Escolar Documentos
Profissional Documentos
Cultura Documentos
Dipartimento di Scienze Botaniche, Universit di Pisa, Via Luca Ghini 5, I-56126 Pisa, Italy
b School of Pharmacy, University of Bradford, Bradford, West Yorkshire, BD7 1DP, UK
c Museo Nazionale Arti e Tradizioni Popolari, Piazza Marconi 8-10, 00144 Rome, Italy
Department of Social Studies, Wageningen University, Gebouuwnummer 201, Hollandseweg 1, NL-6706 KN Wageningen, The Netherlands
Received 5 May 2003; received in revised form 19 June 2003; accepted 8 August 2003
Abstract
We report folk veterinary phytotherapy in Italy collected from ethnobotanical scientific literature of the second half of the 20th Century.
References are cited together with unpublished data gathered recently in the field by the authors. The data have been placed in two databases:
one organized by the names of the plant species (>260) and the other organized by bibliographic references. This represents the basis for the
first national databank for ethnoveterinary botany in Europe. Plants not yet sufficiently studied in pharmacology and veterinary phytotherapy
were also identified.
2003 Elsevier Ireland Ltd. All rights reserved.
Keywords: Ethnoveterinary; Ethnobotany; Medicinal plants; Italy
1. Introduction
The recent return to natural medicine in Italy has emphasized the importance of gathering information about
medicinal plants traditionally used to treat animals. As in
other branches of folk botany, reference to remedies for
animals are often only side comments (Mathias-Mundy and
McCorkle, 1989). Specific references from outside Italy include Mathias-Mundy and McCorkle (1989), McCorkle and
Mathias-Mundy (1992), Goud and Pullaiah (1996), Lans
and Brown (1998a,b), Pieroni (1999a), Lans et al. (2000),
Lopez and Aguilar (2001), and Villareal Quintanilla (2001).
In Italy, many plants were used to treat cows, sheep, poultry, horses and pigs, and these traditions have survived in
some areas. Many of these remedies were handed down by
word of mouth (Guarrera, 1994; Viegi et al., 1999; Viegi
and Pieroni, 2000a,b; Pieroni et al., 2002a,b).
The aim of this paper is to gather information on the
present status of folk veterinary knowledge in Italy. The information on plant remedies for domestic animals and other
effects of plants on various animals, gathered here, was
mostly brief notes in folk botanical literature concerned prin Corresponding
cipally with human therapy. Folk veterinary information collected in various regions of Italy is reported and discussed.
2. Materials and methods
All references available in Italian folk botanical literature
(Caffaro Corti and Gastaldo, 1980) since 1950 up to now
were examined for reports on remedies for animals. We also
cite some earlier publications, the oldest of which, on Helleborus, goes back to 1878 (Padula).
The data, extrapolated by rigorous botanical studies (to
ensure correct determination of plants), was placed in two
databases, one for the taxa used in folk veterinary medicine
and the other for the respective references, so as to create the
basis for a databank. Plant nomenclature (scientific names) is
largely according to Pignatti (1982), Tutin et al. (19641980,
1993), and Greuter et al. (2000).
We then handled and elaborated the data region by region. We examined and added preliminary unpublished
data obtained in the field by interviews in Tuscany, Liguria,
Campania, Calabria, Apulia and Lucania (De Fine, Guazzi,
Maccioni, Marchini, Menale, Muoio, Pieroni and Rizzo,
personal communication). The voucher herbarium specimens are available from the authors and in PI (Herbarium
of the Department of Botanical Science, University of Pisa).
0378-8741/$ see front matter 2003 Elsevier Ireland Ltd. All rights reserved.
doi:10.1016/j.jep.2003.08.003
222
223
Table 1
Plant species cited for ethnoveterinary use in Italy
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Abies sp.
Achillea ligustica All.
Achillea millefolium L.
Aconitum sp.
Adiantum capillus-veneris L.
Aesculus hippocastanum L.
Pinaceae
Asteraceae
Asteraceae
Ranunculaceae
Adiantaceae
Hippocastanaceae
Agave americana L.
Agrimonia eupatoria L.
Agavaceae
Rosaceae
GI
AP
SK
AP
RP
GI
RE
SK
SK
sh
ct
ho
ho
C
C
C
C
C
C
C
C
C
Poaceae
Alchemilla sp.
Allium cepa L.
Rosaceae
Liliaceae
RP
SK
GI, SK
OT
RP
GI
as
do, ho
ct, sh
ch, sh
C
O
C, O
C
O
C
RP, GI, AP
AP
OT
SK, LA
GI, AP
LA
AP
ch, tu
pi
pi
ch, ct
sh
O, C
C
C
C
C
C
C
SK
NE, KI,
RE, AP, SK
GI
C
C
ch
SK
LA
AP
RE
SO
SK
as, ho
ch
ct
ct
ct
C
C
C
C
C
C
NE
AP
SK
SO
KI, SK
FS
GI
AP
GI
GI, AP
AL
OT
OT
RP, AP
GI
GI
OT, GI
OT
AP
ct
ct
pi, sh
ct
sh
sh
ct
ct
ct
ch, ct
C
O
C
C
P, C
O
C
P, C
C
C
C
C
C
P, C
C
P, C
C
C, O
P, C
OT
GI
AP
FS
ch
C
C
C
P
Allium sativum L.
Liliaceae
Allium vineale L.
Allium sp.pl.
Alnus glutinosa (L.) Gartner
Liliaceae
Liliaceae
Betulaceae
Althaea officinalis L.
Malvaceae
Amaranthus sp.
Anagallis arvensis L.
Amaranthaceae
Scrophulariaceae
Angelica sylvestris L.
Apiaceae
Anthemis altissima L.
Anthemis cotula L.
