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Eddy Botanica Malacitana1~P  


Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba Artículo 219
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Fitosociología y sintaxonomía de los manglares y saladares de las


lagunas costeras de los cayos Coco y Sabinal, Cuba

Eddy Martínez-Quesada
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Correspondencia Resumen
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Abstract
Phytosociology and syntaxonomy of mangroves and halophilous plant
communities of coastal lagoons of Coco and Sabinal cays, Cuba.

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Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 221

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Taxon Comunidades Tipo sinantrópico Tipo biológico Tipo de hoja

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Casasia clusiifolia 8 - PLFURIDQHUyÀWR QRWyÀOD
Cienfuegosia yucatanensis  - QDQRIDQHUyÀWR OHSWyÀOD
Conocarpus erectus  - PDFURIDQHUyÀWR PLFUyÀOD
Cynanchum caribaeum  - WUHSDGRUD OHSWyÀOD
Distichlis spicata 9 - KHPLFULSWyÀWR -
Ernodea littoralis 8 - QDQRIDQHUyÀWR QDQyÀOD
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Fimbristylis cymosa  SDUDSyÀWR FDPpÀWR -
Fimbristylis ferruginea 11 - FDPpÀWR -
Flaveria linearis 2 - QDQRIDQHUyÀWR QDQyÀOD
Jacquemontia havanensis 8 LQWUDSyÀWRSLRQHUR QDQRIDQHUyÀWR OHSWyÀOD
Jacquinia keyensis 9 - PLFURIDQHUyÀWR PLFUyÀOD
Iva imbricata  - FDPpÀWR PLFUyÀOD
Lantana involucrata 8 LQWUDSyÀWRUHFXUUHQWH QDQRIDQHUyÀWR QDQyÀOD
Laguncularia racemosa  - PDFURIDQHUyÀWR PLFUyÀOD
Pilosocereus royenii 9 - VXFXOHQWD -
Pithecellobium keyense  - PLFURIDQHUyÀWR PLFUyÀOD
Pithecellobium mucronatum 9 - PLFURIDQHUyÀWR PLFUyÀOD
Rachicallis americana  - QDQRIDQHUyÀWR OHSWyÀOD
Rhizophora mangle  - PDFURIDQHUyÀWR PLFUyÀOD
Sarcocornia perennis  - QDQRIDQHUyÀWR -
6HOHQLFHUHXVJUDQGLÁRUXV  H[WUDSyÀWR VXFXOHQWD -
Senna chapmanii ,VHO\   - PLFURIDQHUyÀWR PLFUyÀOD
Sida ciliaris var. ciliaris  - FDPpÀWR OHSWyÀOD
Sideroxylon horridum 9 - PLFURIDQHUyÀWR PLFUyÀOD
Sesuvium maritimum  - FDPpÀWR OHSWyÀOD
Sesuvium portulacastrum  - FDPpÀWR QDQyÀOD
Sporobolus virginicus  - KHPLFULSWyÀWR -
Suaeda linearis  - QDQRIDQHUyÀWR QDQyÀOD
Suriana maritima  - QDQRIDQHUyÀWR QDQyÀOD
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Tillandsia paucifolia 9 - HStÀWDDUERUtFROD -

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1. Species of studied associations, sinantropic types, life forms and leaf types. 1. Rhizophoretum mangle Batido-Avi-
cennietum germinantisAvicennietum germinantis Avicennio germinantis-Rhizophoretum mangle, Lagunculario
racemosae-Rhizophoretum mangle  Batido-Conocarpodetum erecti  Lagunculario racemosae-Conocarpodetum
erectiConocarpodetum erectiSporobolo virginici-Conocarpodetum erectiBatidetum maritimaeSarcocor-
nietum perennis.
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TXH SHUPLWH LQFOXLU D WRGDV ODV FRPXQLGDGHV (Q 5+,=23+25(7$/,$&XDWUHFDVDV
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\ QRPHQFODWXUD GH ORV UDQJRV SULQFLSDOHV GH ODV KDOODDOQRUWH\VXUGHODJXQD7RUWXJXLOODHQWRGRV
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 Acta Botanica Malacitana1~P 

Figura 2)LWRFHQRVLVGHPDQJODUHVHQODVODJXQDVFRVWHUDVA Rhizophoretum mangleHQHOFDQDOGHHQWUDGDGH/DJXQD


