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Photosynthesis
Autotrophs
Heterotrophs
Photosynthesis
process that converts solar energy into chemical
energy
directly or indirectly nourishes almost the entire
living world
occurs in plants, algae, certain other protists,
and some prokaryotes
(a) Plants
10 m
(d) Cyanobacteria
1.5 m
40 m
Copyright 2008 Pearson Education, Inc., publishing as Pearson Benjamin Cummings
Chloroplast
chlorophyll is in
the membranes
of thylakoids
Mesophyll
Stomata
Chloroplast
CO2
Outer
membrane
chloroplasts also
contain stroma,
a dense fluid
O2
Mesophyll cell
Thylakoid
Stroma
Granum
Thylakoid
space
1 m
5 m
Photosynthesis
summary equation:
Intermembrane
space
Inner
membrane
Photosynthesis
summary equation:
6 CO2 + 12 H2O + Light energy C6H12O6 + 6 O2 + 6 H2O
Word level
Carbon Dioxide + Water + Light Energy Glucose + Oxygen + Water
Products:
12 H2O
6 CO2
C6H12O6
6 H2O
6 O2
CO2
H 2O
Light
NADP+
ADP
+ P
Light
Reactions
Calvin
Cycle
ATP
NADPH
Chloroplast
O2
[CH2O]
(sugar)
105 nm 103 nm
103 nm
1 nm
Gamma
X-rays
rays
UV
106 nm
1m
(109 nm)
Microwaves
Infrared
103 m
Radio
waves
Photosynthetic Pigments
Visible light
380
450
500
550
600
650
700
750 nm
Longer wavelength
Lower energy
Shorter wavelength
Higher energy
Electromagnetic Spectrum
wavelengths that
are not absorbed
are reflected or
transmitted
Light
Reflected
light
Absorbed
light
Granum
Transmitted
light
CH3
CHO
(methyl) in chlorophyll a
(aldehyde/carbonyl) in chlorophyll b
Photosystem
Porphyrin ring:
light-absorbing
head of molecule;
note magnesium
atom at center
Structure of
chlorophyll molecules
in chloroplasts of plants
Hydrocarbon tail:
interacts with hydrophobic
regions of proteins inside
thylakoid membranes of
chloroplasts; H atoms not
shown
Photosystem
Photosystem
Photosystems
2 H+
+
1/ O
2
2
Primary
acceptor
3
e
e
Electron is
transferred to
primary
electron
acceptor
NADP+
reductase
Pc
5
P680
Electrons at a lower
energy level
provide energy for
the synthesis of ATP
Pigment
molecules
NADP+
+ H+
NADPH
P700
Light
6
ATP
Photosystem II
(PS II)
Fd
Cytochrome
complex
1 Light
Photon
energizes
pigment
molecule
and
electrons
and then
relayed up
to P680
Primary
acceptor
e
Pq
2
e
H2O
Electron is
photoexcited and
transferred to PSI's
primary electron
acceptor
Photosystem I
(PS I)
Comparison of
chemiosmosis in
mitochondria and
chloroplasts. In both
kinds of organelles,
electron transport
chains pump protons
(H+) across a membrane
from a
region of low H+
concentration to one of
high H+ concentration.
The protons then diffuse
back across
the membrane through
ATP synthase, driving
the synthesis of ATP.
Mitochondrion
Chloroplast
MITOCHONDRION
STRUCTURE
CHLOROPLAST
STRUCTURE
H+
Intermembrane
space
Inner
membrane
Diffusion
Electron
transport
chain
Thylakoid
space
Thylakoid
membrane
ATP
synthase
Key
Stroma
Matrix
ADP + P
Higher [H+]
Lower [H+]
H+
ATP
Calvin cycle
STROMA
(low H+ concentration)
Cytochrome
complex
Photosystem II
4 H+
Light
Photosystem I
Light
Fd
NADP+
reductase
H2O
e
1
THYLAKOID SPACE
(high H+ concentration)
e
2
Pc
2
1/
NADP+ + H+
NADPH
Pq
O2
4 H+
+2 H+
To
Calvin
Cycle
Thylakoid
membrane
three phases:
Carbon fixation (catalyzed by rubisco)
ATP
synthase
STROMA
(low H+ concentration)
ADP
+
Pi
Reduction
ATP
H+
Light reactions and chemiosmosis: the organization of the thylakoid membrane. Storing energy as a proton-motive force (H+
gradient , attributed by 1) Water is split by photosystem II on the side of the membrane facing tile thylakoid space; 2) as plastoquinone (Pq), a
mobile carrier, transfers electrons to the cytochrome complex, four protons are translocated across the membrane into the thylakoid space; 3)
a hydrogen ion is removed from the stroma when it is taken up by NADP+.
Input
3
CO2
(Entering one
at a time)
Rubisco
3 P
Short-lived
intermediate
3 P
Ribulose bisphosphate
(RuBP)
6
P
3-Phosphoglycerate
ATP
6 ADP
3 ADP
3
Calvin
Cycle
6 P
P
1,3-Bisphosphoglycerate
ATP
6 NADPH
Phase 3:
Regeneration of
the CO2 acceptor
(RuBP)
6 NADP+
6 Pi
P
5
G3P
6
P
Glyceraldehyde-3-phosphate
(G3P)
1
Output
P
G3P
(a sugar)
Phase 2:
Reduction
Glucose and
other organic
compounds