Apium nodiflorum (L.) Lag.
Asteraceae
Asteraceae
Apiaceae
Arctium lappa L.
Arctium sp.pl.
Aristolochia rotunda L.
Artemisia absinthium L.
Asteraceae
Asteraceae
Aristolochiaceae
Asteraceae
Artemisia arborescens L.
Artemisia vulgaris L.
Asteraceae
Asteraceae
Araceae
224
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Asphodelus microcarpus
Salzm. et Viv.
Asplenium trichomanes L.
Atractylis gummifera L.
Avena sativa L.
Liliaceae
SK
ho
Aspleniaceae
Asteraceae
Poaceae
GI
AP
RP
SK
FS
sh
ct
sh
ct
C
C
O
C
P
GI
RE
SK
GI
LO
GI, SK
LO
SK
KI
SK
AP
GI
GI
GI
KI
LO
KI
ct
sh
ct, sh
ct, ho
ct, sh
ct
ct
ct
C
P
C
C
C
C
C
C
C
P
P, C
C
C
C
C
C
C
RP
OT
RP
OT
GI
FS
RP
RE
KI
GI, SO
RP
OT
GI
ch
ct, sh
pi
ct, sh
ct
ct
ho
O
C
P
P
C
O
P, C
C
C
C
C
C
C
ct, ho
tu
C
O
C
P
C
Apiaceae
Betulaceae
Boraginaceae
Brassicaceae
Brassica oleracea L.
Brassicaceae
Bryonia dioica L.
Buxus sempervirens L.
Calamintha nepeta (L.) Savi
Calendula arvensis L.
Calendula officinalis L.
Calendula sp.pl.
Calluna vulgaris (L.) Hull
Cannabis sativa L.
Capsella bursa-pastoris L.
Medik
Capsicum annum L.
Cucurbitaceae
Buxaceae
Lamiaceae
Asteraceae
Asteraceae
Asteraceae
Ericaceae
Cannabaceae
Brassicaceae
Carduus pycnocephalus L.
Castanea sativa Miller
Ceterach officinarum Lam.
et DC
Asteraceae
Fagaceae
Aspleniaceae
Asteraceae
Chelidonium majus L.
Papaveraceae
Chenopodium vulvaria L.
Chrozophora tinctoria (L.)
Juss.
Chrysanthemum parthenium
Bernh.
Cirsium arvense (L.) Scop.
Chenopodiaceae
Euphorbiaceae
RE, GI, RP
OT
SK
OT
SK
Asteraceae
NE
Bandini (1961)
Asteraceae
GI
SK
GI
ra
ct
P, C
C
C
Colchicum autunnale L.
Conium maculatum L.
Liliaceae
Apiaceae
Convolvulus arvensis L.
Convolvulaceae
Cornus sanguinea L.
Cornaceae
SO
OT
GI
RE
AP
SK
AP
RE
FS
GI
FS, OT
OT
sh
go
sh
ct, ho
ra
pi
ra
sh
C, M
C
C, M
C
C
C
C
C
P, C
P, C
P
C
Solanaceae
Ranunculaceae
225
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Crataegus laevigata L.
Crithmum maritimum L.
Cucurbita pepo L.
Cuminum cyminum L.
Cupressus sempervirens L.
Rosaceae
Apiaceae
Cucurbitaceae
Apiaceae
Cupressaceae
Asteraceae
ct
ra
ct
pi, ra
ct, ho
ct
C
P
C
C
C
Cynara cardunculus
ssp. scolymus (L.) Hayek
Cynodon dactylon Pers.
GI
OT
AP
GI
GI
OT
GI
FS
GI
GI, KI
SK
ho
sh
ct
P
C
C
C
Manzi (1989)
Lentini et al. (1988)
Catanzaro (1968)
De Bellis (1978, 1986, 1988)
ct
as
C
C
C
Poaceae
Cynoglossum creticum
Miller (syn. Cynoglossum
pictum Aiton)
Cynoglossum officinale L.
Cytisus scoparius L.
Boraginaceae
Boraginaceae
Fabaceae
Daphne gnidium L.
Thymelaeaceae
SK
OT
SK
OT
Daphne mezereum L.
Thymelaeaceae
AP
Datura stramonium L.
Solanaceae
AP
ch, sh
P, C
Cucurbitaceae
SK
ct
Echinochloa crus-galli
Beauv.
Echium vulgare L.
Poaceae
GI
FS
mu
bi
C
P
Equisetum arvense L.
Equisetum telmateja Ehrh.
Erica arborea L.
Equisetaceae
Equisetaceae
Ericaceae
OT
SK, AP, OT
GI
SK, OT
NE
do, sh
ct
ct
C, M
C
C
C
C
Eryobotrya japonica
(Thunb.) Lindley
Eucalyptus sp.pl.
Euonymus europaeus L.
Rosaceae
RE
ch
Tammaro (1976)
Viegi et al. (1999)
Ciccodicola (1995)
De Capite and Menghini (1973)
De Feo et al. (1992)
De Feo and Senatore (1993)
Maccioni and Marchini (1998b)
Myrtaceae
Celastraceae
SK
AP
C
C
Celastraceae
Asteraceae
Euphorbiaceae
Euphorbiaceae
Euphorbia dendroides L.
Euphorbiaceae
AP
SK
SK, OT
SK, OT
OT
OT
de
P, C
C
C
C, O
O
O
Euphorbia helioscopia L.
Euphorbiaceae
SK, OT
Euphorbia lathyris L.
Euphorbiaceae
Euphorbiaceae
Euphorbiaceae
Apiaceae
Moraceae
AP
GI
AP
SK
GI
RE
GI
ct
ho
ct, sh
P
C
P
C
C
C
C
Apiaceae
RE, OT
RP
ct, go, sh
ct
C
P
GI
FS
ct
ra
C
P
Boraginaceae
226
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Fraxinus excelsior L.
