Elenco y Biogeografía de Los Ácaros Acuáticos de Sudamerica

Graellsia, 61(2): 181-224 (2005)
ELENCO Y BIOGEOGRAFA DE LOS CAROS ACUTICOS (ACARI,

PARASITENGONA, HYDRACHNIDIA) DE SUDAMRICA
B. Rosso de Ferrads1 y H. R. Fernndez2
RESUMEN
Las publicaciones de catlogos y listados de especies determinan una clara conexin entre
taxonoma bsica y temas relacionados con la biodiversidad. Segn algunas estimaciones, solo
el 10-30% del nmero total de especies ha sido nominado.
El trabajo que se presenta aqu insumi aos de trabajo, en la catalogacin de los verdaderos caros acuticos Parasitengona de Amrica del Sur. Se agrega adems informacin sobre la
distribucin por regiones y hbitats conocidos para las especies.
Hasta el presente hay 6 superfamilias, 23 familias, 118 gneros y 916 especies catalogados
en 11 pases de Amrica del Sur. El grado de conocimiento vara enormemente de pas a pas
con numerosos datos de Brasil y ninguno de Guayana Francesa.
Palabras claves: Biodiversidad, Acari, distribucin, hbitat, caros acuticos, taxonoma.
ABSTRACT
Checklist and biogeography of water mites (Acari, Parasitengona,
Hydrachnidia) from South America
Checklist and catalog publications demonstrate a clear connection between basic taxonomy
and biodiversity issues. According to some estimates, only 10-30% of all global species have
been named. As in other fields, catalogs provide an important source of information concerning
species diversity in freshwater ecology.
South America is a continent dominated by freshwater ecosystems. The tremendous habitat
diversity created by this landscape supports a high number of arthropods, including water mites,
which belong to the hyperdiverse group Acari. South America has a substantial task ahead in
cataloging its biodiversity.
Much has been published on water mites in South America. In fact, according to Besch,
water mites were the most studied with the exception of Europe up until the 1960s. Most of
the collections were conducted by two acarologists (Lundblad and Karl Viets) during the
1940s. Today, the collection, identification and description process of water mites is slower.
In the 1980s, the north-american acarologist D. R. Cook produced two lengthy papers about
neotropical water mites in four regions of South America. Recently, several Argentine acarologists have published papers on water mites from diverse habitats and regions in South
America.
The catalog presented here includes information regarding 6 superfamilies, including 23
families in 118 genera of true water mites (Hydrachnidia, Parasitengona, Acari). It also includes
1
2
CONICET, Universidad Nacional de Crdoba, Av. V. Sarsfield 299, 5000 Crdoba, Argentina
CONICET, Universidad Nacional de Tucumn, Av. M Lillo 205, 4000 San Miguel de Tucumn, Argentina.
182
ROSSO DE FERRADS & FERNNDEZ
the references concerning the species, distribution in various regions of South America and as
far as known habitat.
At present, there are 916 species from several authors cataloged in 11 countries in South
America. The degree of knowledge varies greatly from country to country, with numerous
entries for Brazil and none for French Guiana.
Key words: Biodiversity, Acari, distribution, habitat, water mites, taxonomy.
ndice
Introduccin ........................................................ 182
Lista sistemtica ................................................ 184
Comentarios ........................................................ 216
Agradecimientos .................................................. 216
Referencias ........................................................ 216
ndice alfabtico de nombres cientficos ............ 219
Introduccin
La publicacin de catlogos y elencos faunsticos pone en evidencia una clara conexin entre
taxonoma bsica y temas relacionados con la biodiversidad, constituyendo adems del importante
conocimiento de la fauna en s, una ineludible herramienta para estudios de los ecosistemas y punto de
partida para futuros relevamientos y revisiones
taxonmicas. Segn algunas estimaciones, slo 1030% del total de especies ha sido nominada.
Amrica del Sur es un continente dominado por
ecosistemas de aguas continentales; la enorme
diversidad de hbitat creada por este paisaje sostiene un alto nmero de artrpodos, incluyendo caros
acuticos, que constituyen un grupo hiperdiverso
de caros, de lo que se infiere que las investigaciones sobre esta fauna tienen por delante una tarea
enorme en la catalogacin de su biodiversidad.
El rpido crecimiento de la poblacin y la explotacin de los recursos naturales en Sudamrica conllevan una consecuente disminucin de la
biodiversidad en los ecosistemas acuticos, con
extinciones masivas causadas indirectamente por
destruccin de habitats, as como por alteraciones
profundas derivadas directamente de la actividad
humana. Este acelerado proceso de prdida de
riqueza y diversidad se hace ms crtico por la gran
escasez de sistemticos entrenados que puedan
revelar la diversidad animal y vegetal, permitiendo
la desaparicin de un gran nmero de especies an
antes de ser conocidas por la ciencia (Systematic
Agenda, 2000; Valdecasas, 2003).
Se ha publicado mucho sobre los caros acuticos de Amrica del Sur. En efecto, de acuerdo a
Besch, los caros acuticos sudamericanos eran los
ms estudiados con la excepcin de Europa hasta
los aos 60, siendo la mayora de las colecciones

estudiadas por dos acarlogos (Lundblad y Karl
Viets) durante los aos 40. Hoy, el proceso de
coleccin e identificacin y descripcin de caros
acuticos es ms lento. En los aos 80, el acarlogo estadounidense D. R. Cook public dos extensos
trabajos sobre caros acuticos neotropicales en
cuatro regiones de Amrica del Sur, relacionadas
con el corredor andino.
En los ltimos aos hidracarolgos de
Argentina, Rosso de Ferrads y Fernndez, han realizado diversas contribuciones, de Argentina en
particular, y ms recientemente se han extendido a
otros pases de Sudamrica.
Esta contribucin que se presenta aqu, insumi
aos de trabajo, en la catalogacin de los verdaderos caros acuticos en el grupo de la cohorte de
Parasitengona; se incluye tambin informacin
sobre superfamilias, familias, subfamilias, gneros,
subgneros, especies, y subespecies, autores,
fechas de descripcin y el actual estado del taxn.
Se agrega adems informacin sobre la distribucin
por regiones y hbitats conocidos para las especies.
Hasta el presente hay 6 superfamilias, que incluyen 23 familias, 118 gneros y 916 especies citadas
en 11 pases de Amrica del Sur. El grado de conocimiento vara enormemente de pas a pas con
numerosos datos de Brasil y por ejemplo, ninguno
de Guayana Francesa, muy pocos de Ecuador o la
cuenca alta del Amazonas.
De todo esto se puede inferir que an estamos
lejos de tener un conocimiento acabado de esta
fauna; segn estimaciones de algunos acarlogos,
slo se conoceran alrededor del 25% de las especies sudamericanas (Goldschmidt, 2002).
En el presente listado se actualiza el trabajo realizado por Viets en 1987, citndose slo algunos
CAROS ACUTICOS DE SUDAMRICA
183
Cv
Amaz
Cace
Yup
Pun
Par
Chaq
Csc
Aap
Fig. 1. Provincias biogeogrficas de Amrica del Sur (modificado de Morrone, 1996). Abreviaturas en el texto.
Fig. 1. Biogeographic provinces of South America (modified of Morrone, 1996). Abbreviations in the text.
184
trabajos anteriores a esta fecha y no incluidos en el

mismo y, desde luego, todas las contribuciones
posteriores. Para Unionicola (Unionicolidae) con
40 especies conocidas en Sudamrica, se sigui el
trabajo realizado por Vidrine (1996) que incluye
todas las especies del mundo, con importantes
modificaciones, particularmente en el nivel de subgneros.
Las localidades de distribucin de las especies
son reunidas en provincias biogeogrficas, siguiendo la propuesta de Morrone (1996) con algunas
modificaciones. Las mismas surgen del estado del
conocimiento fragmentario y referencias de localidades poco claras de Hydrachnidia de Sudamrica
(Rosso de Ferrads & Fernndez, 2001b). Las reas
de distribucin geogrfica consideradas se presentan a continuacin con la siguiente codificacin
(Fig. 1):
Aap:
Amaz:
Cace:
Chaq:
Csc:
Cv:
Par:
Pun:
Yup:
Andes Australes + Patagonia de Argentina.

Amazonia = Amazonas + Guayanas.
Caatinga + Cerrado Este.
Chaquea: al norte del ro Colorado de
Argentina, Sierras pampeanas de Argentina,
por el oeste y cuenca del Paran de Argentina y
Uruguay.
Chile austral + Chile central desde 30 de latitud
hacia el sur.
Caribea = zonas bajas de Colombia y
Venezuela.
Paran de Morrone.
Chile norte desde 30 de latitud + Bolivia y
Argentina oriental; por encima de 3.000 m.
Yungas + Pramo = Selvas de los Andes orientales.
A causa de la informacin incompleta y compleja sobre los hbitats referidos por los diversos
autores, debimos reunir la informacin existente en
una serie de hbitats cuya codificacin es la
siguiente:
est
fit
hh
ln
lt
lt-ln
ter
Estigoico
Fitotelma
Hbitat del hospedador
Lntico
Ltico
Ambiente remansado de un arroyo
Termales
Esta informacin sobre la distribucin y hbitats

de los hidracridos, es anotada a continuacin de
cada referencia especfica en un orden predeterminado, primeramente las provincias biogeogrficas y
luego los hbitat.
Lista sistemtica
SUPERFAMILIA HYDRACHNOIDEA Leach, 1815
FAMILIA HYDRACHNIDAE Leach, 1815
GNERO Hydrachna Mller, 1776
Hydrachna agilis Gervais, 1849 sp. inc.
Viets, 1987: 324.
Csc; lt.
Hydrachna heterophthalma Viets, 1954
Viets, 1987: 323.
Cace; lt; ln.
Hydrachna pusilla Daday, 1905
Viets, 1987: 307.
Par; ln.
Hydrachna ventrifissa Viets, 1954
Viets, 1987: 324.
Cace; ln.
Subgnero Diplohydrachna Thor, 1916
Hydrachna (Diplohydrachna) chilensis Gervais, 1849
Viets, 1987: 308.
Csc; lt.
Subgnero Rhabdohydrachna Viets, 1931
Hydrachna (Rhabdohydrachna) rectirostris Viets, 1940
Viets, 1987: 321.
Amaz; ln.
Hydrachna (Rhabdohydrachna) silvestrii Ribaga, 1902
Viets, 1987: 321.
Yup; Par; Chaq; ln.
Subgnero Scutohydrachna Viets, 1933
Hydrachna (Scutohydrachna) miliaria miliaria Berlese, 1888
Viets, 1987: 323; Rosso de Ferrads & Mattoni, 1999: 110.
Csc; Chaq, Pun; ln; lt.
Hydrachna (Scutohydrachna) miliaria constricta Lundblad,
1941
Viets, 1987: 323.
Par; Cace; ln; lt.
Hydrachna (Scutohydrachna) nonlamellata Viets, 1940
Viets, 1987: 323.
Cv; ln.
Hydrachna (Scutohydrachna) portigera Lundblad, 1941
Viets, 1987: 323; Rosso de Ferrads, 1996: 1.
Par; Chaq; ln.

SUPERFAMILIA EYLOIDEA Leach, 1815
FAMILIA LIMNOCHARIDAE Grube, 1859
GNERO Neolimnochares Lundblad, 1937
Neolimnochares breviscuta Lundblad, 1941
Viets, 1987: 544.
Par; lt.
Neolimnochares petrophila Lundblad, 1937
Viets, 1987: 544.
Par; lt.
Neolimnochares placophora Lundblad, 1937
Viets, 1987: 544.
Par; lt.
Neolimnochares pugiunculata Viets, 1954
Viets, 1987: 544.
Amaz; lt.
GNERO Rhyncholimnochares Lundblad, 1936
Subgnero Rhyncholimnochares Lundblad, 1936
Rhyncholimnochares (Rhyncholimnochares) andina Lundblad,
1953
Viets, 1987: 679.
Yup; lt.
Rhyncholimnochares (Rhyncholimnochares) expansiseta Cook,
1980
Viets, 1987: 679; Fernndez, 2003: 61.
Yup; lt.
Rhyncholimnochares (Rhyncholimnochares) gracilirostris
Lundblad, 1953
Viets, 1987: 679.
Yup; ln.
Rhyncholimnochares (Rhyncholimnochares) lamellipalpis
Lundblad, 1936
Viets, 1987: 679.
Par; lt.
Rhyncholimnochares (Rhyncholimnochares) longipalpis
Lundblad, 1953
Viets, 1987: 679.
Yup; ln; lt.
Rhyncholimnochares (Rhyncholimnochares) longiscuta
Lundblad, 1941
Viets, 1987: 679.
Par; lt.
Subgnero Paralimnochares Lundblad, 1937
Rhyncholimnochares (Paralimnochares) dipersiai (Rosso de
Ferrads, 1975)
Viets, 1987: 598; 680.
Chaq; lt.
Rhyncholimnochares (Paralimnochares) sursumhians
(Lundblad, 1937)
Viets, 1987: 680.
Par; lt; ln.
FAMILIA EYLAIDAE Leach, 1815
185
Eylais armata Ribaga, 1902
Chaq; lt; ln.
Eylais brasiliensis Viets, 1933
Viets, 1987: 239.
Chaq; ln; lt.
Eylais colpophthalma Ribaga, 1902
Viets, 1987: 240.
Chaq; ln.
Eylais columbiensis Walter, 1912
Viets, 1987: 240.
Cv; ln.
Eylais crawfordi Viets, 1955
Viets, 1987: 240.
Pun; ln.
Eylais glieschi Viets, 1933
Viets, 1987: 246.
Par; ln.
Eylais montana Ribaga, 1902
Viets, 1987: 252.
Yup; ln.
Eylais multispina multispina Ribaga, 1902
Viets, 1987: 252.
Aap; Chaq; lt.
Eylais multispina brevipalpis Ribaga, 1902
Aap; Chaq; ln.
Eylais obliquua Viets, 1933
Viets, 1987: 253.
Par; ln.
Eylais orthophthalma Ribaga, 1902
Viets, 1987: 253.
Chaq; Par; ln.
Eylais parvipons Walter, 1919
Viets, 1987: 254.
Yup; ln.
Eylais perincisa Ribaga, 1902
Aap; Chaq; ln; lt.
Eylais protendens protendens Berlese, 1888
Viets, 1987: 255; Rosso de Ferrads & Bttger, 1997: 180;
Rosso de Ferrads, 2000: 27.
Chaq; Par; ln; lt.
Eylais protendens distendens Ribaga, 1902 sp.inc.
Viets, 1987: 255.
Chaq; ln.
Eylais protendens ornatula Ribaga, 1902 sp.inc.
Viets, 1987: 255.
Yup; Par; ln.
SUPERFAMILIA HYDRYPHANTOIDEA Piersig, 1896
FAMILIA HYDRYPHANTIDAE Piersig, 1896
SUBFAMILIA HYDRYPHANTINAE Piersig, 1896
GNERO Hydryphantes Koch, 1841
Subgnero Hydryphantes Koch, 1841
GNERO Eylais Latreille, 1796

Eylais anisitsi Daday, 1905
Viets, 1987: 237.
Par; ln.
Hydryphantes (Hydryphantes) jujuyensis Nordenskild, 1904

Viets, 1987: 354; Rosso de Ferrads, 1987: 26; Cook,
1988: 7; Smit, 2002: 2; Goldschmidt & Gerecke, 2003: 109.
Pun; Yup; Aap; Csc; ln.
186
Hydryphantes (Hydryphantes) pumilus (Viets, 1954)

Viets, 1987: 356; Goldschmidt & Gerecke, 2003: 84.
Cv; Amaz; ln.
Hydryphantes (Hydryphantes) ramosus Daday, 1905
Viets, 1987: 356; Viets & Bttger, 1986: 109; Goldschmidt
& Gerecke, 2003: 109.
Par; Amaz; Chaq; ln.
Hydryphantes (Hydryphantes) thermacalis Cook, 1988
Cook, 1988: 6; Goldschmidt & Gerecke, 2003: 109.
Csc; ter.
Hydryphantes (Hydryphantes) undulatifrons (Viets, 1954)
Amaz; lt.
Subgnero Papilloporus Walter, 1935
Hydryphantes (Papilloporus) papillosus Lundblad, 1936
Cv; Chaq; Par; Cace; ln; lt.
Subgnero Polyhydryphantes Viets, 1926
Hydryphantes (Polyhydryphantes) alienus Lundblad, 1924
Viets, 1987: 362; Cook, 1988: 8; Goldschmidt & Gerecke,
2003: 109.
Yup; Csc; ln.
Hydryphantes (Polyhydryphantes) coscaroni Cook, 1980
Aap; ln.
Hydryphantes (Polyhydryphantes) pinguipalpis Viets, 1954
Chaq; ln.
SUBFAMILIA THYADINAE Viets, 1926
GNERO Notopanisus Besch, 1964
Notopanisus wetzeli Besch, 1964
2003: 110.
Asp; Csc; lt.
Neocalonyx (Neocalonyx) godeti Walter, 1919

