or, CaCo
High CaCo
High CaCo
CaCo
Ni, CaCo
Br, High CaCo
Or, CaCo
Ni, CaCo
Br, High CaCo
High CaCo
CaCo
CaCo
High CaCo
CaCo
High CaCo