/DUJDB Batido-Avicennietum germinantisDOVXUGHODODJXQDGHO3XHUWRVREUHSDYLPHQWRC 2HVWHGHOD/DJXQD/DV
&RORUDGDV VREUH DUHQD \ SDYLPHQWR &D\R &RFR D  Avicennietum germinantis HQ HO LQWHULRU GH OD /DJXQD 7LEXUyQ
GXUDQWH HO SHUtRGR SRFR OOXYLRVR E  /D PLVPD DVRFLDFLyQ \ ORFDOLGDG HQ SHULRGR OOXYLRVR F  Avicennio germinantis-
Rhizophoretum mangleDOQRUWHGHOD/DJXQD7RUWXJXLOODFD\R6DELQDOFigure 2. Mangrove phytocoenosis in the coastal
lagoons. A) 5KL]RSKRUHWXP PDQJOH at the entrance canal of Larga Lagoon, B) %DWLGR$YLFHQQLHWXP JHUPLQDQWLV, on
limestone pavement, at south of Puerto Lagoon, C) West of Las Coloradas lagoon, on sand and limestone pavement, D)
$YLFHQQLHWXPJHUPLQDQWLV inside Tiburón Lagoon during dry season, E) The same association and locality during rainy
season, Coco Cay, F) $YLFHQQLRJHUPLQDQWLV5KL]RSKRUHWXPPDQJOH at North of Tortuguilla Lagoon, Sabinal Cay.
Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 

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WHPSRUDOPHQWHLQXQGDGRV6XDUHDOELRJHRJUiÀFR XQD DOWXUD GH  D  FP /D HVSHFLH FRQVWDQWH
FRQRFLGRHVOD3HQtQVXODGH=DSDWDSDUWHFHQWUDO \ SRU OR JHQHUDO DEXQGDQWH HV Batis maritima /
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FD\R&RFRVHORFDOL]DSRUHOVXUGHODODJXQDGHO HVWUDWRXQDEXHQDFDQWLGDGGHLQGLYLGXRVMyYHQHV
3XHUWR QRUWH \ VXU GH OD ODJXQD 7LEXUyQ \ QRUWH OD IUHFXHQWH HV Sarcocornia perennis 0LOO  $-
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7DEOD

Batido-Avicennietum germinantis %RUKLGL\'HO5LVFRLQ%RUKLGL


(Rhizophoretea mangle, AvicennietaliaAvicennion occidentalis)

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Características de asociación y unidades superiores
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Sesuvium portulacastrum . .  . +.1 . . . . r.2 . r.1
( 
Rhizophora mangle ( r.1 . . . . . r.1 . . 2.1 . .
Sarcocornia perennis ( . . . . +.1 +.1 +.2  2.2 +.1 1.1 
Compañeras
Sporobolus virginicus . . +.2 . . . . . . r.2 2.2 
Sesuvium maritimum . . . . . .    . . .
Salicornia bigelovi . . . . +.1 +.1 . . . +.1 . .

Otras compañeras: En 2: Borrichia arborescens(QSuriana maritimaFlaveria linearis +.1.


/RFDOLGDGHV  QRUWH GH OD ODJXQD 7LEXUyQ >ƒҋµ1 ƒҋµ2@  RHVWH GH OD ODJXQD /DV &RORUDGDV
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VXUGHODGHODJXQDGHO3XHUWR>ƒҋµ1ƒҋµ2@FD\R&RFR
 Acta Botanica Malacitana1~P 

7DEOD Vt OR VRQ (VWD SXHGH VHU OD UD]yQ SRU OD FXDO
Avicennietum germinantis5H\HV $FRVWD
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(Rhizophoretea mangle, Avicennietalia Avicennietum germinantis FRPR XQ KHUED]DO
Avicennion occidentalis) GRQGH Avicennia germinans DOFDQ]D XQD DOWXUD
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\D TXH ODV HVSHFLHV GH PDQJOHV QR OOHJDQ D VHU QRUWHHQFD\R&RFR 
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7DEOD
Avicennio germinantis-Rhizophoretum mangle5H\HV $FRVWD
(Rhizophoretea mangle, AvicennietaliaAvicennion occidentalis)

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Características de asociación y unidades superiores

Avicennia germinans (   +.1    2.2 

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Batis maritima (   . . . . 1.2 +.2 .

Suriana marítima . . r.1 . . . . .

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Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 