Yup; ln; lt.
Neocalonyx (Neocalonyx) keldomus Cook, 1988
Csc; lt.
Neocalonyx (Neocalonyx) longimaxillaris Viets, 1953
2003: 96.
Yup; Csc; Pun, ln; lt.
Neocalonyx (Neocalonyx) pectunguis Lundblad, 1953
Yup; ln; lt; lt-ln.
Neocalonyx (Neocalonyx) placophorus Cook, 1988
Csc; lt.
Neocalonyx (Neocalonyx) schindowskii Besch, 1963
Viets, 1987: 543; Cook, 1988: 12; Goldschmidt &
Gerecke, 2003: 110.
Csc; lt.
Neocalonyx (Neocalonyx) tenuirostris Lundblad, 1941
Yup; lt.
Subgnero Paracalonyx Lundblad, 1944
Neocalonyx (Paracalonyx) longipalpis Lundblad, 1941
1988: 14; Goldschmidt & Gerecke, 2003: 110.
Aap; ln.
Neocalonyx (Paracalonyx) penai Besch, 1964
2003: 110.
Aap; Csc; lt.
Subgnero Otongacarus Goldschmidt & Gerecke, 2003
Neocalonyx (Otongacarus) pina Goldschmidt & Gerecke, 2003
Goldschmidt & Gerecke, 2003: 99.
Yup; lt.
SUBFAMILIA EUPATRELLINAE Viets, 1935
GNERO Eupatrella Walter, 1935
SUBFAMILIA PROTZIINAE Koenike, 1909

GNERO Protzia Piersig, 1896
Protzia salsa Goldschmidt & Gerecke, 2003
Yup; lt.
Eupatrella platano Goldschmidt & Gerecke, 2003

Yup; lt.
SUBFAMILIA WANDESIINAE Schwoerbel, 1961
GNERO Neocalonyx Walter, 1919

Subgnero Neocalonyx Walter, 1919
GNERO Wandesia Schechtel, 1912

Subgnero Partnuniella Viets, 1938
Neocalonyx (Neocalonyx) desajunos Goldschmidt & Gerecke,

2003
Yup; lt.
Neocalonyx (Neocalonyx) frijolito Goldschmidt & Gerecke, 2003
Yup; lt
Wandesia (Partnuniella) chechoi Cook, 1988

Cook, 1988: 15; W. (Pseudowandesia) chechoi Goldschmidt
& Gerecke, 2003: 111.
Csc; lt; ter.
Wandesia (Partnuniella) lethaea (Besch, 1964)
Aap; est.

Wandesia (Partnuniella) stalagmophila Lundblad, 1941
Par; lt-ln.
Wandesia (Partnuniella) thermalis (Viets, 1938)
Viets, 1987: 821; Schwoerbel 1987: 401; Goldschmidt &
Gerecke, 2003: 111.
Pun; ter.
Wandesia (Partnuniella) walteri Lundblad, 1937
Par; lt; lt-ln.
Subgnero Pseudowandesia Habeeb, 1958
Wandesia (Pseudowandesia) andiana Cook, 1980
Viets, 1987: 821; W. andina Goldschmidt & Gerecke,
2003: 111.
Yup; lt.
187
Hydrodroma peregrina peregrina (Koenike, 1905)
Viets, 1987: 344; Rosso de Ferrads; 1983: 16; Rosso de
Ferrads & Bttger, 1997: 181.
Par; Chaq; Amaz; ln; lt.
Hydrodroma peregrina robusta Viets, 1954
Viets, 1987: 344.
Amaz; ln.
Hydrodroma stalagmophila Lundblad, 1941
Viets, 1987: 344.
Par; lt.
Hydrodroma ungulata Lundblad, 1941
Viets, 1987: 345.
Par; ln.
FAMILIA RHYNCHOHYDRACARIDAE Lundblad, 1936
GNERO Euwandesia Andr & Naudo, 1962

SUBFAMILIA RHYNCHOHYDRACARINAE Lundblad, 1936
Euwandesia sensitiva Andr & Naudo, 1962

Viets, 1987: 235; Cook, 1988: 17; Goldschmidt &
Gerecke, 2003: 111.
Aap; Csc; lt; est.
SUBFAMILIA ANKELOTHYADINAE Besch, 1964
GNERO Ankelothyas Besch, 1964
Ankelothyas emydoides Besch, 1964
2003: 111.
Csc; lt.
GNERO Rhynchohydracarus Lundblad, 1936

Rhynchohydracarus dividuus Lundblad, 1941
Viets, 1987: 679.
Par; lt.
Rhynchohydracarus testudo Lundblad, 1936
Viets, 1987: 679.
Par; ln.
SUBFAMILIA CLATHROSPERCHONTINAE Lundblad, 1936
GNERO Clathrosperchon Lundblad, 1936

FAMILIA HYDRODROMIDAE Viets, 1936
GNERO Hydrodroma Koch, 1837
Hydrodroma despiciens despiciens (Mller, 1776)
1988: 17.
Chaq; Csc; lt; ln.
Hydrodroma despiciens brevirostris Lundblad, 1941
Viets, 1987: 342; Viets & Bttger, 1986: 109.
Par; lt.
Hydrodroma despiciens clavipes Lundblad, 1953
Viets, 1987: 342.
Yup; lt.
Hydrodroma despiciens crassiseta Viets, 1954
Viets, 1987: 342.
Amaz; lt; ln.
Hydrodroma despiciens longiseta Viets, 1954
Viets, 1987: 342.
Amaz; lt; ln.
Hydrodroma despiciens macronyx Viets, 1954
Viets, 1987: 342.
Amaz; lt; ln.
Hydrodroma despiciens micronyx Viets, 1954
Viets, 1987: 342.
Amaz; lt; ln.
Clathrosperchon crassipalpis Lundblad, 1936

Viets, 1987: 211.
Par; lt; lt-ln.
Clathrosperchon minor Lundblad, 1937
Viets, 1987: 211.
Par; lt; lt-ln.
Clathrosperchon punctatus Cook, 1980
Viets, 1987: 211; Rosso de Ferrads, 1984: 129; Rosso de
Ferrads, 2000: 29; Viets & Bttger, 1986: 109; Fernndez,
2003: 61.
Yup; Chaq; Par; lt.
Clathrosperchon transversus Viets, 1977
Viets, 1987: 211; Clathrospercon transversus Gruia, 1988: 21.
Cv; lt.
GNERO Clathrosperchonella Lundblad, 1937
Clathrosperchonella asterifera Lundblad, 1937
Viets, 1987: 211.
Par; lt; ln.
Clathrosperchonella rutae Lundblad, 1938
Viets, 1987: 211.
Par; lt.
188

FAMILIA THERMACARIDAE Sokolow, 1927
GNERO Thermacarus Sokolow, 1927

Thermacarus andinus Martn & Schwoerbel, 2002
Martin & Schwoerbel, 2002: 68.
Pun; ln, ter.
Thermacarus nevadensis Marshall, 1928
Viets, 1987: 729; Schwoerbel, 1987: 403
Pun; ter.
SUPERFAMILIA LEBERTIOIDEA Thor, 1900
FAMILIA SPERCHONTIDAE Thor, 1900
SUBFAMILIA SPERCHONTINAE Thor, 1900
GNERO Notosperchonopsis Besch, 1964
Notosperchonopsis beschi Cook, 1980
Viets, 1987: 579.
Aap; lt.
Notosperchonopsis crassipalpis Besch, 1964
Viets, 1987: 579; Cook, 1988: 21.
Csc; lt.
Notosperchonopsis pauciscutata pauciscutata (Viets, 1953)
Viets, 1987: 579.
Pun; ln.
Notosperchonopsis pauciscutata neuquenensis Cook, 1980
Viets, 1987: 579; Cook, 1988: 20; Smit, 2002: 3.
Aap; Csc; lt.
GNERO Illiesiella Besch, 1964
Illiesiella cataphracta Cook, 1988
Cook, 1988: 26.
Csc; lt.
Illiesiella circularis Besch, 1964
Viets, 1987: 580; Cook, 1988: 22.
Csc; lt.
Illiesiella ischiotricha (Besch, 1964)
Viets, 1987: 579; Cook, 1988: 27.
Aap; Csc; lt.
Illiesiella maceripalpis (Besch, 1964)
Viets, 1987: 579; Cook, 1988: 24.
Csc; lt.
Illiesiella multiscutata (Besch, 1964)
Viets, 1987: 579; Cook, 1988: 24.
Aap; Csc; lt.
Illiesiella weberi (Besch, 1964)
Viets, 1987: 580; Cook, 1988: 25.
Aap; Csc; lt.
Subgnero Mixosperchon Viets, 1926

Sperchon (Mixosperchon) andinus Lundblad, 1953
Viets, 1987: 686.
Yup; lt.
Sperchon (Mixosperchon) motasi Lundblad, 1953
Viets, 1987: 699.
Yup; lt; lt-ln.
Sperchon (Mixosperchon) neotropicus Cook, 1980
Chaq; lt.
SUBFAMILIA APELTOSPERCHONTINAE Cook, 1974
GNERO Apeltosperchon Besch, 1964
Apeltosperchon schmitzi Besch, 1964
Viets, 1987: 29; Cook, 1988: 28.
Csc; lt.
FAMILIA ANISITSIELLIDAE Koenike, 1910
SUBFAMILIA ANISITSIELLINAE Koenike, 1910
GNERO Anisitsiella Daday, 1905
Anisitsiella aculeata Daday, 1905
Viets, 1987: 28.
Par; ln.
GNERO Anisitsiellides Lundblad, 1941
Anisitsiellides australis Smit, 2002
Smit, 2002: 3.
Aap; lt.
Anisitsiellides chilensis Cook, 1988
Cook, 1988: 31.
Csc; lt.
Anisitsiellides lundbladi Cook, 1980
Viets, 1987: 28.
Asp; lt.
Anisitsiellides monticolus Lundblad, 1941
Viets, 1987: 28; Cook, 1988: 30.
Yup; lt.
GNERO Mamersellides Lundblad, 1937
Mamersellides ventriperforatus Lundblad, 1937
Viets, 1987: 506; Viets & Bttger, 1986: 111; Rosso de
Ferrads & Bttger 1997: 181.
Par; Yup; Chaq; ln; lt.
GNERO Sperchon Kramer, 1877
GNERO Rutacarus Lundblad, 1937

Subgnero Rutacarus Lundblad, 1937
Sperchon loaensis Nom. nud.

Schwoerbel, 1987: 402
Pun, lt.
Rutacarus (Rutacarus) pyriformis Lundblad, 1937

Viets, 1987: 682.
Par; lt.
Subgnero Hispidosperchon Thor, 1901
Subgnero Eorutacarus Cook, 1980
Sperchon (Hispidosperchon) brasiliensis Lundblad, 1941

Viets, 1987: 687.
Par; Cace; lt.
Rutacarus (Eorutacarus) ferradasae Cook, 1980

Viets, 1987: 682; Fernndez & Palacios 1989: 234.
Yup; lt.

Subgnero Neorutacarus Orghidan & Gruia, 1983
Rutacarus (Neorutacarus) angelieri Orghidan & Gruia, 1983
Viets, 1987: 1006.
Cv; est.
GNERO Sigthoriella Besch, 1964
Sigthoriella hygropetrica Besch, 1964
Viets, 1987: 685.
Csc; est.
SUBFAMILIA NILOTONIINAE Viets, 1929
GNERO Nilotonia Thor, 1905
Subgnero Bolivartonia Orghidan & Gruia, 1983
Nilotonia (Bolivartonia) simoni Orghidan & Gruia, 1983
Viets, 1987: 1001; Orghidan & Gruia: 104.
Cv; est.
Subgnero Mamersonia Viets, 1954
Nilotonia (Mamersonia) amazonica (Viets, 1954)
Viets, 1987: 577.
Amaz; ln.
FAMILIA OXIDAE Viets, 1926
GNERO Oxus Kramer, 1877
Oxus acutirostris Viets, 1937
Viets, 1987: 584.
Par; Cace; Amaz; lt; ln.
Oxus chilensis Cook, 1988
Cook, 1988: 50.
Csc; lt.
Oxus crassipes (Lundblad, 1936)
Viets, 1987: 585.
Par; Amaz; lt; ln.
Oxus gracilipalpis gracilipalpis (Lundblad, 1941)
Viets, 1987: 585.
Par; Amaz; Cace; lt; ln.
Oxus gracilipalpis aequalipilis Viets, 1954
Viets, 1987: 586.
Cace; ln.
Oxus schubarti Viets, 1954
Viets, 1987: 588.
Cace; lt.
Oxus valdiviensis Besch, 1964
Viets, 1987: 589; Cook, 1988: 49.
Csc; ln.
GNERO Frontipoda Koenike, 1891
Subgnero Frontipoda Koenike, 1891
Frontipoda (Frontipoda) patagonica (Lundblad, 1941)
Viets, 1987: 289; Rosso de Ferrads, 1987: 30; Cook, 1988:
47; Fernndez & Rosso de Ferrads 2001: 31; Smit, 2002: 6.
Csc; Asp; ln.
189
Frontipoda (Frontipoda) trispinosa Lundblad, 1930
Viets, 1987: 289.
Amaz; lt.
Frontipoda (Frontipoda) vicina Cook, 1988
Cook, 1988: 48.
Csc; lt.
Frontipoda (Frontipoda) virescens Lundblad, 1936
Viets, 1987: 289.
Par; Amaz; lt; ln.
Subgnero Flabellifrontipoda Lundblad, 1947
Frontipoda (Flabellifrontipoda) allarka (Cook, 1988)
Cook, 1988: 36; Smit, 2002: 6.
Csc; lt.
Frontipoda (Flabellifrontipoda) allasa (Cook, 1988)
Cook, 1988: 39; Smit, 2002: 7.
Csc; Aap; ln; lt.
Frontipoda (Flabellifrontipoda) alta (Cook, 1980)
Viets, 1987: 290; Cook, 1988: 43; Smit, 2002: 6.
Aap; lt.
Frontipoda (Flabellifrontipoda) aphanoraphtha (Besch, 1964)
Viets, 1987: 290; Cook, 1988: 42; Smit, 2002: 6.
Csc; lt.
Frontipoda (Flabellifrontipoda) ciliata Walter, 1919
Viets, 1987: 290; Cook, 1988: 45; Smit, 2002: 6.
Yup; ln.
Frontipoda (Flabellifrontipoda) dolichopalpis (Cook, 1988)
Cook, 1988: 40; Smit, 2002: 6.
Csc; lt.
Frontipoda (Flabellifrontipoda) neotropica (Lundblad, 1953)
Viets, 1987: 290; Cook, 1988: 44; Fernndez & Rosso de
Ferrads, 2000: 31; Smit, 2002: 7.
Aap; Csc; Yup, Pun; ln, lt; lt-ln.
Frontipoda (Flabellifrontipoda) parva (Cook, 1980)
Viets, 1987: 290; Cook, 1988: 38; Fernndez, 1998: 411;
Smit, 2002: 6; Fernndez, 2003: 61.
Yup; lt.
Frontipoda (Flabellifrontipoda) penai (Cook, 1988)
Cook, 1988: 35; Smit, 2002: 6.
Csc; lt.
Frontipoda (Flabellifrontipoda) polyplacophora (Cook, 1980)
Viets, 1987: 290; Cook, 1988: 46; Smit, 2002: 6.
Aap; Csc; lt.
Frontipoda (Flabellifrontipoda) sneiderni (Lundblad, 1953)
Viets, 1987: 290; Cook, 1988: 37; Smit; 2002: 6.
Yup; Csc; lt.
Frontipoda (Flabellifrontipoda) tranasa (Cook, 1988)
Cook, 1988: 43; Smit, 2002: 6.
Csc; lt.
Frontipoda (Flabellifrontipoda) unoka (Cook, 1988)
Cook, 1988: 41; Smit, 2002: 6.
Csc; lt.
FAMILIA TORRENTICOLIDAE Piersig, 1902
SUBFAMILIA NEOATRACTIDINAE Lundbld, 1941
GNERO Neoatractides Lundblad, 1941
Neoatractides formosus Lundblad, 1941
Viets, 1987: 539.
Yup; lt.
190
Neoatractides inachus Lundblad, 1941

Yup; Chaq; lt.
SUBFAMILIA TORRENTICOLINAE Piersig, 1902
Neomamersa uncipalpis Lundblad, 1953

Viets, 1987: 546.
Yup; lt.
Neomamersa willinki Cook, 1980
Viets, 1987: 546, Fernndez, 2001: 27; Fernndez, 2003: 61.
Yup; lt.
GNERO Torrenticola Piersig, 1896