Oleaceae
GI
ch, sh
P, C
Fraxinus ornus L.
Oleaceae
OT
GI
ch, ra
C
P, C
SK
GI
GI
ch
ch
ch, ct
C
P
C
AP
GI
RP
GI
OT
GI
GI
ch
go, sh
ca, pi
ct
P, C
C
P
C
P
C
C
SK, GI
AP
AP
GI, RE
GI
OT
SK
ho
as, ho
ct, pi
go, sh
ct, mu
C
C
O
C
C
O
C
RP
SK, OT
RP
OT
RE
ct
ct
ch
as
P
C
O
O
C
ct
pi, sh
C
C
C
C
C
Fumaria officinalis L.
Galega officinalis L.
Galium verum L.
Gentiana acaulis L.
Fumariaceae
Fabaceae
Rubiaceae
Gentianaceae
Geraniaceae
Rosaceae
Lamiaceae
Fabaceae
Hedera helix L.
Hedera helix L.
Araliaceae
Araliaceae
Hedera sp.
Helianthus annuus L.
Helichrysum italicum G. Don
Araliaceae
Asteraceae
Asteraceae
Ranunculaceae
Ranunculaceae
GI
OT
NE
RE
RE
Ranunculaceae
KI
GI, OT
OT
C
C
SK
NE
KI
GI
SK
RP, OT
SK
NE, RE,
OT
RP
KI
RE, SK
RE
RP, FS
la
C
C
C
C
C
C
C
C
sh
ct, sh
ct
Helleborus niger L.
Helleborus odorus W. et K.
Ranunculaceae
Ranunculaceae
Helleborus viridis L.
Helleborus sp.
Heracleum sphondilium L.
Ranunculaceae
Apiaceae
C
C
P
227
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Hordeum vulgare L.
Poaceae
Humulus lupulus L.
Hyoscyamus niger L.
Hypericum perforatum L.
Cannabaceae
Solanaceae
Guttiferae
Hyssopus officinalis L.
Ilex aquifolium L.
Lamiaceae
Aquifoliaceae
Asteraceae
Inula sp.
Juglans regia L.
Asteraceae
Juglandaceae
GI
GI
GI
LA
SK, OT
SK
GI, RP
SK, LA
SK
SK
LA
RE, FS
AP
ct, ho
ct, do
ct, sh
ct, sh
ho
do, ho
P, C
C
C
C
C
C
C
C
C
C
C
C
P
do, ho, tu
ho
ra
ct, ho
ra
ra
sh
C
P
C, O
C
O
C
O
C
P, C
C
Ferri (1961)
De Capite and Menghini (1973)
Fossati et al. (1999)
Tammaro and Pietrocola (1975)
Viegi et al. (1999)
Lentini and Aleo (1991)
Guarrera (1987)
Cappelletti et al. (1981)
Renzetti and Taiani (1988)
Tammaro (1984), Viegi et al. (1999)
Viegi et al. (1999)
Picchi (1999)
DAndrea (1982), Guarrera et al. (1984),
Renzetti and Taiani (1988), Guarrera (1994)
Corrain and Zampini (1961)
De Capite and Menghini (1973)
Guarrera et al. (1984), Tammaro (1984)
DAndrea (1982)
Maccioni and Marchini (1999)
De Bellis (1988)
Viegi et al. (1999)
De Capite and Menghini (1973)
Maccioni and Marchini (1998a)
Mearelli and Tardelli (1995)
ct
sh
C
C
Manzi (1989)
Bruni et al. (1997)
Juniperus communis L.
Cupressaceae
Juniperus oxycedrus L.
Cupressaceae
Juniperus oxycedrus L.
subsp. oxycedrus
Juniperus oxycedrus L.
subsp. macrocarpa
(S. et S.) Ball
Juniperus sabina L.
Cupressaceae
GI, OT
AP, RP
OT
LA, SK
FS
LA
GI
KI, RP
FS
LA, AP,
SK
RP
LA, SK
Cupressaceae
GI, FS
ra
Cupressaceae
Juniperus sp.
Kichxia elatine (L.) Dumort.
Cupressaceae
Scrophulariaceae
RP
RP
LA
SK
ct, ho
as, do
C
C, O
C
C
Fabaceae
Pinaceae
Lauraceae
Lavatera arborea L.
Lavatera cretica L.
Malvaceae
Malvaceae
SK
GI
GI, RP
AP, OT
AP
GI
OT
OT
GI
sh
ct
ct
ct
C
C
C
P, C
C
C
C
C
C
Liliaceae
Linaceae
OT
GI
ct, pi
C
C
LA, SK
GI, SK
ct
ct
C
C
ct, pi
ct
as, ho
ct
do, ct, pi
P, C
C
C
C
ct
C
C
Lolium temulentum L.
Lotus corniculatus L.
Lupinus albus L.
Poaceae
Fabaceae
Fabaceae
GI, RP
SK
NE
GI
AP
Lupinus luteus L.
Lycoperdon sp.
Fabaceae
Lycoperdaceae
AP
SK
228
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Lycopersicon esculentum
Miller
Malva neglecta Wallr.
Malva parvifolia L.
Malva sylvestris L.