P/DHVSHFLHGRPLQDQWHDEVROXWDHVA.germinans. (OHVWUDWRDUEyUHRSUHVHQWDXQDFREHUWXUDGHO
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SUHVHQWDQBatis maritima y Sarcocornia perennis. \PX\DEXQGDQWHHVAvicennia germinans7DPELpQ
(V PX\ SREUH HQ HVSHFLHV SRVHH  FRPR HV FRQVWDQWH \ D YHFHV DEXQGDQWH Rizophora
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GHQDQRIDQHUyÀWRV\PLFURIDQHUyÀWRV\SUHGRPLQDQ (O HVWUDWR KHUEiFHR OD PD\RU SDUWH GH ODV YHFHV
ODV KRMDV PLFUyÀODV 3RVHH XQ H[WUDSyÀWR Batis VH SUHVHQWD GLVSHUVR SHUR FXDQGR VH GHVDUUROOD
maritimaHQXQHQODFHQRVLV WDE  WLHQHXQDFREHUWXUDGHO\XQDDOWXUDGHFP
(VWD ÀWRFHQRVLV VH YH IDYRUHFLGD GXUDQWH /R LQWHJUDQ Batis maritima Suriana maritima / \
OD pSRFD OOXYLRVD PRPHQWR HQ HO TXH VXSHUD SOiQWXODVGHAvicennia germinans.
ORV  P GH DOWXUD D OR TXH WDPELpQ FRQWULEX\H (V PX\ SREUH HQ HVSHFLHV SRVHH  FRPR
OD GLVPLQXFLyQ GH ORV YDORUHV GH OD VDOLQLGDG SURPHGLR \ QLQJXQD HV HQGHPLVPR 3UHGRPLQDQ
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  HQ TXH OD SpUGLGD GH DJXD \ HO DXPHQWR ODV KRMDV PLFUyÀODV ([LVWH XQ H[WUDSyÀWR Batis
GH VDOLQLGDG SURYRFDQ OD IRUPD DFKDSDUUDGD GH maritimaHQXQHQODFHQRVLV WDE 
OD PLVPD 6HJ~Q 0DFNHH   ORV PDQJODUHV
GRPLQDGRV SRU A.germinans SUHVHQWDQ VXHOR (VWD ÀWRFHQRVLV VH GHVFULELy VROR FRQ
DOWDPHQWHUHGXFWRU\XQDOWRFRQWHQLGRGHVXOIXURV Avicennia germinans TXH WLHQH ORV PD\RUHV
(VDVFDUDFWHUtVWLFDVXQLGDVDODDOWDFRQFHQWUDFLyQ YDORUHV GH DEXQGDQ   FLD \ Rhizophora mangle
VDOLQDGXUDQWHODpSRFDSRFDOOXYLRVDFRQSHULRGRV FRQ ORV PHQRUHV HOOR FRLQFLGH FRQ HO UHVXOWDGR
GHLQXQGDFLyQVRQODFDXVDGHODEDMDULTXH]DGH GH HVWH HVWXGLR DXQTXH VH SUHVHQWDQ WDPELpQ
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5H\HV 'RPtQJXH] $FRVWD &DQWLOOR LQ 5H\HV Lagunculario racemosae-Rhizophoretum mangle
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SDUWHRULHQWDOGH&XED(QHVWHHVWXGLRVHHQFRQWUy 3XHUWR\DOHVWHGHODODJXQD/DUJDHQFD\R&RFR
DOQRUWHGHODODJXQD7RUWXJXLOODHQFD\R6DELQDO (O HVWUDWR DUEyUHR WLHQH XQD FREHUWXUD GH 

7DEOD
Lagunculario racemosae-Rhizophoretum mangle 'HOJDGLOOR3HLQDGR0DUWtQH]3DUUDV $OFDUD]
(Rhizophoretea mangle, AvicennietaliaAvicennion occidentalis)

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Características de asociación y unidades superiores
Rhizophora mangle (  2.1     . +.1 2.1
Conocarpus erectus   2.1 2.1   2.1  
Laguncularia racemosa +.1 2.1 2.1     +.1 2.1
Avicennia germinans +.1 1.1 +.1 +.1 1.1 +.1 +.1 2.1 .
Compañeras
Batis maritima ( r.1 . r.1 . . . . . .
Sarcocornia perennis . r.2 . . r.2 . . . .
Borrichia arborescens r.1 . . . . . . . .
Sesuvium portulacastrum r.1 . . . . . . . .
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228 Acta Botanica Malacitana1~P 

Figura 3)LWRFHQRVLVGHPDQJODUHV\VDODGDUHVHQODVODJXQDVFRVWHUDVA Lagunculario racemosae-Rhizophoretum