Subgnero Torrenticola Piersig, 1896
GNERO Meramecia Cook, 1963

Subgnero Parameramecia Smith & Cook, 1994
Torrenticola (Torrenticola) columbiana (Lundblad, 1941)

Viets, 1987: 757; Rosso de Ferrads & Mattoni, 1999: 112;
Fernndez 2003: 61; Rosso de Ferrads; Fernndez & Rocabado,
2004: 308.
Yup; Chaq; lt; lt-ln.
Torrenticola (Torrenticola) conirostris (Lundblad, 1941)
Viets, 1987: 757.
Yup; lt.
Meramecia (Parameramecia) saltensis Fernndez, 2001

Fernndez, 2001: 22; Fernndez, 2002: 5.
Yup; lt.
Subgnero Heteratractides Lundblad, 1941

Torrenticola (Heteratractides) serratirostris (Lundblad, 1941)
Viets, 1987: 771.
Yup; lt.
Subgnero Submeramecia Cook, 1980

Meramecia (Submeramecia) diamphida Cook, 1980
Fernndez, 2001: 22, Fernndez, 2002: 4.
Yup; Chaq; Par; lt.
SUBFAMILIA TYRRELLINAE Koenike, 1910
Subgnero Monatractides Viets, 1926
GNERO Tyrrellia Koenike,1895
Torrenticola (Monatractides) brevis brevis (Lundblad, 1941)

Viets, 1987: 773.
Yup; lt.
Torrenticola (Monatractides) brevis clavipes Lundblad, 1953
Viets, 1987: 773.
Yup; lt.
Torrenticola (Monatractides) hesperia (Lundblad, 1941)
Viets, 1987: 775.
Yup; lt.
Tyrrellia australis Besch, 1962

Viets, 1987: 789; Cook, 1988: 61.
Csc; lt.
Tyrrellia crenophila Lundblad, 1938
Viets, 1987: 790.
Par; lt.
Tyrrellia longipes Cook, 1980
Viets, 1987: 790.
Pun, ln.
Tyrrellia noodti Besch, 1962
Viets, 1987: 790; Cook, 1988: 62.
Pun; lt.
Tyrrellia petrophila Viets, 1977
Viets, 1987: 790.
Amaz; ln.
Tyrrellia solivaga Cook, 1980
Viets, 1987: 790; Schwoerbel, 1987: 402.
Yup; Pun; lt.
SUPERFAMILIA HYGROBATOIDEA Koch, 1842

FAMILIA LIMNESIIDAE Thor, 1900
SUBFAMILIA NEOMAMERSINAE Lundblad, 1953
GNERO Neomamersa Lundblad, 1953
Neomamersa andalienensis Cook, 1988
Cook, 1988: 50.
Csc; lt.
Neomamersa apophylonga Gruia, 1988
Gruia, 1988: 22.
Cv; lt-est.
Neomamersa faceta Cook, 1980
Viets, 1987: 545.
Yup; lt.
Neomamersa falcipalpis Cook, 1980
Viets, 1987: 545; Fernndez, 2001: 23
Yup, Aap; lt.
Neomamersa mexicana Cook, 1980
Viets, 1987: 546; Fernndez, 2001: 24.
Aap; lt.
GNERO Neotyrrellia Lundblad, 1938

Neotyrrellia paucipora Nom. nud.
Schwoerbel, 1987: 402.
Pun; lt
Neotyrrellia petricola Lundblad, 1938
Par; Chaq; lt.
Neotyrrellia polypora Lundblad, 1953
Viets, 1987: 548.
Yup; lt.
Neotyrrellia recurva Lundblad, 1953
Viets, 1987: 548.
Yup; lt.

SUBFAMILIA NEOTORRENTICOLINAE Lundblad, 1936
GNERO Neotorrenticola Lundblad, 1936
Neotorrenticola bidens Lundblad, 1953
Viets, 1987: 547.
Yup; lt; lt-ln.
Neotorrenticola chorreronica Gruia, 1988
Gruia, 1988: 25.
Cv; lt-ln.
Neotorrenticola crassipes Lundblad, 1953
Viets, 1987: 547.
Yup; lt; lt-ln.
Neotorrenticola papillata Lundblad, 1953
Viets, 1987: 547.
Yup; lt.
Neotorrenticola plumipes Lundblad, 1953
Viets, 1987: 547.
Yup; lt.
Neotorrenticola sima Viets, 1953
Viets, 1987: 547.
Pun; lt.
Neotorrenticola violacea Lundblad, 1936
Viets, 1987: 547.
Par; lt.
Neotorrenticola walteri Lundblad, 1953
Viets, 1987: 547.
Yup; lt.
SUBFAMILIA PROTOLIMNESIINAE Viets, 1940
GNERO Protolimnesia Lundblad, 1927
Subgnero Protolimnesia Lundblad, 1927
Protolimnesia (Protolimnesia) longa Besch, 1963
Viets, 1987: 659; Smit, 2002: 12.
Yup: lt-est.
Protolimnesia (Protolimnesia) pampaensis Smit, 2002
Smit, 2002: 8.
Aap; lt.
Protolimnesia (Protolimnesia) setifera Cook, 1980
Viets, 1987: 659; Fernndez, 1994a: 26; Fernndez, 2003: 61.
Yup; lt.
Protolimnesia (Protolimnesia) unguiculata unguiculata (Walter,
1919)
Viets, 1987: 659.
Yup: lt.
Protolimnesia (Protolimnesia) unguiculata roseni (Lundblad,
1924)
Viets, 1987: 659.
Yup; lt.
Subgnero Protolimnesella Cook, 1980
Protolimnesia (Protolimnesella) gereckei Goldschmidt, 2004
Godschmidt, 2004: 98.
Yup; lt.
Protolimnesia (Protolimnesella) interstitialis Cook, 1980
Viets, 1987: 660; Fernndez 1987: 144; Fernndez &
Palacios 1989: 233; Fernndez, 2003: 61.
Yup; lt-est.
191
Protolimnesia (Protolimnesella) sorpresa Cook, 1980
Viets, 1987: 660; Fernndez 1994a: 24; Fernndez, 2003: 61.
Yup; lt-est.
GNERO Crenolimnesia Lundblad, 1938
Crenolimnesia placophora Lundblad, 1938
Viets, 1987: 217.
Par; lt.
GNERO Limnesides Lundblad, 1936
Limnesides epimeratus Lundblad, 1936
Viets, 1987: 495.
Par; lt.
SUBFAMILIA RHEOLIMNESIINAE Goldschmidt, 2004

GNERO Rheolimnesia Lundblad, 1953
Rheolimnesia motasi Lundblad, 1953
Viets, 1987: 677.
Yup; lt.
Rheolimnesia placibilis (Gerecke, 1995)
Gerecke, 1995: 418; Goldschmidt, 2004: 106.
Yup; lt.
Rheolimnesia tronchonii Orghidan & Gruia, 1983
Orghidan & Gruia, 1983: 104; Viets, 1987: 1005.
Cv; est.
SUBFAMILIA MIXOLIMNESIINAE Cook, 1988

GNERO Mixolimnesia Cook, 1988
Mixolimnesia aranda Cook, 1988
Cook, 1988: 54.
Csc; est.
Mixolimnesia aspida Cook, 1988
Cook, 1988: 60.
Csc; est.
Mixolimnesia beschi Cook, 1988
Cook, 1988: 56.
Csc; est.
Mixolimnesia eremita Cook, 1988
Cook, 1988: 55.
Csc; lt.
Mixolimnesia lemba Cook, 1988
Cook, 1988: 59.
Csc; est.
Mixolimnesia magnifica Cook, 1988
Cook, 1988: 58.
Csc; est.
Mixolimnesia pallida Cook, 1988
Cook, 1988: 52.
Csc; est.
Mixolimnesia ribagai Cook, 1988
Cook, 1988: 56.
Csc; est.
192

SUBFAMILIA LIMNESIINAE Thor, 1900
GNERO Limnesia Koch, 1836

Subgnero Limnesia Koch, 1836
Limnesia (Limnesia) amazonica Lundblad, 1930
Viets, 1987: 471.
Amaz; ln.
Limnesia (Limnesia) amplipora Viets, 1954
Viets, 1987: 471.
Amaz; ln.
Limnesia (Limnesia) andina Lundblad, 1924
Viets, 1987: 471.
Yup; lt.
Limnesia (Limnesia) angelieri Viets, 1954
Viets, 1987: 471.
Amaz; lt.
Limnesia (Limnesia) angulata Lundblad, 1937
Viets, 1987: 471.
Par; lt.
Limnesia (Limnesia) aymara Rosso de Ferrads, Fernndez &
Rocabado, 2004
Rosso de Ferrads, Fernndez & Rocabado, 2004: 308.
Yup: lt.
Limnesia (Limnesia) besselingi Viets, 1954
Viets, 1987: 472.
Amaz; lt.
Limnesia (Limnesia) brauni Viets, 1954
Viets, 1987: 472.
Amaz; lt; ln.
Limnesia (Limnesia) braytola Cook, 1980
Yup; Chaq; lt.
Limnesia (Limnesia) chilensis Cook, 1988
Cook, 1988: 64.
Csc; lt.
Limnesia (Limnesia) circumcincta Viets, 1936
Viets, 1987: 473.
Cace; Par; lt; ln.
Limnesia (Limnesia) crassiseta Lundblad, 1937
Par; Chaq; lt.
Limnesia (Limnesia) dentipalpis Lundblad, 1941
Cace; Chaq; lt; ln.
Limnesia (Limnesia) dorsalis Lundblad, 1936
Viets, 1987: 474.
Limnesia (Limnesia) dubiosa Daday, 1905
Viets, 1987: 475.
Par; ln.
Limnesia (Limnesia) exigua Walter, 1919
Viets, 1987: 475.
Amaz; lt.
Limnesia (Limnesia) falsificata Lundblad, 1936
Viets, 1987: 475.
Par; Cace; lt.
Limnesia (Limnesia) flosculipora Viets, 1954
Viets, 1987: 475.
Amaz; ln.
Limnesia (Limnesia) fuhrmanni Walter, 1912
Viets, 1987: 475.
Yup; Par; Cace; Chaq; lt; ln.
Limnesia (Limnesia) gessneri Viets, 1956

Viets, 1987: 476.
Cv; lt.
Limnesia (Limnesia) granuligera Viets, 1954
Viets, 1987: 477.
Cace; ln.
Limnesia (Limnesia) hesperia hesperia Lundblad, 1930
Viets, 1987: 477.
Yup; Par; Cace; Amaz; ln, lt.
Limnesia (Limnesia) hesperia porulosa Viets, 1954
Viets, 1987: 477.
Amaz; ln.
Limnesia (Limnesia) hexagona Lundblad, 1938
Viets, 1987: 477.
Par; Cv; ln.
Limnesia (Limnesia) laeta Stoll, 1887
Viets, 1987: 479; Rosso de Ferrads & Bttger, 1997: 181.
Yup; Par; Amaz; ln.
Limnesia (Limnesia) latigenitalis Lundblad, 1937
Par; ln.
Limnesia (Limnesia) longipora Lundblad, 1953
Viets, 1987: 480.
Yup; lt.
Limnesia (Limnesia) longirostris Lundblad, 1941
Viets, 1987: 480.
Par; lt.
Limnesia (Limnesia) longivalvata Viets, 1953
Viets, 1987: 480.
Pun; lt.
Limnesia (Limnesia) minuscula minuscula Ribaga, 1902
Viets, 1987: 483; Cook, 1988: 63; Rosso de Ferrads,
Fernndez & Rocabado, 2004: 309.
Chaq; Par; Amaz; Pun; Csc; ln; lt.
Limnesia (Limnesia) minuscula distorta Viets, 1954
Viets, 1987: 483; Rosso de Ferrads, Fernndez & Rocabado,
2004: 309.
Cace; ln.
Limnesia (Limnesia) minuscula valida Lundblad, 1941
2004: 309.
Par; lt.
Limnesia (Limnesia) minutidentata Viets, 1954
Viets, 1987: 483.
Amaz; ln.
Limnesia (Limnesia) onophora Lundblad, 1944
Viets, 1987: 484.
Yup; lt.
Limnesia (Limnesia) parva Daday, 1905
Viets, 1987: 484.
Par; ln.
Limnesia (Limnesia) patagonica Lundblad, 1941
Viets, 1987: 484; Rosso de Ferrads, Kaisin & Bosnia, 1987:
119; Balseiro, 1992: 1267; Rosso de Ferrads, Fernndez &
Rocabado, 2004: 309.
Aap; ln.
Limnesia (Limnesia) paucipora Lundblad, 1937
Viets, 1987: 485.
Par; lt.
Limnesia (Limnesia) pauciseta pauciseta Ribaga, 1902
Viets, 1987: 485; Schwoerbel, 1987: 402.
Pun; Chaq; Yup; Par; Amaz; Cv; lt; ln.

Limnesia (Limnesia) pauciseta remotipora Viets, 1953
Viets, 1987: 485.
Yup; ln.
Limnesia (Limnesia) praedentata Viets, 1954
Viets, 1987: 485.
Amaz; ln; lt.
Limnesia (Limnesia) reducta Lundblad, 1930
Viets, 1987: 485.
Yup; Chaq; lt; ln.
Limnesia (Limnesia) ribagai Lundblad, 1937
Viets, 1987: 486.
Par; Cace; lt; ln.
Limnesia (Limnesia) semireticulata Viets, 1954
Viets, 1987: 486.
Amaz; ln.
Limnesia (Limnesia) serratipes Viets, 1954
Viets, 1987: 486.
Cace; lt.
Limnesia (Limnesia) slanopa Cook, 1980
Viets, 1987: 487.
Yup; lt.
Limnesia (Limnesia) sutava Cook, 1980
Viets, 1987: 487.
Yup; lt.
Limnesia (Limnesia) tenuicoxalis Viets, 1954
Viets, 1987: 487.
Amaz; ln.
Limnesia (Limnesia) tucumanensis Cook, 1980
Viets, 1987: 488.
Yup; lt.
Subgnero Allolimnesia Viets, 1936
Limnesia (Allolimnesia) angustipalpis Lundblad, 1938
Viets, 1987: 491.
Par; Amaz; ln.
Limnesia (Allolimnesia) bidentata Viets, 1954
Viets, 1987: 491.
Cace; lt.
Limnesia (Allolimnesia) perpusilla Viets, 1956
Viets, 1987: 491
Cv; ln.
Limnesia (Allolimnesia) polypora Viets, 1936
Par; Cace; ln.
Subgnero Limnesiella Daday, 1905
Limnesia (Limnesiella) duricoria Lundblad, 1937
Par; ln.
Limnesia (Limnesiella) loretoensis Rosso de Ferrads & Smit,
1998
Rosso de Ferrads & Smit, 1998: 73.
Chaq; ln.
Limnesia (Limnesiella) malacoderma Lundblad, 1937
Viets, 1987: 492.
Par; Chaq; Amaz; ln.
Limnesia (Limnesiella) plaumanni Lundblad, 1936
Viets, 1987: 492.
Par; Amaz; ln.
Limnesia (Limnesiella) pusilla (Daday, 1905)
Viets, 1987: 493.
Par; Amaz; ln.
193
Subgnero Limnesiellula Viets, 1935
Limnesia (Limnesiellula) brasiliana (Viets, 1935)
Viets, 1987: 493.
Cace; Amaz; lt; ln.
Limnesia (Limnesiellula) schwoerbeli Rosso de Ferrads & Smit,
1998
Chaq; ln.
Subgnero Limnesiopsides Viets, 1938
Limnesia (Limnesiopsides) pectungulata Viets, 1938
Viets, 1987: 493.
Chaq; lt.
Subgnero Paralimnesia Lundblad, 1938
Limnesia (Paralimnesia) microdon Lundblad, 1938
Viets, 1987: 493.
Par; lt.
Subgnero Pilolimnesia Viets, 1938
Limnesia (Pilolimnesia) amazonicola Viets, 1956
Viets, 1987: 471; 493.
Par; Amaz; ln; lt.
Limnesia (Pilolimnesia) rostrata (Viets, 1938)
Viets, 1987 : 493.
Cace; lt.
Subgnero Tetralimnesia Thor, 1922
Limnesia (Tetralimnesia) acanthoscelus Lundblad, 1936
Viets, 1987: 494.
Par; lt.
Limnesia (Tetralimnesia) sinuosa Viets, 1954
Viets, 1987: 495.
Amaz; lt.
Subgnero Seppiella Besch, 1969.
Limnesia (Seppiella) magnipora Besch, 1969
Viets, 1987: 494.
Amaz; lt.
Limnesia (Seppiella) surinamensis Besseling, 1949
Viets, 1987: 494.
Amaz; lt.
GNERO Acantholimnesia Viets, 1954
Acantholimnesia millepora Viets, 1954
Viets, 1987: 14.
Amaz; lt.
GNERO Centrolimnesia Lundbld, 1935
Centrolimnesia bondi Lundblad, 1935
Viets, 1987: 208.
Amaz; lt.
194
Centrolimnesia boopis Gerecke, Fisher-Hartig & SteinitzKannan, 1996