Fabaceae
AP
GI, RP, AP
RP
SK
RP, GI
GI
ct
ct
ct, pi
C
P, C
C
C
C
C
SK
ct
SK, GI
RE, GI
GI, RP,
RE, NE
GI, NE
GI
OT
SK
GI
OT
SK
ct
ct
C
C
C
Guarrera (1994)
Guarrera (1987)
Chimenti Signorini and Fumagalli (1983)
Atzei et al. (1991)
Renzetti and Taiani (1988)
Corrain (1977), Corsi et al. (1981),
Guarrera (1981), De Simoni and Guarrera
(1994), Nardelli (1987), Uncini Manganelli
and Tomei (1995)
Bellomaria (1982), De Bellis (1986),
Fossati et al. (1999), Guarrera (1987)
Corrain and Zampini (1961)
Guarrera (1987)
Manzi (1989)
C
C
C
C
C
O
C
Malvaceae
Malvaceae
Malvaceae
Malva sp.pl.
Malvaceae
Lamiaceae
Lamiaceae
Melilotus sp.pl.
Melissa officinalis L.
Melissa romana Miller
Mentha pulegium L.
Mentha sp.pl.
Mentha x piperita L.
Mercurialis annua L.
Fabaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Lamiaceae
Euphorbiaceae
RP
RP
OT
RE
GI
SK
GI
pi
ct, pi
ct
ct
ct
ct, ho
ct
Lamiaceae
RP, GI
GI, RP, AP
SK, LA
ct
C
C
C
SK
SK, GI
OT
SK
RP
AP, SK
AP
ct, ra
ct
do
C
C
C
C
P, C
C
C
AP
OT
GI, OT
SK, LA
OT
AP
SK
RP
GI
SK
RE
GI
SK, OT
GI
RP
NE
NE
ct
ct
pi
ct
ho
as
pi
ch
pi
as, ho
C
O
C
C
C
P
C
P, C
C
C
C
C
C
C, O
P
C
C
Moraceae
Myrtaceae
Brassicaceae
Nerium oleander L.
Apocynaceae
Nicotiana tabacum L.
Oenanthe crocata L.
Olea europaea L.
Solanaceae
Apiaceae
Oleaceae
Fabaceae
Cactaceae
Papaver somniferum L.
Papaveraceae
Lamiaceae
Poaceae
Papaveraceae
P
P
C
C
C
C
C
229
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Urticaceae
Urticaceae
Parietaria officinalis L.
Urticaceae
Parietaria sp.pl.
Petroselinum crispum
(Miller) A.W.Hill
Urticaceae
Apiaceae
Apiaceae
SK, RP
RE
RP
SK, RP
GI
SK, GI
KI
SK, RP
SK
GI
RP
GI, OT
OT
KI
GI
sh
ct
do
ct
do
as
ra
ch
ra
ra
C
C
C
C
C
P
C
C
C
P, C
O, C
P, C
O
C
C
Nardelli (1987)
Lentini et al. (1988)
Corsi et al. (1981)
Nardelli (1987)
Guarrera (1994)
Catanzaro (1968)
De Capite and Menghini (1973)
Nardelli (1987)
Manzi (1989)
Maccioni and Marchini (1998a, 1999)
Guarrera (1994, 1995)
Martini (1981, 1982)
Lomagno and Lomagno Caramiello (1970)
Corsi et al. (1981)
Coassini Lokar and Poldini (1988)
Poaceae
GI
Oleaceae
Asteraceae
OT
OT, RP
C
P, C
Pimpinella anisum L.
Pinus sp.
Pistacia lentiscus L.
Plantago lanceolata L.
Apiaceae
Pinaceae
Anacardiaceae
Plantaginaceae
Plantago major L.
Plantaginaceae
GI
LA
OT
SK
RP
SK
ct
sh
bi. ct, ho
C
C
O
C
C
C
RP
RE
GI
SK
RE
sh
ch
ra
as
ho
C
C
C
C
C
Guarrera (1981)
De Bellis (1978), Barbini et al. (1999)
Mearelli and Tardelli (1995)
Corrain (1977)
Maccioni and Marchini (1999)
Amici (1992), Corrain and Zampini (1961),
Corsi et al. (1981), Tammaro (1976), Viegi
et al. (1999)
Maccioni and Marchini (1999)
Tammaro (1984)
Guarrera (1987)
Manzi (1989)
De Capite and Menghini (1973)
ra
ho
ct, ho, ra
ct, sh
ct
ct
C
C
C
C
C
P, C
C
C
O
C
C
Plantago sp.pl.
Plantaginaceae
Polygonatum multiflorum
All., Polygonatum
officinale All.
Polygonum aviculare L.
Polygonum bistorta L.
Populus alba L.
Populus nigra L.
Liliaceae
Populus sp.
Potentilla reptans L.
Prunus avium L.
Salicaceae
Rosaceae
Rosaceae
Prunus cerasus L.
Pteridium aquilinum (L.)
Kuhn
Pulicaria dysenterica (L.)
Bernh.
Pulicaria odora (L.) Rchb.
Quercus cerris L.
Quercus cerris L., Quercus
pubescens Willd.
Quercus pubescens Willd.
subsp. pubescens
Quercus pubescens Willd.
Rosaceae
Hypolepidaceae
OT
OT
OT
OT
NE, OT
GI, OT
KI
GI
RP
KI
RP
Asteraceae
AP
do
Guarrera (1994)
Asteraceae
Fagaceae
Fagaceae
RE
SK
SK
as, mu
C
C
C
Fagaceae
SK, OT
Fagaceae
LA, SK,
OT
GI
SK
SK
SK
as
Guarrera (1994)
as, mu
go
C
C
C
C
Guarrera (1981)
De Simoni and Guarrera (1994)
Fossati et al. (1999)
Mearelli and Tardelli (1995)
Quercus robur L.
Quercus suber L.
Polygonaceae
Polygonaceae
Salicaceae
Salicaceae
Fagaceae
Fagaceae
230
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Quercus sp.
Ranunculus acris L.
Ranunculus bulbosus L.
Fagaceae
Ranunculaceae
Ranunculaceae
Ranunculus ficaria L.