mangleDOQRUWHGHODODJXQDGHO3XHUWRFD\R&RFR B Batido-Conocarpodetum erectiDOQRUWHGHODODJXQD7RUWXJXLOOD
FD\R 6DELQDO C  Lagunculario racemosae-Conocarpodetum erecti DO QRUWH GH OD ODJXQD7RUWXJXLOOD FD\R 6DELQDO D 
Conocarpodetum erecti DO RHVWH GH OD ODJXQD /D 6DOLQD FD\R 6DELQDO E  Sporobolo virginici-Conocarpodetum erecti
DOVXUGHODODJXQD7RUWXJXLOODFD\R6DELQDO F Batidetum maritimaeHQHOLQWHULRUGHODODJXQD/RV&DLPDQHVFD\R
6DELQDOG Sarcocornietum perennisDOVXUGHODODJXQD7RUWXJXLOODFD\R6DELQDOGXUDQWHODpSRFDVHFDMangrove
phytocoenosis and salt marshes in coastal lagoons Figure 3. A) /DJXQFXODULR UDFHPRVDH5KL]RSKRUHWXP PDQJOH at
North of Puerto Lagoon, Coco Cay, B) %DWLGR&RQRFDUSRGHWXPHUHFWL at North of Tortuguilla Lagoon, Sabinal Cay, C)
/DJXQFXODULRUDFHPRVDH&RQRFDUSRGHWXPHUHFWL, at North of Tortuguilla Lagoon, Sabinal Cay, D) &RQRFDUSRGHWXPHUHFWL
at West of La Salina Lagoon, Sabinal Cay, E) 6SRURERORYLUJLQLFL&RQRFDUSRGHWXPHUHFWL at South of Tortuguilla Lagoon,
Sabinal Cay, F) %DWLGHWXPPDULWLPDH inside of Los Caimanes Lagoon, Sabinal Cay, G) 6DUFRFRUQLHWXPSHUHQQLV at South
of Tortuguilla Lagoon, Sabinal Cay, during dry season.
Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 229

7DEOD
Batido-Conocarpodetum erecti Martínez-Quesada ass. nova
(Rhizophoretea mangle, Combretalia, Batido maritimae-Conocarpodion erecti)

Variantes Sesuvium portulacastrum Fimbristylis cymosa


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Características de asociación y de alianza
Conocarpus erectus  (       
Batis maritima ( 1.2 1.2  +.2 +.2  
Diferenciales de variantes
Sesuvium portulacastrum (  1.2  . . . .
Rachicallis americana r.1 r.1 +.1 . . . .
Fimbristylis cymosa . . .   r.2 r.2
Sesuvium maritimum . . . +.2 +.2 1.2 1.1
Sporobolus virginicus . . . +.2 +.2 1.2 
Broughtonia lindenii . . . r.1 r.1 r.1 .
6HOHQLFHUHXVJUDQGLÁRUXV . . . . r.1 r.1 .
Sarcocornia perennis ( . . . . 2.2 1.2 .
Evolvulus convolvuloides . . . r.1 r.1 . .
Cienfuegosia yucatanensis . . . +.1 r.1 . .
Ambrosia hispida . . . r.1 +.1 . .
Compañeras
Suaeda linearis ( r.1 . . . . 1.1 .
Otras compañeras:(QSalicornia bigellovii(QIva imbricataTabebuia trachycarpaUSida ciliaris var. ciliarisU
Cynanchum caribaeumUSenna chapmaniiUPithecellobium keyenseU(QAvicennia germinans r.1.

D  \ XQD DOWXUD GH  D  P /DV HVSHFLHV FRQXQDEDMDDEXQGDQFLD(VFRLQFLGHQWHHOKHFKR


FRQVWDQWHV VRQ Conacarpus erectus y Rizophora GHODEUXSWRHFRWRQRFRQRWUDVFRPXQLGDGHVFRPR
mangleODPiVDEXQGDQWHVAvicennia germinans HOERVTXHVLHPSUHYHUGHDOQRUWHGHODODJXQDGHO
y Laguncularia racemosa(OHVWUDWRKHUEiFHRHVWi 3XHUWR\HOPDWRUUDO[HURPRUIRFRVWHURDOHVWHGH
GLVSHUVRQRREVWDQWHVHREVHUYDQBatis maritima ODODJXQD/DUJD(Q&XEDQRVHKDEtDUHJLVWUDGR
y Sarcocornia perennis$GHPiVVHDSUHFLDQFRPR
SDUWHGHODUHJHQHUDFLyQGHOERVTXHDMXYHQLOHVGH Combretalia &XDWUHFDVDV
R. mangleA. germinans y L. racemosa. Batido-Conocarpodetum erecti Martínez-Quesada
(VSREUHHQHVSHFLHVSRVHHFRPRSURPHGLR ass. nova
\ QLQJXQD HV HQGHPLVPR 6H SUHVHQWDQ ORV >7DEODholotypus assLQYÀJ%@
QDQRIDQHUyÀWRV\PDFURIDQHUyÀWRVFRQSUHGRPLQLR &DUDFWHUtVWLFDV GH DVRFLDFLyQ Conocarpus
GHODVKRMDVPLFUyÀODV3RVHHXQH[WUDSyÀWRBatis erectus y Batis maritima.
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DSDUHFH FRPR HVSHFLH GRPLQDQWH KD VLGR GHO 3XHUWR HQ FD\R &RFR \ DO QRUWH GH ODJXQD
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'HOJDGLOOR HW DO   \ 3HLQDGR HW DO   (OHVWUDWRDUEyUHRWLHQHXQDFREHUWXUDGHOD
DXQTXHVLQConocarpus erectusHQWUHODVHVSHFLHV \XQDDOWXUDGHDP/DHVSHFLHFRQVWDQWH
GH PDQJOHV \ HQ JHQHUDO SRU OD IDOWD GH RWUDV \ PX\ DEXQGDQWH HV Conocarpus erectus. El
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 Acta Botanica Malacitana1~P 