Gerecke Fisher-Hartig & Steinitz-Kannan, 1996: 195.
Yup; ln.
Centrolimnesia geijskesi Besseling, 1949
Viets, 1987: 208.
Amaz; ln.
Centrolimnesia geniculata Lundblad, 1938
Viets, 1987: 208.
Par; lt.
Centrolimnesia guarani Rosso de Ferrads & Bttger, 1997
Rosso de Ferrads & Bttger, 1997: 184.
Par; ln.
Centrolimnesia lundbladi Viets, 1954
Viets, 1987: 208.
Amaz; ln.
Centrolimnesia schadei Lundblad, 1938
Par; Yup; ln.
Centrolimnesia schubarti Viets, 1938
Viets, 1987: 208.
Par; Cace; Amaz; ln.
Centrolimnesia vietsi Lundblad, 1941
Viets, 1987: 208.
Par; ln.
GNERO Tubophorella Viets, 1978
Tubophorella multiacetabulata Fernndez, 1992
Fernndez, 1992: 254.
Yup; lt.
FAMILIA OMARTACARIDAE Cook, 1963
GNERO Omartacarus Cook, 1963
Omartacarus ferradasae Cook, 1988
Cook, 1988: 65.
Csc; est.
Omartacarus paraelongatus Fernndez & Grosso, 1991
Fernndez & Grosso: 1991: 43.
Aap; est.
Omartacarus tucumanensis Fernndez, 1987
Fernndez, 1987: 395; Fernndez & Grosso, 1991: 43; Fernndez, 1993: 112; 2002: 2.
Yup, Aap, est.
FAMILIA HYGROBATIDAE Koch, 1842
SUBFAMILIA HYGROBATINAE Koch, 1842
GNERO Hygrobates Koch, 1837
Subgnero Hygrobates Koch, 1837
Hygrobates (Hygrobates) ampliatus ampliatus Viets, 1936
Viets, 1987: 367; Viets & Bttger, 1986: 111; Rosso de
Ferrads, 1987: 31; 2000: 33; Smit, 2002: 17.
Amaz; Par; Yup; Chaq; Aap; lt; ln.
Hygrobates (Hygrobates) ampliatus interpositus Viets, 1954
Viets, 1987: 368.
Cace; lt.
Hygrobates (Hygrobates) ampliatus productus Viets, 1936

Viets, 1987: 368.
Cace; lt.
Hygrobates (Hygrobates) amplipalpis Lundblad, 1953
Viets, 1987: 368.
Yup; lt.
Hygrobates (Hygrobates) apertus apertus Viets, 1936
Viets, 1987: 368.
Amaz; Par; lt; ln.
Hygrobates (Hygrobates) apertus apertulus Viets, 1954
Viets, 1987: 368.
Amaz; ln; lt.
Hygrobates (Hygrobates) clevamus Cook, 1980
Viets, 1987: 370.
Yup; lt.
Hygrobates (Hygrobates) crassipes Lundblad, 1937
Viets, 1987: 370.
Par; lt.
Hygrobates (Hygrobates) curvipalpis Besseling, 1949
Viets, 1987: 370.
Amaz; lt.
Hygrobates (Hygrobates) discrepans Lundblad, 1937
Viets, 1987: 371.
Par; lt.
Hygrobates (Hygrobates) diversidentatus Viets, 1954
Viets, 1987: 371.
Amaz; lt.
Hygrobates (Hygrobates) gracilidens Lundblad, 1936
Viets, 1987: 374.
Par; lt; ln.
Hygrobates (Hygrobates) obtusidens Lundblad, 1953
Viets, 1987: 380.
Yup; lt.
Hygrobates (Hygrobates) oxyrhynchus Viets, 1954
Viets, 1987: 380.
Cace; ln.
Hygrobates (Hygrobates) plaumanni Lundblad, 1936
Viets, 1987: 381.
Par; Cace; lt.
Hygrobates (Hygrobates) plebejus plebejus Lundblad, 1930
Fernndez, 2003: 61.
Yup; Chaq; lt.
Hygrobates (Hygrobates) plebejus tamboensis Lundblad, 1953
Viets, 1987: 381.
Yup: lt.
Hygrobates (Hygrobates) plicatus Lundblad, 1942
Viets, 1987: 381.
Par; lt.
Hygrobates (Hygrobates) porosus Lundblad, 1938
Viets, 1987: 381.
Par; lt.
Hygrobates (Hygrobates) procursus Viets, 1936
Viets, 1987: 381.
Par; Cace; lt.
Hygrobates (Hygrobates) sterrodermus Lundblad, 1953
Viets, 1987: 383.
Yup; lt.
Hygrobates (Hygrobates) triangularis triangularis Lundblad,
1937
Viets, 1987: 384.
Par; Chaq; lt.

Hygrobates (Hygrobates) triangularis lundbladi Viets, 1938
Viets, 1987: 384.
Cace; lt.
Subgnero Hygrobatides Lundblad, 1936
Hygrobates (Hygrobatides) angustiporus Viets, 1954
Viets, 1987: 386.
Amaz; lt.
Hygrobates (Hygrobatides) distendens Lundblad, 1953
Viets, 1987: 386.
Yup; lt.
Hygrobates (Hygrobatides) longimanus Lundblad, 1942
Viets, 1987: 375.
Par; lt.
Hygrobates (Hygrobatides) pachydermis (Lundblad, 1936)
Viets, 1987: 386.
Par; Amaz; lt.
Hygrobates (Hygrobatides) rufus (Lundblad, 1937)
Viets, 1987: 386.
Par; Amaz; lt; ln.
Hygrobates (Hygrobatides) transversalis Viets, 1954
Viets, 1987: 386.
Cace; ln.
Subgnero Schubartella Viets, 1937
Hygrobates (Schubartella) longipes (Viets, 1937)
Viets, 1987: 390.
Amaz; ln.
Hygrobates (Schubartella) paraensis (Besch, 1969)
Viets, 1987: 390.
Amaz; lt.
GNERO Actinacarus Lundblad, 1953
Actinacarus affinis Lundblad, 1953
Viets, 1987: 18.
Yup; lt-ln.
Actinacarus cardioporus Lundblad, 1953
Viets, 1987: 18.
Yup; lt-ln.
GNERO Aspidiobates Lundblad, 1941
Aspidiobates harveyi Cook, 1988
Cook, 1988: 93.
Csc; lt.
GNERO Atractidella Lundblad, 1936
Atractidella atractidellides (Besch, 1965)
Viets, 1987: 126.
Amaz; lt.
Atractidella biscutata Lundblad, 1937
Viets, 1987: 126.
Par; Cace; lt.
Atractidella coriacea Lundblad, 1942
Viets, 1987: 126.
Par; lt.
Atractidella hamata Lundblad, 1936
Viets, 1987: 126.
Par; lt; ln.
195
Atractidella longidens Lundblad, 1941
Viets, 1987: 126.
Yup; lt.
Atractidella magna Lundblad, 1936
Viets, 1987: 126.
Par; lt; ln.
Atractidella magniacetabulae Orghidan & Gruia, 1983
Viets, 1987: 987.
Cv; lt.
Atractidella obtusidens Lundblad, 1953
Viets, 1987: 126.
Yup; lt.
Atractidella porophora Viets, 1953
Fernndez, 2003: 61.
Yup; Chaq; lt.
Atractidella thermophila Cook, 1980
Viets, 1987: 126.
Yup; ter.
GNERO Atractides Koch, 1837
Subgnero Atractides Koch, 1837
Atractides (Atractides) brasiliensis (Lundblad, 1937)
Par; Yup; Chaq; lt; lt-ln.
Atractides (Atractides) plaumanni plaumanni Lundblad, 1953
Viets, 1987: 150.
Yup, lt; lt-ln.
Atractides (Atractides) plaumanni novus Lundblad, 1953
Viets, 1987: 150.
Yup, lt.
Atractides (Atractides) porosus porosus (Lundblad, 1936)
Viets, 1987: 150.
Par; lt.
Atractides (Atractides) porosus columbianus Lundblad, 1953
Viets, 1987: 151.
Yup; lt.
Atractides (Atractides) radilofus Cook, 1980
Viets, 1987: 152.
Yup; lt.
Atractides (Atractides) rostratus Lundblad, 1953
Viets, 1987: 153.
Yup; lt.
Atractides (Atractides) schadei (Lundblad, 1942)
Viets, 1987: 153.
Par; Yup; lt.
Atractides (Atractides) sinuatipes Lundblad, 1953
Viets, 1987: 154; Fernndez, 2003: 61.
Yup; lt.
Atractides (Atractides) validipalpis validipalpis (Lundblad, 1937)
Viets, 1987: 160.
Par; lt; ln.
Atractides (Atractides) validipalpis crassipes (Lundblad, 1942)
Viets, 1987: 160.
Par; lt.
Atractides (Atractides) zoldomus Cook, 1980
Viets, 1987: 161.
Yup; lt.
GNERO Australiobatella Lundblad, 1953
Australiobatella vietsi Lundblad, 1953
Viets, 1987: 177.
Yup; lt; lt-ln.
196
GNERO Australiobates Lundblad, 1941
GNERO Callumobates Cook, 1988
Australiobates cekalovici Besch, 1964

Viets, 1987: 177; Cook, 1988: 66; Smit, 2002: 14.
Aap; Csc; lt.
Australiobates curtipalpis Cook, 1988
Cook, 1988: 67.
Csc; lt.
Australiobates gomorus Cook, 1988
Cook, 1988: 67.
Csc; lt.
Australiobates klaasseni Besch, 1964
Viets, 1987: 178; Cook, 1988: 69.
Aap; Csc; lt; ln.
Australiobates litatus Cook, 1988
Cook, 1988: 68.
Csc; lt.
Australiobates ogavus Cook, 1988
Cook, 1988: 71.
Csc; lt.
Australiobates vantermus Cook, 1988
Cook, 1988: 69.
Csc; lt.
Callumobates kurtvietsi Cook, 1988

Cook, 1988: 85.
Csc; lt.
GNERO Brevaturus Schwoerbel, 1986

Subgnero Brevaturus Schwoerbel, 1986
Brevaturus (Brevaturus) chilensis (Schwoerbel, 1986)
Viets, 1987: 1005; Cook, 1988: 133.
Csc; lt.
Brevaturus (Brevaturus) geometricus Cook, 1988
Cook, 1988: 137.
Csc; lt.
Brevaturus (Brevaturus) gledhilli Cook, 1988
Cook, 1988: 138.
Csc; lt.
Brevaturus (Brevaturus) gonseri (Schwoerbel, 1986)
Viets, 1987: 1005; Cook, 1988: 136.
Csc; lt.
Brevaturus (Brevaturus) schwoerbeli Cook, 1988
Cook, 1988: 134.
Csc; lt.
Brevaturus (Brevaturus) stellatus (Schwoerbel, 1986)
Viets, 1987: 1006; Cook, 1988: 132.
Csc; lt.
Brevaturus (Brevaturus) triangularis Cook, 1988
Cook, 1988: 135.
Csc; lt.
Subgnero Paraspidiobates Schwoerbel, 1986
Brevaturus (Paraspidiobates) lembus Cook, 1988
Cook, 1988: 139.
Csc; lt.
Brevaturus (Paraspidiobates) similis Cook, 1988
Cook, 1988: 141.
Csc; lt.
Brevaturus (Paraspidiobates) wetzlari (Schwoerbel, 1986)
Viets, 1987: 1003; Cook, 1988; 142.
Csc; lt.
GNERO Camposea Schwoerbel, 1986

Camposea phreaticola Schwoerbel, 1986
Viets, 1987: 992; Cook, 1988: 92.
Csc; est.
GNERO Corticacarus Lundblad, 1936
Corticacarus incurvatus Viets, 1953
Viets, 1987: 215.
Pun, Yup; lt.
Subgnero Corticacarus Lundblad, 1936
Corticacarus (Corticacarus) coarctipalpis Lundblad, 1942
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) coeruleus Lundblad, 1942
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) columbianus Lundblad, 1941
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) crassipalpis Lundblad, l942
Viets, 1987: 213.
Par; lt; ln.
Corticacarus (Corticacarus) davecooki (Cook, 1988)
Cook, 1988: 113; Fernndez 1997: 92.
Csc; lt.
Corticacarus (Corticacarus) dentipalpis dentipalpis Lundblad,
1936
Viets, 1987: 213.
Par; lt; ln.
Corticacarus (Corticacarus) dentipalpis schadei Lundblad, 1942
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) edentulus Lundblad, 1942
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) exilis Cook, 1988
Cook, 1988: 114.
Csc; lt.
Corticacarus (Corticacarus) jemobus Cook, 1988
Cook, 1988: 120.
Csc; lt.
Corticacarus (Corticacarus) malleifer malleifer Lundblad, 1936
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) malleifer affinis Lundblad, 1942
Viets, 1987: 213.
Par; lt.
Corticacarus (Corticacarus) patagonicus Besch, 1964
Viets, 1987: 213; Cook, 1988: 118; Smit, 2002: 15.
Csc; Aap; lt.
Corticacarus (Corticacarus) placophorus Schwoerbel, 1986
Viets, 1987: 992; Cook, 1988: 110; Smit, 2002: 16.
Csc; Aap; lt.

Corticacarus (Corticacarus) pravanus Cook, 1988
Cook, 1988: 114.
Csc; lt.
Corticacarus (Corticacarus) rimorus Cook, 1988
Cook, 1988: 116.
Csc; lt.
Corticacarus (Corticacarus) rostratus Cook, 1988
Cook, 1988: 117.
Csc; lt.
Corticacarus (Corticacarus) sovus Cook, 1988
Cook, 1988: 119; Smit, 2002: 16.
Csc; Aap; lt.
Corticacarus (Corticacarus) stukabus Cook, 1988
Cook, 1988: 115.
Csc; lt.
Corticacarus (Corticacarus) umadus Cook, 1988
Cook, 1988: 111.
Csc; lt.
Subgnero Corticacarellus Lundblad, 1937
Corticacarus (Corticacarellus) acutidens (Lundblad, 1938)
Viets, 1987: 214.
Par; ln.
Corticacarus (Corticacarellus) labialis Lundblad, 1936
Viets, 1987: 214.
Par; lt; ln.
Corticacarus (Corticacarellus) latidens (Viets, 1953)
Viets, 1987: 214.
Pun; lt.
Corticacarus (Corticacarellus) vietsi Lundblad, 1953
Viets, 1987: 214.
Yup; lt-ln.
Subgnero Corticacarides Lundblad, 1941
Corticacarus (Corticacarides) nilsoni Besch, 1964
Viets, 1987: 214; Cook, 1988: 100; Smit, 2002: 14.
Csc; Aap; lt.
Corticacarus (Corticacarides) plaumanni Lundblad, 1941
Viets, 1987: 214.
Par; lt.
Subgnero Lundbladacarus Motas & Tanasachi, 1960
Corticacarus (Lundbladacarus) anchistus Cook, 1988
Cook, 1988: 106.
Csc; lt.
Corticacarus (Lundbladacarus) angelescui Motas & Tanasachi,
1960
Viets, 1987: 214.
Aap; Csc; lt.
Corticacarus (Lundbladacarus) argentinensis Cook, 1988
Cook, 1988: 108.
Aap; lt.
Corticacarus (Lundbladacarus) brundini Besch, 1964
Viets, 1987: 213; Cook, 1988: 104; Smit, 2002: 15.
Aap; Csc; lt.
Corticacarus (Lundbladacarus) chilensis Viets, 1967
Viets, 1987: 214; Cook, 1988: 101.
Csc; Asp; lt; ln.
197
Corticacarus (Lundbladacarus) divisus Cook, 1988
Cook, 1988: 102.
Csc; lt.
Corticacarus (Lundbladacarus) icotus Cook, 1988
Cook, 1988: 105.
Csc; lt.
Corticacarus (Lundbladacarus) penai Cook, 1988
Cook, 1988: 109.
Csc; lt.
Corticacarus (Lundbladacarus) schusteri Viets, 1977
Viets, 1987: 214.
Amaz; lt.
Corticacarus (Lundbladacarus) xystus Besch, 1964
Viets, 1987: 214.
Csc; lt.
Subgnero Paracorticacarus Lundblad, 1953
Corticacarus (Paracorticacarus) brassanus Cook, 1980
Yup; Chaq; lt.
Corticacarus (Paracorticacarus) brogavus Cook, 1980
Viets, 1987: 215.
Yup; lt.
Corticacarus (Paracorticacarus) coldomus Cook, 1980
Viets, 1987: 215.
Yup; lt.
Corticacarus (Paracorticacarus) copinae Rosso de Ferrads,
1991
Chaq; lt.
Corticacarus (Paracorticacarus) curvirostris Lundblad, 1953
Viets, 1987: 215.
Yup; lt; ln; lt-ln.
Corticacarus (Paracorticacarus) longirostris Lundblad, 1953
Viets, 1987: 215.
Yup; lt-ln.
Corticacarus (Paracorticacarus) minutissimus (Lundblad, 1953)
Viets, 1987: 215.
Yup; lt-ln.
Corticacarus (Paracorticacarus) motasi Lundblad, 1953
Viets, 1987: 215.
Yup; lt.
Corticacarus (Paracorticacarus) multiscutus multiscutus
Lundblad, 1953
Viets, 1987: 215.
Yup; lt; ln.
Corticacarus (Paracorticacarus) multiscutus divisus Lundblad,
1953
Viets, 1987: 216.
Yup; lt-ln.
Corticacarus (Paracorticacarus) multiscutus reductus Lundblad,
1953
Viets, 1987: 216.
Yup; lt-ln.
Corticacarus (Paracorticacarus) noodti Besch, 1964
Viets, 1987: 216; Cook, 1988: 103.
Csc; lt.
Corticacarus (Paracorticacarus) schwoerbeli Cook, 1980
Viets, 1987: 216.
Yup; lt.
198
Corticacarus (Paracorticacarus) setipes Lundblad, 1953
Viets, 1987: 216.
Yup; lt; lt-ln.
Corticacarus (Paracorticacarus) smithi Cook, 1980
Viets, 1987: 216; Fernndez, 2003: 61.
Yup; lt.
Corticacarus (Paracorticacarus) suchezae Fernndez & Rosso
de Ferrads, 2001
Fernndez & Rosso de Ferrads, 2001: 33.
Pun; lt.
Corticacarus (Paracorticacarus) testudo testudo Lundblad, 1953
Viets, 1987: 216.
Yup; lt; lt-ln.
Corticacarus (Paracorticacarus) testudo aberratus Lundblad,
1953
Viets, 1987: 216.
Yup; lt; ln; lt-ln.
Subgnero Polycorticacarellus Lundblad, 1953
Corticacarus (Polycorticacarellus) multiporus Lundblad, 1953
Viets, 1987: 216.
Yup; lt; ln; lt-ln.
Corticacarus (Polycorticacarellus) pereirai Cook, 1980
Viets, 1987: 216.
Aap; lt.
Subgnero Tetracorticacarellus Lundblad, 1941
Corticacarus (Tetracorticacarellus) scutatus Lundblad, 1941
Viets, 1987: 217.
Par; lt.
GNERO Crenohygrobates Lundblad, 1938
Crenohygrobates multiporus Lundblad, 1938
Viets, 1987: 217.
Par; ln.
GNERO Decussobates Cook, 1988
Decussobates angustus Cook, 1988
Cook, 1988: 88.
Csc; lt.
Decussobates planus Cook, 1988
Cook, 1988: 87.
Csc; lt.
GNERO Diamphidaxona Cook, 1963
Diamphidaxona guacharensis Orghidan & Gruia, 1987
Orghidan & Gruia, 1987: 129.
Cv; lt.
Diamphidaxona hiporreica Fernndez, 1988
Fernndez, 1988: 140; Fernndez & Palacios 1989: 234;
Fernndez 1991: 26.
Yup; lt-est.
Diamphidaxona separata Cook, 1980
Viets, 1987: 223.
Yup; lt-est.
Diamphidaxona spinaxa Orghidan & Gruia, 1987
Cv; lt.