Ranunculus lanuginosus L.
Raphanus raphanistrum L.
Ricinus communis L.
Robinia pseudacacia L.
Rosa canina L.
Ranunculaceae
Ranunculaceae
Brassicaceae
Euphorbiaceae
Fabaceae
Rosaceae
Rosmarinus officinalis L.
Rubia peregrina L.
Rubia tinctorum L.
Rubus fruticosus L.
Lamiaceae
Rubiaceae
Rubiaceae
Rosaceae
Rubus ulmifolius L.
Rubus sp. pl.
Rumex crispus L.
Rosaceae
Rosaceae
Polygonaceae
Rumex sp.
Polygonaceae
Ruta chalepensis L.
Rutaceae
Ruta graveolens L.
Kutacean
GI
LA
LA
SK
SK
LA
RP
SK
OT
RP
OT
GI
RP
RP
SK
GI
RE, OT
GI
SK
GI
FS
GI
SO
GI
GI
ct
ct
ra
sh
ct
sh
ct
sh
ca
du, gs
ct, ho
sh
do
ca, ch, ct
C
C
C
C
C
C
P
C
C
O
C
C
C
O
C
C
C
C
C
C
O
C
C
C
C
AP
ch, ct, ho
Saccharomicetaceae
AP
LA
GI
ho
ca
ct
P
C
C
Salicaceae
GI
ct, sh
SK, GI
GI, OT
GI
GI
AP, GI
GI
SK
AP
GI
ct
sh
ct
ct
ct
ct
pi
ct, do
C
C
C
C
C, O
C
C
P
C
RP, SK
AP
SK
ct
ch
ct
C
P
C
LA
RE
AP
SK
RP
GI
LA
SK
RP
SK
ct
ct
ct
ct
sh
ct, sh
C
C
O
C
P
C
C
C
P
C
SK, RP
Saccharomices cerevisiae
Rees
Salix alba L. subsp. alba
Salix alba L.
Salix viminalis L.
Salix sp.
Salicaceae
Salicaceae
Salicaceae
Salvia verbenaca L.
Sambucus ebulus L.
Sambucus nigra L.
Lamiaceae
Loniceraceae
Loniceraceae
Sambucus sp.
Sanguisorba minor Scop.
Santolina marchii Arrigoni
Satureja montana L.
Loniceraceae
Rosaceae
Asteraceae
Lamiaceae
Scorpiurus subvillosus L.
Scrophularia canina L.
Fabaceae
Scrophulariaceae
Scrophularia nodosa L.
Scrophulariaceae
KI
231
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Secale cereale L.
Sedum sp.pl.
Sempervivum tectorum L.
Poaceae
Crassulaceae
Crassulaceae
GI, RP
SK
GI
ca, pi
as, ho
ct, sh
C
C
C
GI
GI, OT
SK
SK
OT
GI
NE
RP
GI
GI
SK, RP
GI
GI
SK
SK
RP
OT
SK
AP
ch
ch
do
bi, pi
ct
sh
as, ho
ch
ct
do
P
C
C
C
O
C
C
O
C
C
C
C
C
C
C
O
C, M
C
P
OT
LA
RP
GI
GI
OT
OT
RE
GI
GI
LA, SK,
RP
SK
OT
GI
RP
GI
SK
RP, GI
GI
AP
ct, ho
ct
ct
ct
ct
ct
ct
ct
O
C
C
C
C
C
C
C
C
C
C
mu
ct, ho
sh
ct, ra
as
ct, sh
ct
C
C
C
P
C
C
P, M
P
P, O
SK
RP, OT
OT
GI
RP
RP
sh
tu
ct
ct, sh, tu
ch, do
C
P, O
P
C
C
O
SK, OT
RP, GI
RP
GI
OT
RP
AP
RP
RP
OT
ho
ct
ch
tu
ct
ch
ch, chs
O
C
O
P, C
P, O
O
C
P
O
P
Smilax aspera L.
Solanum melongena L.
Solanum nigrum L.
Liliaceae
Solanaceae
Solanaceae
Solanum tuberosum L
Solanaceae
Sonchus asper L.
Sonchus oleraceus L.
Sorbus domestica L.
Sorbus sp.pl.
Stachys germanica L.
Stachys officinalis (L.) Trev.
Stellaria media (L.) Vill.
Tamarix gallica L.
Tamus communis L.
Tanacetum vulgare L.
Asteraceae
Asteraceae
Rosaceae
Rosaceae
Lamiaceae
Lamiaceae
Caryophyllaceae
Tamaricaceae
Dioscoreaceae
Asteraceae
Teucrium chamaedrys L.
Lamiaceae
Thymus pulegioides L.
Lamiaceae
Thymus vulgaris L.
Thymus sp.
Lamiaceae
Lamiaceae
Triticum aestivum L.
Poaceae
Poaceae
Ulmaceae
Ulmus sp.
Urginea maritima (L.) Baker
Ulmaceae
Liliaceae
Urtica dioica L.
Urticaceae
Urticaceae
Urtica urens L.
Urticaceae
Urtica sp.pl.
Urticaceae
Guarrera (2002)
Viegi et al. (1999)
Manzi (1989)
Fossati et al. (1999)
Viegi et al. (1999)
Guarrera (1981, 1994, 2002)
Manzi (1989)
Corrain and Zampini (1961)
Lentini et al. (1988), Lentini and Aleo
(1991)
Lentini (1987)
Maccioni and Marchini (1998a, 1999)
De Bellis (1988), Fossati et al. (1999)
Corsi et al. (1981)
Manzi (1989), Nardelli (1987)
Bruni et al. (1997), Tammaro and
Pietrocola (1975)
Tammaro (1984)
Guarrera (1987)
Lentini et al. (1988)
Viegi et al. (1999)
De Bellis (1988), Fossati et al. (1999)
Nardelli (1987)
Tammaro and Pietrocola (1975)
Guarrera (1994)
Catanzaro (1968), Guarrera (1994)
Tammaro (1984), Guarrera (1994)
232
Table 1 (Continued )
Botanical taxon
Botanical family
Veterinary use
Animals
Cat.