7DEOD VHKDQGHVFULWRGRVDVRFLDFLRQHVDistichli spicatae-


Lagunculario racemosae-Conocarpodetum erecti
Conocarpodetum erecti5H\HV'RPtQJXH]\
3HLQDGR$OFDUD] 'HOJDGLOOR(Rhizophoretea mangle,
Lycio caroliniani-Conocarpodetum erecti 5H\HV
Combretalia, Conocarpo-Laguncularion)
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,QYHQWDULRQƒ      FRQGLFLRQHV HFROyJLFDV HQ ODV TXH VH GHVDUUROOD
(VWUDWRDUEyUHR(       ODVTXHVHSXHGHQFRQWUDVWDUFRQODVGHODDOLDQ]D
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GH GLFKD DOLDQ]D FRQ XQD DOWD SUHVHQFLD HQ ORV
Características de asociación y unidades superiores LQYHQWDULRVGHODDVRFLDFLyQ
Laguncularia racemosa     
( Lagunculario racemosae-Conocarpodetum erecti
Conocarpus erectus +.1 r.1   2.1 3HLQDGR$OFDUD] 'HOJDGLOOR
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FREHUWXUDGHOD\XQDDOWXUDGHDFP [HURPRUIRFRVWHURLQWHUUXPSLGRSRUXQWHUUDSOpQ\
/DFRQVWDQWH\DYHFHVSXQWXDOPHQWHDEXQGDQWHHV XQD FRPXQLGDG GH Rhizophoretum mangle. Está
Batis maritima0HQRVIUHFXHQWHVVRQBroughtonia HQVXHORVDUHQRVRVFRQLQXQGDFLRQHVWHPSRUDOHV
lindenii /LQGO  'UHVVOHU  Fimbristylis cymosa GH PHQRU GXUDFLyQ WUHV D FXDWUR PHVHV  TXH ORV
5 %U  Rachicalis americana -DFT  +LWFKF PDQJODUHVGHAvicennietalia.
Sesuvium maritimum /  / S. portulacastrum (OHVWUDWRDUEyUHRHVHO~QLFRH[LVWHQWHFXEUH
/  / \ Sporobolus virginicus, \ ODV RFDVLRQDOHV HQWUH HO  \  GH OD VXSHUÀFLH \ DOFDQ]D XQD
VRQ Ambrosia hispida 3XUVK Cienfuegosia DOWXUDHQWUH\PLaguncularia racemosa
yucatanensis 0LOOVS Evolvulus convolvuloides HV OD HVSHFLH FRQVWDQWH \ GRPLQDQWH$GHPiV HV
:LOOG H[ 6FKXOW  6WHDUQ Sarcocornia perennis, FRQVWDQWHConocarpus erectus.
6HOHQLFHUHXV JUDQGLÁRUXV /  %ULWWRQ  5RVH \ (VH[WUHPDGDPHQWHSREUHHQHVSHFLHVSRVHH
Suaeda linearis (OOLRWW 0RT/DVLQXVLDGHOLDQDV GRV QR HQGpPLFDV TXH VRQ PDFURIDQHUyÀWRV GH
VH FRPSRQH GH Metastelma cubense *ULVHE \ KRMDVPLFUyÀODVQRVLQDQWUySLFDV WDE 
6HOHQLFHUHXV JUDQGLÁRUXV \ OD GH HStÀWDV SRU (Q OD OLWHUDWXUD ÀWRVRFLROyJLFD VH DSUHFLD TXH
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3UHVHQWD GRV YDULDQWHV GH Fimbristylis Laguncularia racemosa 3UHYLDPHQWH 'HOJDGLOOR
cymosaTXHVHDSUHFLDHQORVLQYHQWDULRVD HW DO   \ 3HLQDGR HW DO   VH UHÀULHURQ
y de Sesuvium portulacastrum HQ ORV LQYHQWDULRV a Lagunculario racemosae-Rhizophoretum
\/DSULPHUDVHGLIHUHQFLDSRUVHUPiV mangle y a Lagunculario racemosae-Avicennietum
ULFD ÁRUtVWLFDPHQWH DO FROLQGDU FRQ XQ ERVTXH germinantisGHODFRVWDGH%DMD&DOLIRUQLD0p[LFR
VLHPSUHYHUGH GH GRQGH SUREDEOHPHQWH SURYLHQH 0iV UHFLHQWHPHQWH 5H\HV 'RPtQJXH]  
XQDSRUWHGHQXWULHQWHVSRUDUUDVWUHGHODVOOXYLDV GHVFULELy HO Laguncularietum racemosae XQ
/DDVRFLDFLyQHVPX\SREUHHQHVSHFLHVSRVHH PDQJODUTXHURGHDODODJXQD%LUDPDHQODSURYLQFLD
 FRPR SURPHGLR \ QLQJXQD HV HQGHPLVPR /DV7XQDV &XED GRQGHL. racemosaHVODHVSHFLH
6H FRPSRQH SULQFLSDOPHQWH GH FDPpÀWRV \ GRPLQDQWHDEVROXWD6HJ~Q0HQpQGH]&DUUHUDHW
QDQRIDQHUyÀWRV \ SUHGRPLQDQ ODV KRMDV OHSWyÀODV DO   HQ HO HFRVLVWHPD 6DEDQD&DPDJH\
\ PLFUyÀODV 3RVHH FLQFR HVSHFLHV VLQDQWUySLFDV QR H[LVWH QLQJ~Q ERVTXH FRQ GRPLQDQFLD GH L.
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JUDQGLÁRUXV XQ SDUDSyÀWR Fimbristylis cymosa UHSUHVHQWDFLyQ GH OD HVSHFLH HQ ORV PDQJODUHV
XQ LQWUDSyÀWR SLRQHUR Evolvulus convolvuloides Lagunculario racemosae-Conocarpodetum erecti
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*ULVHE  .6FKXP HQ XQ  HQ OD FHQRVLV GHPXHVWUDORLQVXÀFLHQWHPHQWHHVWXGLDGRTXHHVWi
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(VWDÀWRFHQRVLVVHXELFDHQODDOLDQ]DBatido GRPLQDQWH SRU VX DOWD DEXQGDQFLD (Q &XED QR
maritimae-Conocarpodion erectiTXHVHGHVFULELy VHFRQRFtDGHVXSUHVHQFLD6LQWD[RQyPLFDPHQWH
GHODVGXQDVDUHQRVDV\FRQFKDVPDULQDV 5H\HV Lagunculario racemosae-Conocarpodetum erecti
'RPtQJXH] RVRORVREUHHVWDV~OWLPDVHQ VH LQFOX\y SULPHUR HQ /DJXQFXODULR UDFHPRVDH
ODRULOODGHOPDURHQWUHORVPDQJODUHVRDOERUGH $YLFHQQLRQ JHUPLQDQWLV DKRUD Avicenion
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Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 