Diamphidaxona yungasa Cook, 1980
Viets, 1987: 223; Fernndez, 1995: 114; Fernndez, 2003: 61
Yup; lt-est.
GNERO Dodecabates Viets, 1926
Dodecabates dodecaporus (Nordenskild, 1904)
Viets, 1987: 227; Schwoerbel, 1987: 402; Rosso de
Ferrads, 2000: 34; Smit, 2002: 18; Fernndez, 2003: 61.
Pun; Yup; Aap; Chaq; lt.
GNERO Dubiobates Cook, 1988
Dubiobates bodus Cook, 1988
Cook, 1988: 153.
Csc; lt.
Dubiobates evolus Cook, 1988
Cook, 1988: 156.
Csc; lt.
Dubiobates kolus Cook, 1988
Cook, 1988: 150.
Csc; lt.
Dubiobates minutus Cook, 1988
Cook, 1988: 147.
Csc; lt.
Dubiobates penai Cook, 1988
Cook, 1988: 145.
Csc; lt.
Dubiobates recurvus Cook, 1988
Cook, 1988: 152.
Csc; lt.
Dubiobates schwoerbeli Cook, 1988
Cook, 1988: 154.
Csc; lt.
Dubiobates securipes Cook, 1988
Cook, 1988: 146.
Csc; lt.
Dubiobates similis Cook, 1988
Cook, 1988: 149.
Csc; lt.
Dubiobates valdiviensis Cook, 1988
Cook, 1988: 151.
Csc; lt.
GNERO Eocorticacarus Besch, 1964
Eocorticacarus siolii Besch, 1964
Viets, 1987: 231.
Yup; lt.
GNERO Hygrobatella Viets, 1926
Subgnero Hygrobatella Viets, 1926
Hygrobatella (Hygrobatella) chillanensis Cook, 1988
Cook, 1988: 77.
Csc; lt.
Hygrobatella (Hygrobatella) coriacea coriacea (Viets, 1953)
Viets, 1987: 364.
Pun; lt.
Hygrobatella (Hygrobatella) coriacea quiponensis Cook, 1980
Viets, 1987: 365.
Yup; Chaq; lt.

Hygrobatella (Hygrobatella) elegantula Lundblad, 1953
Viets, 1987: 365; Cook, 1988: 79.
Yup; Csc; lt.
Hygrobatella (Hygrobatella) karinae Besch, 1964
Viets, 1987: 365; Cook, 1988: 77.
Csc; ln.
Hygrobatella (Hygrobatella) longigenitalis longigenitalis
Lundblad, 1953
Viets, 1987: 365.
Yup; lt.
Hygrobatella (Hygrobatella) longigenitalis elata Lundblad, 1953
Viets, 1987: 365.
Yup; lt; lt-ln.
Hygrobatella (Hygrobatella) neopuberula Cook, 1988
Cook, 1988: 80
Csc; lt.
Hygrobatella (Hygrobatella) papillata Lundblad, 1941
Viets, 1987: 365.
Yup; lt.
Hygrobatella (Hygrobatella) parvula Besch, 1964
Viets, 1987: 365; Cook, 1988: 76.
Csc; lt.
Hygrobatella (Hygrobatella) placophora (Walter, 1919)
Viets, 1987: 365.
Yup; lt.
Hygrobatella (Hygrobatella) puberula puberula Viets, 1935
Viets, 1987: 365; Schwoerbel, 1987: 402; Cook, 1988: 79;
Smit, 2002: 17.
Pun; Csc; Aap; lt.
Hygrobatella (Hygrobatella) puberula arcuata Lundblad, 1953
Viets, 1987: 365.
Yup; lt.
Hygrobatella (Hygrobatella) puberula coriacea (Viets, 1953)
Viets, 1987: 365.
Yup; lt; ln.
Hygrobatella (Hygrobatella) puberula minuta Lundblad, 1953
Viets, 1987: 365.
Yup; lt.
Hygrobatella (Hygrobatella) puberula montana (Lundblad, 1941)
Viets, 1987: 366.
Yup; lt.
Hygrobatella (Hygrobatella) puberula validipalpis Lundblad,
1953
Viets, 1987: 366.
Yup; lt.
Hygrobatella (Hygrobatella) valdiviensis Cook, 1988
Cook, 1988: 78
Csc; lt.
Subgnero Schwoerbelobatella Rosso de Ferrads, Fernndez
& Rocabado, 2004
Hygrobatella (Schwoerbelobatella) multiacetabulata Cook, 1980
Viets, 1987: 365; Fernndez, 1994b: 221; Fernndez, 2003:
61; Rosso de Ferrads, Fernndez & Rocabado, 2004: 311.
Yup; lt.
Hygrobatella (Schwoerbelobatella) polygramma Lundblad, 1953
2004: 311.
Yup; lt; lt-ln.
199
GNERO Kyphohygrobatella Lundblad, 1936
Kyphohygrobatella serratipalpis Lundblad, 1936
Viets, 1987: 423.
Par; lt.
GNERO Kyphohygrobates Viets, 1935
Kyphohygrobates verrucifer (Daday, 1905)
Viets, 1987: 423.
Par; lt.
GNERO Mapuchacarus Besch, 1964
Mapucharacus condiscipulorum Besch, 1964
Viets, 1987: 507; Cook, 1988: 121.
Csc; lt.
GNERO Megapella Lundblad, 1936
Megapella longimaxillaris Lundblad, 1936
Viets, 1987: 509.
Par; lt.
Megapella flabellum (Lundblad, 1941)
Viets, 1987: 509.
Par; lt.
GNERO Motasia Lundblad, 1953
Motasia placoderma Lundblad, 1953
Viets, 1987: 533.
Yup; lt; lt-ln.
GNERO Neocorticacarus Lundblad, 1953
Neocorticacarus validipalpis Lundblad, 1953
Viets, 1987: 543.
Yup; lt; lt-ln.
GNERO Osornobates Cook, 1988
Osornobates gennadus Cook, 1988
Cook, 1988: 82.
Csc; lt.
GNERO Paraschizobates Lundblad, 1937
Paraschizobates columbianus (Lundblad, 1953)
Viets, 1987: 598.
Yup; lt; lt-ln.
Paraschizobates megapoides Lundblad, 1937
Viets, 1987: 598.
Par; Amaz; lt; ln.
GNERO Polyhygrobatella Lundblad, 1953
Polyhygrobatella polypora (Lundblad, 1953)
Viets, 1987: 656.
Yup; lt.
200
GNERO Schizobates Thor, 1927
Schizobates clanopus Cook, 1988
Cook, 1988: 75.
Csc; lt.
Schizobates critus Cook, 1988
Cook, 1988: 72.
Csc; lt.
Schizobates disjunctus (Walter, 1925)
Viets, 1987: 683.
Aap; Csc; ln; lt.
Schizobates similis Cook, 1988
Cook, 1988: 74.
Csc; lt.
GNERO Stylohygrobates Viets, 1935
Stylohygrobates longipalpis Viets, 1935
Viets, 1987: 718.
Cace; ln.
GNERO Subcorticacarus Lundblad, 1937
Subcorticacarus digitatus Lundblad, 1937
Viets, 1987: 719.
Par; lt.
Subcorticacarus trispinosus Lundblad, 1942
Viets, 1987: 719.
Par; lt.
GNERO Tetrahygrobatella Lundblad, 1953
Tetrahygrobatella argentinensis Cook, 1980
Viets, 1987: 726; Fernndez, 2003: 61.
Yup; lt.
Tetrahygrobatella bovala Cook, 1980
Viets, 1987: 726; Rosso de Ferrads, 2000: 33; Fernndez
& Rosso de Ferrads, 2001: 31.
Yup; Chaq, Pun; lt.
Tetrahygrobatella chilensis (Besch, 1964)
Viets, 1987: 726; Smit, 2002: 16.
Csc; Aap; lt.
Tetrahygrobatella dictyoderma (Lundblad, 1953)
Viets, 1987: 726.
Yup; lt; lt-ln.
Tetrahygrobatella longipalpis Cook, 1980
Viets, 1987: 726.
Yup; lt.
GNERO Thoracohygrobates Lundblad, 1936
Thoracohygrobates cancellatus Lundblad, 1936
Viets, 1987: 729.
Par; lt.
GNERO Zabobates Cook, 1988
Zabobates alphus Cook, 1988
Cook, 1988: 89.
Csc; lt.
Zabobates facetus Cook, 1988
Cook, 1988: 91.
Csc; lt.

Zabobates gledhilli Cook, 1988
Cook, 1988: 90.
Csc; lt.
SUBFAMILIA RHYNCHATURINAE Schwoerbel, 1986
GNERO Rhynchaturus Besch, 1964
Rhynchaturus beschi Cook, 1988
Cook, 1988: 125.
Csc; lt.
Rhynchaturus cautinensis Cook, 1988
Cook, 1988: 127.
Csc; lt.
Rhynchaturus circularis Cook, 1988
Cook, 1988: 126.
Csc; lt.
Rhynchaturus hexaporus Besch, 1964
Viets, 1987: 678; Cook, 1988: 124.
Csc; Aap; lt.
Rhynchaturus octoporus Schwoerbel, 1986
Viets, 1987: 1005; Cook, 1988: 129.
Csc; lt.
Rhynchaturus proyectus Cook, 1988
Cook, 1988: 128.
Csc; lt.
Rhynchaturus tricornis Cook, 1988
Cook, 1988: 129.
Csc; lt.
GNERO Andesobates Smit, 2002
Andesobates longipalpis Smit, 2002
Smit, 2002: 19.
Aap; Csc; lt.
GNERO Szalayella Lundblad, 1953
Szalayella incisa Lundblad, 1953
Viets, 1987: 721.
Yup; lt.
Szalayella lundbladi Cook, 1980
Viets, 1987: 721; Cook, 1988: 84; Smit, 2002: 18.
Aap; Csc; lt.
FAMILIA FERRADASIIDAE Cook, 1980
GNERO Ferradasia Cook, 1980
Ferradasia musicola Cook, 1980
Viets, 1987: 278.
Aap; lt.
FAMILIA UNIONICOLIDAE Oudemans, 1909
SUBFAMILIA UNIONICOLINAE Oudemans, 1909
GNERO Unionicola Haldeman, 1842
Subgnero Unionicola Haldeman, 1842
Unionicola (Unionicola) iheringi (Koenike, 1890)
Viets, 1987: 797; Vidrine, 1996: 5.
Par; lt.

Unionicola (Unionicola) inermis Lundblad, 1941
Viets, 1: 797; Vidrine, 1996: 5.
Aap; lt.
Unionicola (Unionicola) levipalpis Besseling, 1949
Viets, 1987: 810; Vidrine, 1996: 5.
Amaz; lt.
Unionicola (Unionicola) motasi Viets, 1959
Viets, 1987: 811; Vidrine, 1996: 5.
Amaz; ln.
Unionicola (Unionicola) perpusilla Viets, 1954
Viets, 1987: 812; Vidrini, 1996: 5.
Amaz, ln.
Unionicola (Unionicola) pugionipalpis Viets, 1954
Viets, 1987: 812; Vidrine, 1996: 5.
Amaz; ln.
Unionicola (Unionicola) pusuligera Viets, 1954
Viets, 1987: 812; Vidrine, 1996: 5.
Cace; ln.
Unionicola (Unionicola) siolii Viets, 1954
Viets, 1987: 801; Vidrine, 1996: 5.
Amaz; lt.
Subgnero Ampullariatax Vidrine, 1985
Unionicola (Ampullariatax) ampullariae (Koenike, 1890)
Viets, 1987: 814; Vidrine, 1996: 2.
Par; Chaq, hh: Pomacea insularum; Amaz; ln; lt.
Subgnero Atacella Lundblad, 1937
Unionicola (Atacella) clathrata (Lundblad, 1937)
Viets, 1987: 118; Vidrine, 1996: 2.
Par, Chaq; hh: Anodontites obtusus lucidus; Anodontites
trapesialis spixii; Diplodon rhuacoicus; Diplodon charruanus;
lt; ln.
Unionicola (Atacella) entrerrianensis (Rosso de Ferrads, 1976)
Viets, 1987: 118; Vidrine, 1996: 2.
Chaq, hh: Anodontites trapesialis spixii; lt.
Unionicola (Atacella) fissipes (Koenike, 1891)
Viets, 1987: 118; Vidrine, 1996: 2.
Par, hh, Anodontites patagonicus; Chaq, hh: Anodontites
trapesialis susannae; lt.
Unionicola (Atacella) gigantea (Cachs & Ma-Garzn, 1973)
Viets, 1987: 118; Vidrine, 1996: 2.
Chaq; lt; ln.
Unionicola (Atacella) perforata (Koenike, 1890)
Viets, 1987: 118; Vidrine, 1996: 2.
Par, hh: Anodontites patagonicus; Chaq, hh, Anodontites
trapesialis spixii Chaq; lt.
Unionicola (Atacella) rugosa (Koenike, 1890)
Viets, 1987: 119; Vidrine, 1996: 2.
Par; lt.
Unionicola (Atacella) schubarti (Viets, 1954)
Viets, 1987: 119; Vidrine, 1996: 2.
Cace, hh: Diplodon delodontus expansus; ln; lt.
Unionicola (Atacella) subrecta (Cachs & Ma-Garzn, 1973)
Viets, 1987: 119; Vidrine, 1996: 2.
Chaq; lt.
Subgnero Australatax Vidrine, 1985
Unionicola (Australatax) sinuata Lundblad, 1938
Viets, 1987: 813; Vidrine, 1996: 2.
Par, hh: Anodontites sp; Chaq, hh: Anodontites trapesialis
susanae; lt.
201
Subgnero Crameratax Vidrine, 1988
Unionicola (Crameratax) acutidens (Lundblad, 1936)
Viets, 1987: 806; Vidrine, 1996: 3.
Par; lt.
Unionicola (Crameratax) gracilipes (Lundblad, 1936)
Viets, 1987: 809; Vidrine, 1996: 3.
Par; lt.
Unionicola (Crameratax) tumidipalpis (Lundblad, 1936)
Viets, 1987: 813; Vidrine, 1996:3.
Par; Cace; lt.
Subgnero Curryatax Vidrine, 1992
Unionicola (Curryatax) prehendens Viets, 1954
Viets, 1987: 812; Vidrine, 1996: 3.
Amaz; ln.
Subgnero Ferradasatax Vidrine, 1988
Unionicola (Ferradasatax) procurvipes (Koenike, 1890)
Viets, 1987: 812; Vidrine, 1996: 3.
Par, hh: Anodonta sp., Anodontites sp.; Chaq, hh:
Anodontites trapesialis susannae; lt.
Subgnero Gledhillatax Vidrine, 1992
Unionicola (Gledhillatax) angustipalpis (Lundblad, 1937)
Viets, 1987: 806; Vidrine, 1996: 3.
Par; lt.
Unionicola (Gledhillatax) brevisuturata Viets, 1959
Viets, 1987: 807; Vidrine, 1996: 3.
Amaz; ln.
Unionicola (Gledhillatax) longipes Lundblad, 1942
Viets, 1987: 811; Vidrine, 1996: 3.
Par; Amaz; lt.
Unionicola (Gledhillatax) simplicipes (Lundblad, 1936)
Viets, 1987: 813; Vidrine, 1996: 3.
Par; lt.
Subgnero Lasalleatax Vidrine, 1992
Unionicola (Lasalleatax) brasiliensis (Lundblad, 1936)
Viets, 1987: 807; Vidrine, 1996: 4.
Par; lt.
Unionicola (Lasalleatax) conjuncta Viets, 1954
Viets, 1987: 807; Vidrine, 1996: 4.
Cace; ln.
Unionicola (Lasalleatax) unidens Lundblad, 1942
Viets, 1987: 801; Vidrine, 1996: 4.
Par; ln.
Subgnero Lundbladatax Vidrine, 1988
Unionicola (Lundbladatax) fissipalpis Lundblad, 1942
Viets, 1987: 808; Vidrine, 1996: 4.
Par; Cace; ln; lt.
Subgnero Mitchellatax Vidrine, 1992
Unionicola (Mitchellatax) curvitarsis Lundblad, 1941
Viets, 1987: 808; Vidrine, 1996: 4.
Par; lt.
202
Unionicola (Mitchellatax) longidens Lundblad, 1942