References
Valeriana officinalis L.
Valerianaceae
NE
ch, do
Veratrum album L.
Liliaceae
Liliaceae
SK
OT
OT
AP
SK, OT
mu
pi
Sh
ct, sh
C
C
O
C
C
Liliaceae
Liliaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Scrophulariaceae
Verbascum thapsus L.
Scrophulariaceae
Verbascum sp.pl.
Verbena officinalis L.
Vicia ervilia (L.) Willd.
Vicia faba L.
Vitex agnus-castus L.
Vitis vinifera L.
Scrophulariaceae
Verbenaceae
Fabaceae
Fabaceae
Verbenaceae
Vitaceae
AP
AP
SK
SK, LA
OT
OT
SK, LA
SK
SK, LA
GI
OT
SK
SK
FS
GI, OT
OT
SK
OT, RP
sh
sh
sh
mu
sh
ho, mu, sh
ct
mu
pi
do
bi, sh
ca, pi, sh
ho
sh
C
C
C
C
O
C
C
C
C
C
O
C
C
O
C, P
C
LA
FS
GI
SK
OT, AP, LA
SK
GI
RP
chs
pi, sh
ca
ho, mu
sh
C
P, C
C
C
C
C
C, M
O
Zea mays L.
Poaceae
Guarrera (1994)
Renzetti and Taiani (1988), Guarrera (1995)
Barbagallo et al. (1979a)
Tammaro (1976)
Mearelli and Tardelli (1995)
De Bellis (1988)
Catanzaro (1970), Tammaro (1984)
Bellomaria (1982)
Tammaro (1984)
Corsi et al. (1981)
Mearelli and Tardelli (1995)
Guarrera and Tammaro (1991)
Amici (1992)
Guarrera (1990)
Manzi (1989)
Picchi (1999)
Viegi et al. (1999)
Nardelli (1987), Maccioni and Marchini
(1999)
Amici (1992), Guarrera (1994)
Guarrera (1994)
De Bellis (1988), Bellomaria (1982)
Guarrera (1981)
Guarrera (1994)
Corsi et al. (1981)
Guarrera (1987)
Manzi (1989)
Animals: ass, asses; ca, calves; ch, chickens; chs, chicks; ct, cattle; de, deers; do, dogs; du, ducks; go, goats; gs, gooses; ho, horses; la, lambs; mu,
mules; pi, pigs; ra, rabbits; sh, sheeps; tu, turkeys.
Categories: C, curative use; eu, external use; iu, internal use; M, magic use; O, other uses; P, preventive use.
Veterinary use: AP, antiparasitics and repellents; FS, food supplement; GI, gastrointestinal complaints; KI, kidney disorders; LA, ailments affecting the
locomotor apparatus; NE, nervous system disorders; OT, other ailments; RE, respiratory ailments; RP, reproductive disorders; SO, ailments of the sensory
organs; SK, problems of the skin and wounds.
Table 2
Personal communications on plant species reported for ethnoveterinary use in some regions of Italy
Family
Region
Part used
Use modality
Veterinary use/animals
Cat.
Personal communication
Allium sativum L.
Anemone hortensis L.
Apium nodiflorum (L.) Lag.
Artemisia absinthium L.
Avena sativa L.
Capsella bursa-pastoris (L.)
Medicus
Cistus monspeliensis L.
Cyclamen repandum S. et S.
Cynara cardunculus
subsp. scolymus (L.) Hayek
Daphne gnidium L.
Datura stramonium L.
Equisetum telmateja Ehrh.
Fraxinus ornus L.
Liliaceae
Ranunculaceae
Apiaceae
Asteraceae
Poaceae
Brassicaceae
Tuscany
Calabria
Tuscany
Tuscany
Campania
Apulia
bu
ap
ap
ap
bu
ep
iu
eu
Decoction, iu
iu
Fresh plant, iu
Decoction, iu
GI (ca)
SK (do)
GI
GI (ca)
RP (ct)
RP (ct)
C
C
C
C
P
Cistaceae
Primulaceae
Asteraceae
Apulia
Calabria
Liguria
le
ep
le
Cataplasms, eu
iu
Macerate, iu
SK (ho)
RP (pi)
GI (ct, tu)
C
C
C
Thymeleaceae
Solanaceae
Equisetaceae
Oleaceae
Calabria
Calabria
Calabria
Tuscany
ba
le, se
ep
ba
eu
Hung eu in the cowsheds
eu
Macerate in water, iu
O
M
O
P, O
De Fine
De Fine
De Fine
Pieroni
Oleaceae
Oleaceae
Ranunculaceae
Tuscany
Tuscany
Calabria
iba
ap
Basal le, pe
iu
Fumigations, eu
Inserted on the back of the ear or
under the fur of the neck, eu
SK (papillomas)
OT
OT (litterct, sh, ho)
GI (to prevent intestinal
diseases) (chs); RP (to increase
the output of eggs) (ch)
GI (ch)
RE (go)
RE
C
C
C, M
Ranunculaceae
Ranunculaceae
Ranunculaceae
Malvaceae
Tuscany
Tuscany
Tuscany
Campania
ap
ro
rh
ap
P, C
C
C
C
Pieroni
Maccioni and Guazzi
Maccioni and Marchini
Menale and Muoio
Fagaceae
Liliaceae
Tuscany
Calabria
ybr
ep
Decoction, eu
Inserted under the skin, eu
Inserted under the skin, eu
Boiled with Urtica dioica and bran
of oats, iu
Hydrolyte, eu
With broad beans, eu
C
O, P
Urtica dioica L.