7DEOD

Conocarpodetum erecti5H\HV'RPtQJXH]
(Rhizophoretea mangle, Combretalia, Conocarpo-Laguncularion)

,QYHQWDULRQƒ    81 82
(VWUDWRDUEyUHR(      
(VWUDWRKHUEiFHR(  GLVS GLVS GLVS GLVS GLVS
1ƒHVSHFLHV     

Características de asociación y unidades superiores

Conocarpus erectus (     

Compañeras

Suriana maritima E1 +.1 +.1 +.1 r.1 r.1


Lantana involucrata +.1 r.1 +.1 2.1 r.1
Casasia clusiifolia( r.1 . r.1 . +.1
Erithalis fruticosa E1 . r.1 r.1 r.1 r.1
Ernodea littoralis . . . r.1 r.1

Otras compañeras:(QRachicalis americanaU(QJacquemontia havanensis.

/RFDOLGDGHV: QRUWHGHODODJXQD7RUWXJXLOOD>ƒҋµ1ƒҋµ2@FD\R6DELQDOGLVSGLVSHUVR

Tabla 10

Sporobolo virginici-Conocarpodetum erecti 0DUWtQH]4XHVDGD


Rhizophoretea mangleCombretaliaConocarpo-Laguncularion

,QYHQWDULRQƒ      88 89  91
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Características de asociación y unidades superiores

Conocarpus erectus (         
Laguncularia racemosa . . . . . . r.1 +.1 +.1
Avicennia germinans ( . . . . r.1 . . r.1 r.1

Compañeras

Sporobolus virginicus ( 2.2 2.2    +.2   
Batis maritima . . +.2 +.2 1.2 1.2 1.2 +.2 +.2
Sarcocornia perennis . . 2.2 +.2 1.2 +.2 . 1.2 +.2
Jacquinia keyensis r.1 r.1 r.1 +.1 . . . . .
Pilosocereus royenii ( r.1 r.1 r.1 . . . . . .
Sesuvium maritimum r.1 . +.2 2.1 . . . . .
Sesuvium portulacastrum . . . . . . 1.2 +.2 .
Pithecellobium mucronatum r.1 +.1 . . . . . . .
Tillandsia paucifolia r.1 . . +.2 . . . . .