Viets, 1987: 811; Vidrine, 1996: 4.
Par; Cace; lt.
Subgnero Polyatacides Lundblad, 1941
Unionicola (Polyatacides) prominens (Koenike, 1914)
Viets, 1987: 119 ; Vidrine, 1996: 4.
Par; lt.
Subgnero Poundsatax Vidrine, 1992
Unionicola (Poundsatax) retractidens Lundblad, 1937
Viets, 1987: 812; Vidrine, 1996: 4.
Par; lt.
Subgnero Wilsonatax Vidrine, 1992
Unionicola (Wilsonatax) plaumanni Lundblad, 1937
Viets, 1987: 812; Vidrine, 1996: 6.
Par; lt.
Unionicola (Wilsonatax) viciniseta (Lundblad, 1936)
Viets, 1987: 814; Vidrine, 1996: 6.
Par; Cace; lt.
Subgnero Unionicolides Lundblad, 1937
Unionicola (Unionicolides) bonariensis Mauri y Alzuet, 1972
Viets, 1987: 815; Viddrine, 1996: 6.
Chaq, hh, Diplodon variabilis; lt.
Unionicola (Unionicolides) sica Lundblad, 1937
Viets, 1987: 815; Vidrine, 1996: 6.
Par, hh, Anodontites sp.; Chaq, hh, Anodontites obtusus
lucidus, Anodontites crispatus tenebricusus, Anodontites trapezeus, lt.
Subgnero Unionicolella Lundblad, 1941
Unionicola (Unionicolella) pachyscelus Lundblad, 1941
Viets, 1987: 815; Vidrine, 1996: 6.
Par, hh, Anodontites sp.; Chaq, hh, Diplodon charruanus; lt.
SUBFAMILIA PIONATACINAE Viets, 1916

GNERO Amazonella Lundblad, 1930
Amazonella crassipalpis Lundblad, 1941
Viets, 1987: 26.
Par; lt.
Amazonella polypora polypora Lundblad, 1936
Viets, 1987: 26.
Par; lt; ln.
Amazonella polypora curvimarginata Viets, 1959
Viets, 1987: 26.
Amaz; lt.
Amazonella ribagai Lundblad, 1930
Viets, 1987: 26.
Par; Amaz; lt; ln.
GNERO Koenikea Wolcott, 1900

Koenikea longiuscula Lundblad, 1936
Viets, 1987: 401.
Par; lt.
Koenikea pauciseta Cook, 1980
Viets, 1987: 402.
Yup; lt.
Koenikea plaumanni Lundblad, 1937
Viets, 1987: 402.
Par; lt; ln.
Koenikea schubarti Viets, 1977
Viets, 1987: 403.
Cace; lt.
Subgnero Koenikea Wolcott, 1900
Koenikea (Koenikea) acuta acuta Lundblad, 1943
Par; ln.
Koenikea (Koenikea) acuta dentata Viets, 1975
Viets, 1987: 397.
Amaz; lt.
Koenikea (Koenikea) affinis Lundblad, 1938
Viets, 1987: 397.
Par; lt; ln.
Koenikea (Koenikea) angustipalpis Viets, 1975
Viets, 1987: 397.
Amaz; lt.
Koenikea (Koenikea) bicornis Lundblad, 1941
Viets, 1987: 398.
Par; lt.
Koenikea (Koenikea) bipapillata Lundblad, 1941
Viets, 1987: 398.
Par; Amaz; ln; lt.
Koenikea (Koenikea) biscutata Daday, 1905
Viets, 1987: 398.
Par; lt; ln.
Koenikea (Koenikea) brunnea Lundblad, 1937
Viets, 1987: 398.
Par; lt.
Koenikea (Koenikea) coerulescens coerulescens Lundblad, 1936
Viets, 1987: 398.
Par; lt; ln.
Koenikea (Koenikea) coerulescens rosea Lundblad, 1936
Viets, 1987: 398.
Par; lt.
Koenikea (Koenikea) coerulescens vietsi Lundblad, 1937
Viets, 1987: 398.
Par; Amaz; lt
Koenikea (Koenikea) convexa Daday, 1905 sp. inc.
Viets, 1987: 399.
Par; ln.
Koenikea (Koenikea) corniculata Viets, 1975
Viets, 1987: 399.
Amaz; ln.
Koenikea (Koenikea) falcaria Viets, 1977
Viets, 1987: 400.
Amaz; ln.
Koenikea (Koenikea) fluctuata Viets, 1975
Viets, 1987: 400.
Amaz; ln.

Koenikea (Koenikea) furcatimaculata Lundblad, 1936
Viets, 1987: 400.
Par; lt.
Koenikea (Koenikea) hastata Viets, 1977
Viets, 1987: 401.
Cace; ln.
Koenikea (Koenikea) lata Viets, 1975
Viets, 1987: 401.
Amaz; lt.
Koenikea (Koenikea) longipes Lundblad, 1936
Viets, 1987: 401.
Par; lt.
Koenikea (Koenikea) medioposita medioposita Lundblad, 1936
Viets, 1987: 401.
Par; lt; ln.
Koenikea (Koenikea) medioposita asterifera Lundblad, 1943
Viets, 1987: 401.
Par; lt.
Koenikea (Koenikea) medioposita transversa Lundblad, 1943
Viets, 1987: 401.
Par; lt.
Koenikea (Koenikea) octopapillata Lundblad, 1930
Viets, 1987: 402.
Par; Amaz; lt; ln.
Koenikea (Koenikea) pallida Lundblad, 1937
Viets, 1987: 402.
Par; Amaz; lt; ln.
Koenikea (Koenikea) papillata Viets, 1975
Viets, 1987: 402.
Amaz; lt.
Koenikea (Koenikea) paraguayensis Rosso de Ferrads &
Bttger, 1997
Par; ln.
Koenikea (Koenikea) plicata Lundblad, 1936
Viets, 1987: 402.
Par; lt.
Koenikea (Koenikea) procursa Viets, 1977
Viets, 1987: 403.
Amaz; lt.
Koenikea (Koenikea) quadricornuta Walter, 1919
Viets, 1987: 403.
Par; lt; ln.
Koenikea (Koenikea) rectangulata Lundblad, 1930
Viets, 1987: 403.
Par; Cace; lt.
Koenikea (Koenikea) rectirostris rectirostris Lundblad, 1936
Par; Yup; Chaq; lt.
Koenikea (Koenikea) rectirostris ramosa Lundblad, 1943
Viets, 1987: 403.
Par; lt.
Koenikea (Koenikea) retractidens retractidens Lundblad, 1938
Viets, 1987: 403.
Par; lt; ln.
Koenikea (Koenikea) retractidens crassoalata Lundblad, 1943
Viets, 1987: 403.
Par; ln; lt.
Koenikea (Koenikea) retrocornuta Rosso de Ferrads & Bttger,
1997
Par; ln.
203
Koenikea (Koenikea) siolii Viets, 1975
Viets, 1987: 403.
Amaz; lt.
Koenikea (Koenikea) triangularis Lundblad, 1938
Par; ln.
Koenikea (Koenikea) unguiculata Lundblad, 1943
Viets, 1987: 404.
Par; lt.
Koenikea (Koenikea) victoriola Rosso de Ferrads, 1998
Chaq; ln.
Subgnero Diplokoenikea Lundblad, 1936
Koenikea (Diplokoenikea) clavigera clavigera Lundblad, 1943
Viets, 1987: 405; Cook, 1988: 159.
Par; Csc; lt.
Koenikea (Diplokoenikea) clavigera assimilis Lundblad, 1943
Viets, 1987: 406.
Par; lt; ln.
Koenikea (Diplokoenikea) curvirostris Walter, 1919
Rosso de Ferrads & Mattoni, 1999: 118.
Par; Chaq; ln; lt
Koenikea (Diplokoenikea) flagellaris Lundblad, 1938
Viets, 1987: 406.
Par; ln.
Koenikea (Diplokoenikea) grossa Lundblad, 1938
Viets, 1987: 406.
Par; ln; lt.
Koenikea (Diplokoenikea) pectinata Lundblad, 1936
Viets, 1987: 406.
Par; lt.
Koenikea (Diplokoenikea) pectinifera Lundblad, 1938
Viets, 1987: 406; Cook, 1988: 160.
Par; Csc; lt; ln.
Koenikea (Diplokoenikea) quinquemaculata Lundblad, 1943
Viets, 1987: 406.
Par; ln.
Koenikea (Diplokoenikea) scutica Viets, 1977
Viets, 1987: 406.
Cace; lt; ln.
Koenikea (Diplokoenikea) simulans Lundblad, 1943
Viets, 1987: 407.
Par; ln.
Subgnero Jurucuia Viets, 1954
Koenikea (Jurucuia) oblongata (Viets, 1954)
Viets, 1987: 407.
Amaz; lt; ln.
Subgnero Koenikella Lundbblad, 1936
Koenikea (Koenikella) crassipalpis Lundblad, 1936
Viets, 1987: 407.
Par; lt.
Koenikea (Koenikella) cyanozona Lundblad, 1937
Viets, 1987: 407.
Par; lt.
204
Subgnero Neokoenikea Lundblad, 1936
Koenikea (Neokoenikea) armipes Lundblad, 1936
Viets, 1987: 407.
Par; lt.
Subgnero Notomideopsis Wolcott, 1905
Koenikea (Notomideopsis) acanthifera Lundblad, 1953
Viets, 1987: 407.
Par; lt.
Koenikea (Notomideopsis) acutidens Lundblad, 1943
Viets, 1987: 408.
Par; lt.
Koenikea (Notomideopsis) brasiliensis Lundblad, 1930
Viets, 1987: 408.
Par; ln.
Koenikea (Notomideopsis) chilensis Cook, 1988
Cook, 1988: 157.
Csc; lt.
Koenikea (Notomideopsis) circumcincta Lundblad, 1937
Viets, 1987: 408.
Par; lt.
Koenikea (Notomideopsis) consimilis Viets, 1975
Viets, 1987: 408.
Amaz; lt; ln.
Koenikea (Notomideopsis) crassa Lundblad, 1937
Viets, 1987: 408.
Par; lt.
Koenikea (Notomideopsis) curicola Viets, 1975
Viets, 1987: 408.
Amaz; lt; ln.
Koenikea (Notomideopsis) curvipes Lundblad, 1938
Par; Chaq; ln; lt.
Koenikea (Notomideopsis) diabolica Viets, 1975
Viets, 1987: 408.
Amaz; ln.
Koenikea (Notomideopsis) elegans Lundblad, 1938
Par; ln; lt.
Koenikea (Notomideopsis) hamulata Lundblad, 1936
Viets, 1987: 408.
Par; lt.
Koenikea (Notomideopsis) imitata Lundblad, 1930
Viets, 1987: 408.
Cace; Amaz; lt.
Koenikea (Notomideopsis) lunipes Lundblad, 1938
Viets, 1987: 408.
Par; ln.
Koenikea (Notomideopsis) nordenskioldi Cook, 1980
Viets, 1987: 408.
Yup; ln.
Koenikea (Notomideopsis) obscura Lundblad, 1937
Viets, 1987: 409.
Par; lt.
Koenikea (Notomideopsis) rectipes Lundblad, 1943
Viets, 1987: 409.
Par; ln.
Koenikea (Notomideopsis) soona Cook, 1988
Cook, 1988: 158.
Csc; lt.

Koenikea (Notomideopsis) spinosa Daday, 1905
Rosso de Ferrads & Mattoni, 1999: 120.
Par; Chaq; ln; lt.
Koenikea (Notomideopsis) tigrina tigrina Lundblad, 1936
Viets, 1987: 409.
Par; lt.
Koenikea (Notomideopsis) tigrina crassiseta Lundblad, 1943
Viets, 1987: 409.
Par; lt.
Subgnero Parakoenikea Lundblad, 1936
Koenikea (Parakoenikea) curvipalpis Lundblad, 1936
Viets, 1987: 410.
Par; lt; ln.
Koenikea (Parakoenikea) scutellum Besch, 1969
Viets, 1987: 410.
Amaz; lt.
Subgnero Pseudokoenikea Lundblad, 1941
Koenikea (Pseudokoenikea) horrida Lundblad, 1930
Viets, 1987: 410.
Par; Amaz; lt; ln.
Koenikea (Pseudokoenikea) rutae rutae Lundblad, 1930
Viets, 1987: 410.
Par; Amaz; lt; ln.
Koenikea (Pseudokoenikea) rutae furcaticornis Viets, 1956
Viets, 1987: 410.
Amaz; ln.
Subgnero Tamboella Lundblad, 1953
Koenikea (Tamboella) multiseta Lundblad, 1953
Viets, 1987: 411.
Yup; lt.
GNERO Neumania Lebert, 1879
Subgnero Neoneumania Lundblad, 1941
Neumania (Neoneumania) xiphophora Lundblad, 1941
Viets, 1987: 569.
Par; lt.
Subgnero Tetraneumania Lundblad, 1930
Neumania (Tetraneumania) arpodos Rosso de Ferrads, 1982
Viets, 1987: 571.
Chaq; ln.
Neumania (Tetraneumania) barbifera Lundblad, 1937
Viets, 1987: 571.
Par; lt.
Neumania (Tetraneumania) bituberculata Lundblad, 1942
Viets, 1987: 571.
Par; Amaz; ln; lt.
Neumania (Tetraneumania) breviseta Viets, 1959
Viets, 1987: 571.
Amaz; lt.
Neumania (Tetraneumania) circummarginata Viets, 1954
Viets, 1987: 571.
Cace; ln.