Verbascum thapsus L.
Urticaceae
Scrophulariaceae
Tuscany
Tuscany
ap
ro
iu
With Allium sativum, iu
SK (do)
To take away the parasites in
granaries, in silos, etc.
OT(ch)
GI (ca)
C
C
Pieroni
Maccioni and Marchini
Taxon
Animals: ass, asses; ca, calves; ch, chickens; chs, chicks; ct, cattle; de, deers; do, dogs; du, ducks; go, goats; gs, gooses; ho, horses; la, lambs; mu, mules; pi, pigs; ra, rabbits; sh, sheeps; tu, turkeys.
Categories: C, curative use; eu, external use; iu, internal use; M, magic use; O, other uses; P, preventive use.
Part(s) used: ap, aerial parts; ba, bark; bu, buds; ep, entire plant; fr, fruits; iba, internal bark; le, leaves; pe, petiole; rh, rhizomes; ro, roots; se, seeds; ybr, young branches.
Veterinary use: AP, antiparasitics and repellents; GI, gastrointestinal complaints; KI, kidney disorders; LA, ailments affecting the locomotor apparatus; NE, nervous system disorders; OT, other ailments;
RE, respiratory ailments; RP, reproductive disorders; SO, ailments of the sensory organs; SK, problems of the skin and wounds.
233
234
Table 3
Botanical classification of taxa reported for ethnoveterinary use in Italy
Fungi
Ferns
Gymnosperms
Angiosperms
Dicots
Monocots
Crassulaceae (2)
Cucurbitaceae (3)
Ericaceae (2)
Euphorbiaceae (10)
Fabaceae (14)
Fagaceae (6)
Fumariaceae (1)
Gentianaceae (1)
Geraniaceae (2)
Hippocastanaceae (1)
Juglandaceae (1)
Lamiaceae (21)
Lauraceae (1)
Linaceae (1)
Loniceraceae (3)
Malvaceae (7)
Moraceae (2)
Myrtaceae (2)
Oleaceae (5)
Papaveraceae (3)
Plantaginaceae (3)
Polygonaceae (4)
Primulaceae (2)
Ranunculaceae (14)
Rosaceae (15)
Rubiaceae (3)
Rutaceae (2)
Salicaceae (6)
Scrophulariaceae (9)
Solanaceae (7)
Tamaricaceae (1)
Thymelaeaceae (2)
Ulmaceae (2)
Urticaceae (8)
Valerianaceae (1)
Verbenaceae (2)
Vitaceae (1)
Agavaceae (1)
Araceae (1)
Cactaceae (1)
Dioscoreaceae (1)
Liliaceae (14)
Poaceae (12)
242
30
gain, growth and development in various regions (EmiliaRomagna, Tuscany, Umbria) (Fossati et al., 1999; De
Bellis, 1988; Uncini Manganelli et al., 2001; Nardelli,
1987). Greater weight gain has been documented in fowl
fed with nettle compared to those fed with synthetic feed
(Nardelli, 1987). Species of the genus Urtica are rich in
aminoacids, proteins, minerals and vitamins, but they also
contain tannins, formic acid and salicylic acid (Guarrera,
1994). Urtica is a polyvalent genus. Indeed in Tuscany,
nettle (Urtica dioica) is given in the feed to cure diseases of
fowl (Pieroni, personal communication). In Abruzzo, nettle
boiled for a few minutes given to pigs would protect them
from contagious diseases; in the same region, to cows were
given a decoction consisting of one part of Urtica dioica
tips and two parts of Malva sylvestris and bran (nutritious,
Table 4
Number of taxa reported for ethnoveterinary use in various parts of Italy and number of taxa of the most represented families, compared with number
reported either in Gastaldo (1987) for the Italian medicinal flora or in Pignatti (1982) for the Italian Flora
Families
Apiaceae
Asteraceae
Euphorbiaceae
Fabaceae
Lamiaceae
Liliaceae
Poaceae
Ranunculaceae
Rosaceae
Central Italy
Southern Italy
Islands
3
12
4
3
2
3
4
8
4
9
19
3
11
15
5
14
5
10
0
1
1
0
1
2
0
3
0
2
3
5
0
2
2
3
3
1
Total number
(this paper)
Number, Gastaldo
(1987)
Number Pignatti
(1982)
13
30
10
14
21
14
12
14
15
60
116
12
51
74
27
18
46
49
223
688
73
394
194
176
443
172
238
235
100
90
80
70
n. of plants
60
50
40
30
20
Sicily
Sardinia
Apulia
Calabria
Molise
Campania
Latium
Abruzzo
Marches
Umbria
Liguria
Tuscany
Emilia Romagna
Venetum
Trentino-Alto Adige
Friuli-Venezia Giulia
10
236
Table 5
Use of plants in the Italian folk veterinary medicine (% of reports)
Antiinflammatory/digestive/antiparasitic/vulnerary
Antidiarrhoea/post partum problems/antiseptic/purgative and laxative/egg production/vermifuge
Antiecchymosis/appetiser/eutrophic/galactagogue/sedative
Antitussive/carminative/diuretic
Abortifacient/analgesic/dropsy/arthritis/antipyretic/respiratory diseases/antiviral/skin diseases/
aphrodisiac/hemostatic/ophthalmic/rubefacient
Toothache/snake bite/antitumoural/detergent/emollient/stimulant/antirheumatic/antigalactagogue/diseases of reproductive
system/rectal prolapse/kidney stones
>10%
510%
35%
23%
12%
<1%
Table 6
Plant species of folk veterinary remedies fairly or not yet investigated so far in phytopharmacology and phytotherapy
Species
Family
References
Asteraceaes
Apiaceae
Asteraceae
Liliaceae
Aspleniaceae
Aspleniaceae
Euphorbiaceae
Convolvulaceae
Malvaceae
Fabaceae
Lamiaceae
Liliaceae
Leguminosae
Poaceae
Asteraceae
Asteraceae
Fabaceae
Scrophulariaceae
Scrophularia nodosa L.