Otras compañeras(Q%XFLGDPROLQHWLLU(Q6LGHUR[\ORQKRUULGXPU(Q'LVWLFKOLVVSLFDWD

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 Acta Botanica Malacitana1~P 

HQOD]RQDGHSOHDPDU 3HLQDGRHWDO VLQ Casasia clusiifolia -DFT  8UE Erithalis fruticosa


HPEDUJRHQXQWUDEDMRSRVWHULRU&RUWpV&DVWLOOR  / Ernodea littoralis 6Z Lantana involucrata /
5DQJHO&K   OD WUDQVÀULHURQ D /DJXQFXODULR y Suriana maritima \ ODV HVSHFLHV FRQVWDQWHV
UDFHPRVDH5KL]RSKRULRQ PDQJOH DKRUD VRQ Ernodea littoralis L. involucrata, y Suriana
Rhizophorion occidentalis TXHVRQPDQJODUHVTXH maritima PLHQWUDV TXH Casasia clusiifolia es
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LJXDOHVFRQGLFLRQHVHFROyJLFDVTXHODVREVHUYDGDV SURPHGLR \ QLQJXQD HV HQGHPLVPR 6H FRPSRQH
HQHVWHHVWXGLR6HFRQVLGHUDTXHOD/DJXQFXODULR HQVXPD\RUtDGHORVQDQRIDQHUyÀWRV\SUHGRPLQDQ
&RQRFDUSRGHWXPGHEHVHUXELFDGDHQ&RQRFDUSR ODV KRMDV QDQyÀODV 3UHVHQWD XQ LQWUDSyÀWR
/DJXQFXODULRQ 6LQ /DJXQFXODULR UDFHPRVDH UHFXUUHQWH Lantana involucrata \ XQ LQWUDSyÀWR
$YLFHQQLRQ JHUPLQDQWLV  \D TXH HV XQ PDQJODU SLRQHUR Erithalis fruticosa HQ XQ  HQ OD
GH PX\ EDMD DOWXUD HVWDFLRQDOPHQWH LQXQGDGR FHQRVLV WDE 
FDVL VLHPSUH DOHMDGR GH OD LQÁXHQFLD GLUHFWD GH 5H\HV'RPtQJXH] 5H\HV'RPtQJXH] $FRVWD
ODVPDUHDV3RUHOORORVYDORUHVGHVDOLQLGDGHQHO &DQWLOOR GHVFULELyHO&onocarpodetum erecti
VXHORGHEHQÁXFWXDUDQXDOPHQWH)ORUtVWLFDPHQWH FRPRXQPDQJODUSXURGHConocarpus erectus y sin
OD SUHVHQFLD FRQVWDQWH GH &RQDFDUSXV HUHFWXV XQHVWUDWRKHUEiFHR(QFD\R6DELQDOVHSUHVHQWD
DXQTXH QR VLHPSUH FRQ DOWRV YDORUHV GH VX XQERVTXHGRQGHHOHVWUDWRDUEyUHRHVWiGRPLQDGR
DEXQGDQFLDGRPLQDQFLD WDPELpQ MXVWLÀFD VX SRU HVD HVSHFLH \ DSDUHFHQ DOJXQDV KHUEiFHDV
LQFOXVLyQHQHVWDDOLDQ]D TXHWLHQHQXQDEDMDDEXQGDQFLDGRPLQDQFLDFRPR
RFXUUHHQFD\R$QFOLWDV 0DUWtQH]4XHVDGD 
Conocarpodetum erecti 5H\HV 'RPtQJXH] LQ (VD HV OD UD]yQ SRU OD FXDO QR VH FRQVLGHUD XQD
5H\HV'RPtQJXH] $FRVWD&DQWLOOR DVRFLDFLyQGLVWLQWD
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SDtV6HREVHUYyDOQRUWHGHODODJXQD7RUWXJXLOOD\ REVHUYyDOVXUGHODODJXQD/RV&DLPDQHVVXUGH
DORHVWHGHODODJXQD/D6DOLQD ODODJXQD7RUWXJXLOOD\VXUGHODJXQD/D6DOLQDHQ
(OHVWUDWRDUEyUHRFXEUHGHODO\WLHQH FD\R6DELQDO
XQD DOWXUD TXH YD GHVGH  D  P /D HVSHFLH (O HVWUDWR DUEyUHR SRVHH XQD FREHUWXUD GHO
GRPLQDQWH DEVROXWD HV Conocarpus erectus. El  D  \ XQD DOWXUD GH  D  P /D FRQVWDQWH
HVWUDWR KHUEiFHR HVWi GLVSHUVR 6H REVHUYDQ \ PX\ DEXQGDQWH HV Conocarpus erectus.