Neumania (Tetraneumania) curvipes Lundblad, 1930
Viets, 1987: 571.
Par; Cace; lt; ln.
Neumania (Tetraneumania) dorsoscutata Lundblad, 1938
Viets, 1987: 571.
Par; ln.
Neumania (Tetraneumania) gaethgensi Besch, 1965
Viets, 1987: 571.
Amaz; lt.
Neumania (Tetraneumania) lateribarbata Lundblad, 1937
Viets, 1987: 572.
Par; Cace; len.
Neumania (Tetraneumania) loricata Lundblad, 1936
Viets, 1987: 572.
Neumania (Tetraneumania) paupercula Viets, 1954
Viets, 1987: 572.
Cace; Amaz; ln.
Neumania (Tetraneumania) polytricha Lundblad, 1938
Par; ln; lt.
Neumania (Tetraneumania) schadei Lundblad, 1942
Viets, 1987: 572.
Par; ln; lt.
GNERO Recifella Viets, 1935
Recifella columbiana (Lundblad, 1953)
Viets, 1987: 674.
Yup; lt.
Recifella inka (Lundblad, 1944)
Viets, 1987: 674.
Yup; lt.
Recifella paucipora (Lundblad, 1943)
Viets, 1987: 675.
Par; lt.
Recifella pulchra (Lundblad, 1937)
Viets, 1987: 675.
Par; lt.
Recifella truncata (Besseling, 1949)
Viets, 1987: 675.
Amaz; lt.
Subgnero Recifella Viets, 1935
Recifella (Recifella) angulipes (Lundblad, 1938)
Viets, 1987: 674.
Par; ln; lt.
Recifella (Recifella) clavata (Lundblad, 1937)
Viets, 1987: 674
Par; lt.
Recifella (Recifella) gibberipalpis (Lundblad, 1936)
Viets, 1987: 674.
Par; lt; ln.
Recifella (Recifella) laminipes (Viets, 1935)
Viets, 1987: 674.
Par; Amaz; lt; ln.
Recifella (Recifella) latipes (Lundblad, 1936)
Viets, 1987: 674.
Par; Amaz; lt.
205
Recifella (Recifella) lundbladi (Besseling, 1949)
Viets, 1987: 675.
Amaz; lt, ln.
Recifella (Recifella) sinuatipes (Lundblad, 1941)
Viets, 1987: 675.
Par; ln.
Subgnero Eorecifella Cook, 1980
Recifella (Eorecifella) balteata (Lundblad, 1943)
Viets, 1987: 675.
Par; lt.
Recifella (Eorecifella) cruciata cruciata (Lundblad, 1936)
Viets, 1987: 675.
Par; lt.
Recifella (Eorecifella) cruciata brevis (Lundblad, 1943)
Viets, 1987: 675.
Par; lt.
Recifella (Eorecifella) diplothoracica (Lundblad, 1937)
Viets, 1987: 675.
Par; lt.
Recifella (Eorecifella) elliptica elliptica (Walter, 1919)
Viets, 1987: 675.
Par; Amaz; lt; ln.
Recifella (Eorecifella) elliptica amabilis (Lundblad, 1943)
Viets, 1987: 676.
Par; lt; ln.
Recifella (Eorecifella) elliptica schadei (Lundblad, 1943)
Viets, 1987: 676.
Par; lt.
Recifella (Eorecifella) elliptica validipalpis (Lundblad, 1937)
Viets, 1987: 676.
Par; lt; ln.
Recifella (Eorecifella) excavata (Lundblad, 1936)
Par, Amaz; lt.
Recifella (Eorecifella) excavatoides (Viets, 1975)
Viets, 1987: 676.
Amaz; lt; ln.
Recifella (Eorecifella) opimipalpis (Viets, 1977)
Viets, 1987: 676.
Amaz; lt.
Recifella (Eorecifella) pusilla (Lundblad, 1937)
Viets, 1987: 676.
Par; lt.
Recifella (Eorecifella) undulata (Lundblad, 1936)
Par; lt; ln.
Subgnero Monokoenikea Lundblad, 1941
Recifella (Monokoenikea) melini (Lundblad, 1930)
Viets, 1987: 677.
Par; ln.
GNERO Schadeella Lundblad, 1938
Schadeella crassipalpis Lundblad, 1938
Viets, 1987: 683.
Par; Amaz; ln; lt.
206

FAMILIA PIONIDAE Thor, 1900
SUBFAMILIA PIONINAE Thor, 1900
GNERO Piona Koch, 1842

Piona aculeatipalpis Viets, 1954
Viets, 1987: 606.
Amaz; Par; ln.
Piona acutidens Lundblad, 1941
Viets, 1987: 606.
Par; lt.
Piona amazonica Viets, 1954
Viets, 1987: 608.
Amaz; ln.
Piona anisitsi Daday, 1905
Viets, 1987: 609.
Par; ln.
Piona atrorubra Lundblad, 1937
Viets, 1987: 609.
Par; Chaq; Amaz; ln; lt.
Piona chilensis Viets, 1968
Viets, 1987: 612; Cook, 1988: 160.
Csc; ln.
Piona confinis Lundblad, 1943
Viets, 1987: 617.
Par; ln.
Piona davidcooki Rosso de Ferrads & Bttger, 1997
Par; ln.
Piona deformis Lundblad, 1941
Par; Chaq; Par; ln; lt.
Piona dejecta Viets, 1937
Viets, 1987: 621.
Cace; Amaz; ln.
Piona dissipata Lundblad, 1938
Viets, 1987: 622.
Par; ln.
Piona erratica Marshall, 1940
Viets, 1987: 623.
Chaq; Aap; ln.
Piona fissa Viets, 1940
Viets, 1987: 623.
Cace; ln.
Piona juncta juncta Lundblad, 1936
Viets, 1987: 626.
Par; Amaz; ln.
Piona juncta inconspicua Lundblad, 1943
Viets, 1987: 626.
Par; Cace; ln.
Piona oxyura Viets, 1954
Viets, 1987: 634.
Amaz; ln.
Piona panniculata Viets, 1940
Viets, 1987: 634.
Amaz; ln.
Piona plaumanni Lundblad, 1943
Viets, 1987: 635.
Par; Amaz; ln.
Piona plenipalpis Viets, 1954
Viets, 1987: 635.
Cace; ln.
Piona praegracilis Viets, 1937

Viets, 1987: 636.
Cace; ln.
Piona punctatissima Viets, 1954
Viets, 1987: 636.
Cace; ln.
Piona rectimarginata Viets, 1954
Viets, 638.
Amaz; ln.
Piona reflexidens Viets, 1940
Viets, 1987: 639.
Cace; Par; lt.
Piona robustipalpis Viets, 1954
Viets, 1987: 639.
Amaz; ln.
Piona rotunda rotunda (Kramer, 1879)
Viets, 1987: 639.
Amaz; Cv; Chaq; ln; lt.
Piona rotunda acutipes Viets, 1954
Viets, 1987: 639.
Cace; ln.
Piona rotunda rotundiformis Lundblad, 1938
Viets, 1987: 639.
Par; ln.
Piona serratiseta Viets, 1956
Viets, 1987: 641.
Cv; lt.
Piona setipes Cook, 1980
Viets, 1987: 641; Rosso de Ferrads, Kaisin & Bosnia,
1987: 119.
Aap; ln.
Piona sicaria Lundblad, 1937
Viets, 1987: 641.
Par; ln.
Piona sudamericana Viets, 1910
Viets, 1987: 642.
Par; Chaq; Cace; Amaz; ln; lt.
Piona tenuipalpis Viets, 1954
Viets, 1987: 642.
Amaz; ln.
Piona thoracica Lundblad, 1941
Viets, 1987: 642.
Par; Chaq; lt; ln.
Piona ungularis Lundblad, 1943
Viets, 1987: 644.
Par; Chaq; ln, lt.
Piona unidens Viets, 1954
Viets, 1987: 644.
Amaz; ln.
Piona venezuelensis Viets, 1956
Viets, 1987: 645.
Cv; lt.
FAMILIA ATURIDAE Thor, 1900
SUBFAMILIA FRONTIPODOPSINAE Viets, 1931
GNERO Frontipodopsis Walter, 1919
Frontipodopsis chilensis Cook, 1988
Cook, 1988: 161.
Csc; lt.

Frontipodopsis staheli staheli Walter, 1919
Viets, 1987: 292.
Amaz; lt-est.
Frontipodopsis staheli mandibularis Lundblad, 1943
Viets, 1987: 292.
Par; lt.
SUBFAMILIA ATURINAE Thor, 1900

GNERO Aturides Lundbld, 1937
Aturides dentatus (Lundblad, 1936)
Viets, 1987: 164.
Par; Amaz; lt.
GNERO Aturus Kramer, 1875
Aturus andinus Lundblad, 1953
Viets, l987: 165.
Yup; lt.
Aturus kronestedti Cook, 1988
Cook, 1988: 170.
Csc; lt.
207
SUBFAMILIA AXONOPSINAE Viets, 1929
GNERO Axonopsella Lundblad, 1930
Axonopsella affinis Lundblad, 1943
Viets, 1987: 181.
Par; lt.
Axonopsella andina Lundblad, 1953
Viets, 1987: 181.
Yup; Csc; lt, lt-ln.
Axonopsella henrechi Orghidan & Gruia, 1981
Viets, 1987: 181.
Cv; lt.
Axonopsella plaumanni Lundblad, 1936
Viets, 1987: 181.
Par; lt.
Axonopsella spinigera Lundblad, 1930
Viets, 1987: 181.
Par; lt.
Axonopsella vaginalis Viets, 1936
Viets, 1987: 181.
Cace; lt.
Axonopsella vietsi Lundblad, 1937
Viets, 1987: 181.
Par; Cace; lt.
GNERO Kongsbergia Thor, 1899

Subgnero Chilaxonopsella Cook, 1988
Kongsbergia globipalpis Lundblad, 1953
Yup; Chaq; lt.
GNERO Neoaturus Lundblad, 1941
Neoaturus caudatus Lundblad, 1953
Viets, 1987: 539.
Yup; lt; ln; lt-ln.
Neoaturus dentipes (Lundblad, 1936)
Viets, 1987: 539.
Par; lt; ln.
Neoaturus multiporus (Walter, 1919)
Viets, 1987: 540.
Amaz; ln.
Neoaturus novus novus Lundblad, 1943
Viets, 1987: 540.
Par; lt-ln.
Neoaturus novus plaumanni Lundblad, 1943
Viets, 1987: 540.
Par; lt.
Neoaturus paucidens (Lundblad, 1937)
Viets, 1987: 540.
Par; lt.
Neoaturus projectus (Lundblad, 1936)
Viets, 1987: 540.
Par; lt-ln.
Neoaturus setipes (Lundblad, 1936)
Viets, 1987: 540.
Par; lt.
Neoaturus tornquistensis Cook, 1980
Viets, 1987: 540.
Chaq; lt.
Axonopsella (Chilaxonopsella) barri Cook, 1988

Cook, 1988: 165.
Csc; lt.
Axonopsella (Chilaxonopsella) chilensis Cook, 1988
Cook, 1987: 162.
Csc; lt.
Axonopsella (Chilaxonopsella) doloma Cook, 1988
Cook, 1988: 164.
Csc; lt.
Axonopsella (Chilaxonopsella) tanda Cook, 1988
Cook, 1988: 166.
Csc; lt.
Subgnero Coaxonopsella Cook, 1980
Axonopsella (Coaxonopsella) jujuyensis Cook, 1980
Viets, 1987: 182.
Yup; lt.
Subgnero Humboldtaxonopsella Orghidan & Gruia, 1981
Axonopsella (Humboldtaxonopsella) linaresi Orghidan & Gruia,
1981
Viets, 1987: 182.
Cv; lt.
Subgnero Luciaxonopsella Rosso de Ferrads, 1995
Axonopsella (Luciaxonopsella) misionum Rosso de Ferrads,
1995
Chaq; lt.
208
Subgnero Neoaxonopsella Lundblad, 1937
Subgnero Miraxonidella Cook, 1980
Axonopsella (Neoaxonopsella) argentinensis Cook, 1980

Viets, 1987: 182.
Yup; lt.
Axonopsella (Neoaxonopsella) caripense Orghidan & Gruia, 1981
Viets, 1987: 182.
Cv; lt.
Axonopsella (Neoaxonopsella) hamatoides Lundblad, 1937
Par; Chaq; lt.
Axonopsella (Neoaxonopsella) mamillata Viets, 1954
Viets, 1987: 182.
Amaz; ln.
Axonopsella (Neoaxonopsella) trifida Lundblad, 1937
Par; Chaq; lt.
Miraxonides (Miraxonidella) karlvietsi Cook, 1980

Viets, 1987: 529.
Yup; lt.
Subgnero Paraxonopsella Lundbblad, 1937.

Axonopsella (Paraxonopsella) bifida Lundblad, 1937
Par; Chaq; lt.
Axonopsella (Paraxonopsella) filunguis Lundblad, 1937
Par; Chaq; lt.
Axonopsella (Paraxonopsella) hamata Lundblad, 1936
Par; Chaq; lt.
GNERO Neoalbia Lundblad, 1936

Subgnero Neoalbia Lundblad, 1936
Neoalbia (Neoalbia) violacea Lundblad, 1936
Viets, 1987: 538.
Par; lt.
Subgnero Neoalbiella Viets, 1954
Neoalbia (Neoalbiella) bidentata Viets, 1959
Viets, 1987: 538.
Amaz; lt.
Neoalbia (Neoalbiella) octopora Viets, 1954
Viets, 1987: 538.
Amaz; lt.
GNERO Neoaxona Lundblad, 1936
Neoaxona aculeata Besseling, 1949
Viets, 1987: 540.
Amaz; lt.
Neoaxona plaumanni Lundblad, 1937
Viets, 1987: 540.
Par; lt-ln.
Subgnero Rostaxonopsella Orghidan & Gruia, 1987
Subgnero Neoaxona Lundblad, 1936
Axonopsella (Rostaxonopsella) nana Orghidan & Gruia, 1987

Cv; est.
Neoaxona (Neoaxona) oblonga Lundblad, 1936

Viets, 1987: 540.
Par; lt-ln.
GNERO Axonopsis Piersig, 1893

Subgnero Brachypodopsis Piersig, 1903
Subgnero Lamellaxona Lundblad, 1937
Axonopsis (Brachypodopsis) boutata Cook, 1980

Viets, 1987: 186.
Yup; lt.
Axonopsis (Brachypodopsis) columbicola Lundblad, 1953
Viets, 1987: 186.
Yup; lt; ln.
GNERO Lethaxonella Cook, 1963
Lethaxonella argentinensis Cook, 1980
Viets, 1987: 470.
Yup; est.
GNERO Miraxonides Lundblad, 1938
Subgnero Miraxonides Lundblad, 1938
Miraxonides (Miraxonides) alatus Lundblad, 1938
Viets, 1987: 529.
Par; Cace; ln.
Neoaxona (Lamellaxona) abnormipes Lundblad, 1937

Viets, 1987: 540.
Par; lt.
GNERO Neoaxonopsis Lundblad, 1938
Neoaxonopsis odontogaster Lundblad, 1938
Viets, 1987: 541.
Par; lt.
GNERO Polyaxonopsella Lundblad, 1943
Polyaxonopsella bordoni (Orghidan & Gruia, 1981)
Viets, 1987: 655.
Cv; lt.
Polyaxonopsella dentifera Lundblad, 1943
Viets, 1987: 655.
Par; lt.
Polyaxonopsella polypora Lundblad, 1953
Viets, 1987: 656.
Yup; lt.