Sempervivum tectorum L.
Scrophulariaceae
Crassulaceae
237
238
under the tails of ewes which did not come into estrus
(Versilia) (Corsi et al., 1981). Documentation exists on
treatment with aerial parts of Rubia tinctorum for eight
consecutive days (Bertagnon, 1955). However, plants such
as Helleborus viridis and Juniperus sabina are potentially
toxic, and are feared by shepherds and farmers.
Infusion of Agrimonia eupatoria is used in Sicily as an antiinflammatory in cases of acute orchitis (Catanzaro, 1970).
3.4.6. Nervous system disorders
Few plants are reported for these disorders, six are
sedatives, seven antipyretics and three antiodontalgics. Unusual sedatives were Chrysanthemum parthenium (Liguria)
(Bandini, 1961) and Erica arborea (Campania) (De Feo
et al., 1992; De Feo and Senatore, 1993), whereas Papaver
rhoeas (Pantelleria) was regarded as facilitating assimilation of nutrients (Catanzaro, 1968). One use of Papaver
somniferum (stupefacient latex) was to calm down particularly lively horses and donkeys before taking them to market (Manzi, 1989). An unusual use of Lolium temulentum
as sedative (Manzi, 1989; Catanzaro, 1970) described for
Sicily and Abruzzo can be attributed to traces of alkaloids,
especially perloline (or temuline) and a mixture of fungi
of the genus Chaetomium which make it toxic (Gastaldo,
1987; Maugini, 1994). Indeed, cases of poisoning due to
mixing seeds of Lolium temulentum with wheat flour have
been documented. Against toothache, dangerous plants such
as Solanum nigrum (Picchi, 1999), Helleborus foetidus and
Helleborus bocconei (Leporatti and Pavesi, 1989) were used
in Calabria. The anesthetic properties of a similar species,
Helleborus niger, are known (Gastaldo, 1987; Schauenberg
and Paris, 1977). As antipyretics, atypical use of the latter
two species is documented, as well as Helleborus viridis,
used to treat a series of complaints known by local names
(Corsi and Pagni, 1978; Guarrera, 1994).
3.4.7. Ailments affecting the locomotor apparatus
Many plants are documented for treating dislocations
(seven), contusions (five), inflammation of the hooves
(three), fractures (three), lameness (three) and rheumatism
(two). Besides the usual plants (Brassica oleracea, Parietaria diffusa, Verbena officinalis) (Guarrera, 1987; De
Capite and Menghini, 1973; Nardelli, 1987; Amici, 1992)
for dislocations and contusions, also used in human folk
medicine, other less known plants such as Ranunculus acris
(Liguria) (Gastaldo et al., 1978) and Ranunculus bulbosus
(Sicily) (Barbagallo et al., 1979a) were reported. Many of
these plants are rubefacients, for example, Solanum nigrum
(solanin), Ranunculus sp.pl. (protoanemonin) and fruits
of Laurus nobilis; others are antineuralgics, for example,
Brassica oleracea (mirosin, mucilages, etc.) and Verbena
officinalis (verbenalin, verbenin, essential oil). Fractures
were immobilized with rudimentary casts made of spiny
fruits of Arctium lappa (Guarrera, 1990) or wheat flour
(Guarrera, 1994). Lameness was treated with Hyoscyamus niger and Satureja montana (Abruzzo) (Tammaro and
239
240
241
namic damage prevention, wound healing, antiinflammatory, immunomodulatory and antiviral activities, explaining
use of the bark in folk veterinary medicine (Kostova, 2001
and references therein).
Marrubium incanum and Marrubium vulgare, considered
to be medicinally equivalent in Lucania, were used against
foot and mouth disease in the northern part of this region.
Marrubium vulgare extracts were analysed in recent phytopharmacological studies and antioxidant, anti-spasmodic,
hypotensive, anti-asthmatic and analgesic properties were
found (Hnsel et al., 1999), most ascribed to its furanic labdane diterpenes. No other pharmacological studies have yet
been done and its folk veterinary properties still have to be
investigated.
4. Conclusions
The data gathered will be used to create the first national
databank of folk veterinary medicine. Further studies are underway into the veterinary heritage of Tuscany, Liguria, Calabria, Apulia, Lucania (Pieroni, 1999a, 2000, 2002a; Viegi
et al., 1999) and Molise (Guarrera, 2002), to complete the
databank. The databank, actually in Italian, will be available as soon as possible, in English, to all vets, botanists
and pharmacologists interested in the development of alternative therapies or involved in programs of sustainable land
use. For less known species, the databank could prompt phytochemical, biological and phytopharmacological research
into new therapeutic substances for veterinary drugs.
Acknowledgements
Many thanks to all persons who kindly provided references and interviews (Val di Cecina, Tuscany); to Giuseppe
De Fine (Cir, KR, Calabria) for use of his index; to Dott.
Rossella Muoio and Bruno Menale (Napoli, Campania),
Emanuele Guazzi (Grosseto, Tuscany), Giuseppe Marchini
(Castelnuovo Magra, SP, Liguria), Simonetta Maccioni
(Pisa, Tuscany) and Antonietta Rizzo (Arcola, SP, Liguria)
for unpublished data. The study was made possible by finance from Pisa University and MARS Nutritional Research
Council, Urbana, IL, USA.
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