7DEOD

Batidetum maritimae6DPHN
(Batido-SarcocornieteaBatido -SarcocornietaliaFimbrystilo-Sarcocornion perennis)

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Característica de asociación y unidades superiores
Batis maritima (        
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Salicornia bigelovi . . . . . +.1 +.1 r.1
Compañeras
Avicennia germinans . . . . . +.1 +.1 +.1
Sarcocornia perennis . . . . . . . +.2
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Eddy Martínez-Quesada. Fitosociología y sintaxonomía de los manglares y saladares de Cuba 

6RQ PHQRV IUHFXHQWHV Avicennia germinans, GH ODV HVSHFLHV GH DGDSWDUVH D HOODV R GH VX
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(V FRQVWDQWH Sporobolus virginicus IUHFXHQWHV ÀWRFHQRVLV Bucida molinetii 0*yPH] $OZDQ 
VRQ Batis maritima y Sarcocornia perennis FRPR 6WDFH Jacquinia keyensis, Pilosocereus royenii,
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portulacastrum. /D VLQXVLD GH HStÀWDV VROR HVWi PLVPDDVRFLDFLyQDXQTXHVHHQFXHQWUHHQ]RQDV
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FD\R $QFOLWDV VREUH DUHQDV FDOFiUHDV HyOLFR Batidetum maritimae 6DPHN
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WHPSRUDOPHQWH \ DXQTXH VH HQFRQWUDURQ WDPELpQ FRVWD QRUWH GH OD SURYLQFLD /D +DEDQD HQ OD
HQ HO HVWUDWR DUEyUHR D Avicennia germinans y SDUWH FHQWUDO GH &XED SHUR GHEH HVWDU PiV
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\ WLHQHQ EDMD DEXQGDQFLDGRPLQDQFLD  $GHPiV HQ HO LQWHULRU GH OD ODJXQD /RV &DLPDQHV SRU HO
de Sporobolus virginicus, HQ HO HVWUDWR KHUEiFHR QRUWH GRQGH FRQVWLWX\H XQ JUDQ SDUFKH GH Batis
VH SUHVHQWDQ PXFKDV PiV HVSHFLHV TXH HQ FD\R maritima FDVL SXUR TXH VH GHVDUUROOD VREUH XQ
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7DEOD

Sarcocornietum perennis 6DPHN


(Batido-SarcocornieteaBatido -Sarcocornietalia
Fimbrystilo-Sarcocornion perennis)

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Características de asociación y unidades superiores

Sarcocornia perennis (     

Suaeda linearis +.1 +.1 +.1 +.1 1.1

Fimbristylis cymosa +.2 +.2 . . .

Compañeras

Conocarpus erectus (  2.1 1.1 1.1 1.1

Sesuvium maritimum +.2 +.2 r.2 +.2 1.2

Sporobolus virginicus  2.2 2.2 2.2 2.2

Frimbristylis ferruginea +.2 . . . .

/RFDOLGDGHVVXUGHODODJXQD7RUWXJXLOOD>ƒҋµ1ƒҋµ2@FD\R6DELQDO
 Acta Botanica Malacitana1~P 

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/DJXQD /DV &RORUDGDV B  /DJXQD /DUJD H[WUHPR
RULHQWDO \ C  /DJXQD 7LEXUyQ  Batido-Avicennietum
germinantis, 2- Rhizophoretum mangle,Lagunculario
racemosae-Rhizophoretum mangle,  Avicennietum
germinantis,  %RVTXH VHPLGHFtGXR PLFUyÀOR Figure Figura 5 =RQDFLyQ GH ODV ÀWRFHQRVLV GH PDQJODU A 
4. Mangrove phytocoenosis zoning. A) Las Coloradas /DJXQD/RV&DLPDQHVB /DJXQDGHO3XHUWRC /DJXQD
Lagoon, B) Larga Lagoon, west part, and C) Tiburón 7RUWXJXLOODD /DJXQD/D6DOLQDBatido-Avicennietum
Lagoon. germinantis, 2- Batidetum maritimae,  Sporobolo
virginici-Conocarpodetum erecti,  Lagunculario
HQFD\R6DELQDO racemosae-Rhizophoretum mangle,  Rhizophoretum
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GHO  D   \ XQD DOWXUD GH  D  FP /D germinantis-Rhizophoretum mangle, 8- Sarcocornietum
perennis,%RVTXHVLHPSUHYHUGHPLFUyÀOR0DWRUUDO
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[HURPRUIR FRVWHUR Figure 5. Mangrove phytocoenosis
marítima,PHQRVIUHFXHQWHVHREVHUYDDSesuvium zoning. A) Los Caimanes Lagoon, B) El Puerto Lagoon,
portulacastrum /  / 6H HQFXHQWUDQ WDPELpQ C) Tortuguilla Lagoon, D) La Salina Lagoon.
VREUHSDYLPHQWRFDUVWLÀFDGRAvicennia germinans
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