GNERO Stygalbiella Cook, 1974
Stygalbiella tucumanensis Cook, 1980
Viets, 1987: 714; Fernndez, 2003: 61.
Yup; lt.
GNERO Submiraxona Lundblad, 1937
Subgnero Submiraxona Lundblad, 1937
Submiraxona (Submiraxona) crassipes Lundblad, 1937
Par; Chaq; lt.
Submiraxona (Submiraxona) rutae Lundblad, 1937
Viets, 1987: 719.
Par; lt; ln.
Subgnero Pentalbia Lundlbad, 1937
Submiraxona (Pentalbia) walteri (Lundblad, 1937)
Viets, 1987: 720.
Par; lt-ln.
SUBFAMILIA ALBIINAE Viets, 1915
GNERO Albia Thor, 1899
Subgnero Anchistalbia Cook, 1974
Albia (Anchistalbia) nobilis Lundblad, 1936
Par; Chaq; lt.
SUBFAMILIA NOTOATURINAE Besch, 1964
209
Noesaturus otendus Cook, 1988
Cook, 1988: 180.
Csc; lt.
GNERO Notaxona Besch, 1964
Notaxona ochiepus Besch, 1964
Viets, 1987: 578; Cook, 1988: 170.
Csc; lt.
SUPERFAMILIA ARRENUROIDEA Thor, 1900
FAMILIA MOMONIIDAE Viets, 1926
GNERO Momoniella Viets, 1929
Momoniella sudamericana Cook, 1988
Cook, 1988: 181.
Csc; lt.
GNERO Notomomonia Cook, 1988
Notomomonia alza Cook, 1988
Cook, 1988: 182.
Csc; lt.
Notomomonia anchista Cook, 1988
Cook, 1988: 183.
Csc; lt.
Notomomonia crassipes Cook, 1988
Cook, 1988: 184.
Csc; lt.
FAMILIA MIDEOPSIDAE Koenike, 1910
GNERO Notoaturus Besch, 1964
GNERO Mideopsis Neuman, 1880
Notoaturus arrenuripalpis Cook, 1988

Cook, 1988: 175.
Csc; lt.
Notoaturus beschi Cook, 1988
Cook, 1988: 173.
Csc; lt.
Notoaturus leptoglyphus Besch, 1964
Viets, 1987: 578; Cook, 1988: 172.
Csc; lt.
Notoaturus phreaticus Schwoerbel, 1986
Viets, 1987: 1001.
Csc; est.
Mideopsis dentifera Lundblad, 1943

Viets, 1987: 517.
Par; lt.
Mideopsis plaumanni Lundblad, 1937
Viets, 1987: 519.
Par; Amaz; lt.
Mideopsis singularis Viets, 1954
Viets, 1987: 520.
Amaz; lt.
Mideopsis venusta Lundblad, 1943
Viets, 1987: 520.
Par; lt.
Mideopsis violacea Lundblad, 1943
Viets, 1987: 520.
Par; lt.
GNERO Noesaturus Cook, 1988

Noesaturus chiloensis Cook, 1988
Cook, 1988: 179.
Csc; lt.
Noesaturus expansipes Cook, 1988
Cook, 1988: 178.
Csc; lt.
Noesaturus hopkinsi Cook, 1988
Cook, 1988: 176.
Csc; lt.
Subgnero Mideopsis Neuman, 1880

Mideopsis (Mideopsis) choconensis Cook, 1980
Viets, 1987: 516; Rosso de Ferrads, 1987: 34; Cook, 1988:
185.
Aap; Csc; ln; lt.
Mideopsis (Mideopsis) clavipes Lundblad, 1936
Viets, 1987: 516.
Par; lt.
210
Mideopsis (Mideopsis) elongata (Viets, 1935)
Viets, 1987: 517.
Cace; Par; lt.
Mideopsis (Mideopsis) gradatipalpis Viets, 1954
Viets, 1987: 517.
Amaz; lt; ln.
Mideopsis (Mideopsis) infractipalpis Viets, 1954
Viets, 1987: 517.
Amaz; ln.
Mideopsis (Mideopsis) longidens Lundblad, 1943
Viets, 1987: 517.
Par; Yup; ln; lt.
Mideopsis (Mideopsis) microdactylus Viets, 1954
Viets, 1987: 518.
Cace; lt.
Mideopsis (Mideopsis) oviformis Lundblad, 1944
Viets, 1987: 519.
Yup; lt.
Mideopsis (Mideopsis) ovum Lundblad, 1937
Viets, 1987: 519.
Par; Cace; lt.
Mideopsis (Mideopsis) pacrodactylus Lundblad, 1938
Viets, 1987: 519.
Par; Chaq; ln; lt.
Mideopsis (Mideopsis) rectangularis Lundblad, 1937
Viets, 1987: 520.
Par; Amaz; lt.
Mideopsis (Mideopsis) rostrata Lundblad, 1943
Viets, 1987: 520.
Par; lt.
Mideopsis (Mideopsis) trifoliata Viets, 1959
Viets, 1987: 520.
Amaz; lt.

Mideopsis (Neoxystonotus) sica Lundblad, 1943
Par; ln; lt.
Mideopsis (Neoxystonotus) trilobata trilobata Viets, 1935
Viets, 1987: 523.
Cace; Par; lt; ln.
Mideopsis (Neoxystonotus) trilobata lineata Lundblad, 1936
Viets, 1987: 523.
Par; Cace; lt; ln.
Mideopsis (Neoxystonotus) trilobata remotipora Viets, 1954
Viets, 1987: 523.
Cace; ln.
Mideopsis (Neoxystonotus) trilobata vicinipora Viets, 1954
Viets, 1987: 524.
Cace; lt.
GNERO Mideopsella Lundblad, 1937
Mideopsella forficipalpis Lundblad, 1937
Viets, 1987: 514.
Par; lt.
GNERO Phreatomideopsis Schwoerbel, 1986
Phreatomideopsis arrenuripalpis Schwoerbel, 1986
Viets, 1987: 1003; Cook, 1988: 186.
Csc; lt.
FAMILIA KRENDOWSKIIDAE Viets, 1926
GNERO Krendowskia Piersig, 1895
Subgnero Krendowskia Piersig, 1895
Subgnero Mideopsides Lundblad, 1943

Mideopsis (Mideopsides) gibberipalpis Lundblad, 1943
Viets, 1987: 522.
Par; Cace; Amaz; lt.
Subgnero Neoxystonotus Lundblad, 1927
Mideopsis (Neoxystonotus) affinis Lundblad, 1943
Viets, 1987: 522.
Par; lt.
Mideopsis (Neoxystonotus) biverrucata Viets, 1954
Viets, 1987: 522.
Cace; Amaz; ln.
Mideopsis (Neoxystonotus) caviventris Lundblad, 1941
Viets, 1987: 523.
Par; lt.
Mideopsis (Neoxystonotus) curvipes Viets, 1954
Viets, 1987: 523.
Amaz; ln.
Mideopsis (Neoxystonotus) genitalis Lundblad, 1936
Viets, 1987: 523.
Par; Cace; lt.
Mideopsis (Neoxystonotus) maculata Viets, 1935
Viets, 1987: 523.
Cace; ln.
Mideopsis (Neoxystonotus) paprzyckii Lundblad, 1944
Viets, 1987: 523.
Yup; lt.
Krendowskia (Krendowskia) convexa convexa (Ribaga, 1902)

Viets, 1987: 421; Rosso de Ferrads, Fernndez &
Rocabado, 2004: 312.
Csc; Chaq; Yup; ln; lt.
Krendowskia (Krendowskia) dorsolineata Lundblad, 1936
Viets, 1987: 421.
Par; lt.
Krendowskia (Krendowskia) frontalis frontalis Lundblad, 1938
Viets, 1987: 421.
Par; ln.
Krendowskia (Krendowskia) frontalis tricolor Lundblad, 1938
Viets, 1987: 421.
Par; ln.
Krendowskia (Krendowskia) laceripalpis Lundblad, 1943
Viets, 1987: 421.
Par; Cace; lt; ln.
Subgnero Krendowskiella Viets, 1931
Krendowskia (Krendowskiella) megalopsis Lundblad, 1930
Viets, 1987: 422.
Par; Amaz; ln; lt.
Krendowskia (Krendowskiella) quadrimaculata Lundblad, 1938
Viets, 1987: 422.
Par; Amaz; ln; lt.
Krendowskia (Krendowskiella) rubicunda Viets, 1954
Viets, 1987: 423.
Amaz; ln.

Krendowskia (Krendowskiella) rufolineata Lundblad, 1936
Viets, 1987: 423.
Par; lt.
Subgnero Neokrendowskia Lundblad, 1941
Krendowskia (Neokrendowskia) mediolineata Viets, 1959
Viets, 1987: 423.
Amaz; lt.
Krendowskia (Neokrendowskia) quadripustulata Lundblad, 1941
Viets, 1987: 423.
Par; ln.
GNERO Roqueella Lundblad, 1930
Subgnero Roqueella Lundblad, 1930
Roqueella (Roqueella) flabellifera Lundblad, 1930
Viets, 1987: 681.
Yup; lt.
Roqueella (Roqueella) lenzi (Viets, 1935)
Viets, 1987: 681.
Cace; lt.
Subgnero Decaroqueella Viets, 1954
Roqueella (Decaroqueella) actinophora Viets, 1954
Viets, 1987: 681.
Cace; lt.
Subgnero Hexaroqueella Viets, 1959
Roqueella (Hexaroqueella) longimaxilaris Viets, 1959
Viets, 1987: 681.
Amaz; lt.
Subgnero Neoroqueella Lundblad, 1937
Roqueella (Neoroqueella) lundbladi Viets, 1954
Viets, 1987: 681.
Amaz; lt-ln.
Roqueella (Neoroqueella) maculata Lundblad, 1937
Viets, 1987: 681.
Par; lt.
211
Subgnero Geayia Lundblad, 1936
Geayia (Geayia) venezuelae Thor, 1897
Viets, 1987: 293.
Cv; Amaz; lt.
Subgnero Geayella Thor, 1897
Geayia (Geayella) amazonica Viets, 1954
Viets, 1987: 293.
Amaz; lt.
Geayia (Geayella) catharinensis Lundblad, 1936
Viets, 1987: 293.
Par; lt.
Subgnero Geayidea Lundblad, 1941
Geayia (Geayidea) avinotata Viets, 1959
Viets, 1987: 293.
Amaz; ln.
Geayia (Geayidea) coeruleocruciata Lundblad, 1936
Viets, 1987: 294.
Par; lt.
Geayia (Geayidea) coeruleolineata Lundblad, 1936
Viets, 1987: 294.
Par; lt.
Geayia (Geayidea) inchoata Viets, 1954
Viets, 1987: 294.
Cace; lt.
Geayia (Geayidea) latirostris Viets, 1954
Viets, 1987: 294.
Amaz; lt.
Geayia (Geayidea) micronycha Viets, 1954
Viets, 1987: 294.
Cace; ln.
Geayia (Geayidea) monochroa Viets, 1954
Viets, 1987: 294.
Cace; lt.
Subgnero Pirapama Viets, 1935
Geayia (Pirapama) schubarti Viets, 1935
Viets, 1987: 294.
Cace; Amaz, Par; ln; lt.
Subgnero Pararoqueella Lundblad, 1937

FAMILIA ATHIENEMANNIIDAE Viets, 1922
Roqueella (Pararoqueella) conimaxillaris Viets, 1954
Viets, 1987: 681.
Amaz; lt.
Roqueella (Pararoqueella) cruciata Lundblad, 1937
Viets, 1987: 681.
Par; lt.
Roquella (Pararoqueella) papillata Viets, 1954
Viets, 1987: 681.
Amaz; lt.
Roqueella (Pararoqueella) striata Lundblad, 1937
Viets, 1987: 681.
Par; lt.
GNERO Plaumannia Lundblad, 1936

Plaumannia arrenuripalpis Lundblad, 1936
Viets, 1987: 654.
Par; lt.
Plaumannia crenophila Lundblad, 1938
Viets, 1987: 654.
Par; ln.
FAMILIA ARRENURIDAE Thor, 1900
GNERO Geayia Thor, 1897
GNERO Arrenurus Dges, 1834
Geayia flagellifera Viets, 1954

Viets, 1987: 293.
Cace; lt.
Arrenurus acutidentatus Viets, 1959

Viets, 1987: 104.
Amaz; ln.
212
Arrenurus acutiepimeratus Viets, 1954
Viets, 1987: 104.
Cace; lt.
Arrenurus amplipenicillatus Viets, 1954
Viets, 1987: 104.
Cace; ln.
Arrenurus anisitsi Daday, 1905
Viets, 1987: 104.
Par; ln.
Arrenurus apertus Daday, 1905
Viets, 1987: 104.
Par; ln.
Arrenurus boettgeri Viets, 1968
Viets, 1987: 104; Cook, 1988: 189.
Csc; ln.
Arrenurus caquetiorum Rosso de Ferrads & Fernndez, 2001
Rosso de Ferrads & Fernndez, 2001a: 54.
Cv; fit.
Arrenurus cometes Lundblad, 1937
Viets, 1987: 38.
Par; ln.
Arrenurus confertus Viets, 1954
Viets, 1987: 105.
Cace; ln; lt.
Arrenurus dentipalpis Lundblad, 1936
Viets, 1987: 42
Par; lt; ln.
Arrenurus hirsutipalpis Walter, 1919
Viets, 1987: 107.
Yup; lt.
Arrenurus honoratus Thor, 1911
Viets, 1987: 107.
Par; ln.
Arrenurus multangulus Daday, 1905
Viets, 1987: 108.
Par; ln.
Arrenurus multipapillatus Viets, 1954
Viets, 1987: 108.
Cace; ln.
Arrenurus ovum Viets, 1954
Viets, 1987: 109.
Cace; ln.
Arrenurus papilliger Viets, 1959
Viets, 1987: 109.
Amaz; ln.
Arrenurus pennapodus Rosso de Ferrads, 1989
Par; ln.
Arrenurus promacrus Lundblad, 1936
Viets, 1987: 53.
Par; ln.
Arrenurus propinquus Daday, 1905
Viets, 1987: 109.
Par; ln; lt.
Arrenurus quadrisetosus Viets, 1954
Viets, 1987: 109.
Amaz; ln.
Arrenurus tortus Viets, 1954
Viets, 1987: 110.
Amaz; ln.
Arrenurus tumulosus tumulosus Viets, 1954
Viets, 1987: 111.
Amaz; ln.

Arrenurus tumulosus intercursor Viets, 1954
Viets, 1987: 111.
Cace; ln.
Arrenurus uberifer Viets, 1954
Viets, 1987: 111.
Amaz; lt.
Subgnero Arrenurus Dges, 1834
Arrenurus (Arrenurus) brasiliensis Lundblad, 1941
Par; Chaq; ln; lt.
Arrenurus (Arrenurus) brevipetiolatus Lundblad, 1936
Viets, 1987: 36.
Par; lt.
Arrenurus (Arrenurus) conspicuos Viets & Bttger, 1986
Viets, 1987: 986.
Par; ln.
Arrenurus (Arrenurus) erectipetiolatus Viets, 1954
Viets, 1987: 43.
Amaz; ln.
Arrenurus (Arrenurus) flagellatus Lundblad, 1936
Viets, 1987: 44.
Par; lt.
Arrenurus (Arrenurus) inflatipalpis Lundblad, 1944
Viets, 1987: 46.
Par; ln.
Arrenurus (Arrenurus) lobatus Lundblad, 1941
Viets, 1987: 48.
Par; ln.
Arrenurus (Arrenurus) mystrophorus Lundblad, 1938
Par; Chaq; ln.
Arrenurus (Arrenurus) oxyurus Ribaga, 1902
Viets, 1987: 51; Rosso de Ferrads, 1987a: 35.
Yup; Aap; Chaq; Par; ln; lt.
Arrenurus (Arrenurus) plaumanni Lundblad, 1936
Viets, 1987: 52.
Par; ln; lt.
Arrenurus (Arrenurus) rectipetiolatus Lundblad, 1941
Viets, 1987: 54.
Yup; ln.
Arrenurus (Arrenurus) ringueleti Rosso de Ferrads, 1980
Viets, 1987: 54.
Chaq; ln; lt.
Arrenurus (Arrenurus) ripiphorus Lundblad, 1937
Viets, 1987: 55.
Par; ln.
Arrenurus (Arrenurus) sellatus Viets, 1954
Viets, 1987: 56.
Chaq; lt.
Arrenurus (Arrenurus) subulifer Lundblad, 1938
Viets, 1987: 56.
Par; ln.
Arrenurus (Arrenurus) trichophorus Daday, 1905
Viets, 1987: 57.
Par; Chaq; ln.
Arrenurus (Arrenurus) valdiviensis Viets, 1964
Viets, 1987: 58; Rosso de Ferrads, 1987b: 119; Cook,
1988: 188.
Csc; Aap; ln.
213
Arrenurus (Arrenurus) valencius Marshall, 1919

Viets, 1987: 58.
Cv; ln; lt.
Arrenurus (Arrenurus) vicinus Lundblad, 1944
Viets, 1987: 59.
Par; ln.
Arrenurus (Dadayella) rotunda subrotundata (Viets, 1954)

Viets, 1987: 220; Cramer & Cook, 1992: 221.
Cace; Amaz; ln; lt.
Arrenurus (Dadayella) walteri (Lundblad, 1953)
Yup; lt.
Subgnero Arrhenuropsides Viets, 1954
Subgnero Megaluracarus Viets, 1911.
Arrenurus (Arrhenuropsides) parviscutatus (Viets, 1954)

Viets, 1987: 114.
Cace; ln.
Arrenurus (Arrhenuropsides) rionegrensis Rosso de Ferrads,
1978
Viets, 1987: 114.
Chaq; ln.
Arrenurus (Megaluracarus) adulterinus Viets, 1954

Viets, 1987: 61.
Amaz; Cace; Par; ln.
Arrenurus (Megaluracarus) amazonicus Viets, 1954
Viets, 1987: 61.
Amaz; lt; ln.
Arrenurus (Megaluracaraus) andrewfieldi Orghidan & Gruia,
1983
Viets, 1987: 986; Rosso de Ferrads & Fernndez, 2001a: 57.
Cv; fit.
Arrenurus (Megaluracarus) angulicauda Lundblad, 1937
Viets, 1987: 61; Rosso de Ferrads; 1984: 81.
Par; Chaq; ln; lt.
Arrenurus (Megaluracarus) angulosus Viets, 1974
Viets, 1987: 61.
Amaz; lt.
Arrenurus (Megaluracarus) archangelskii Rosso de Ferrads,
1998
Chaq; ln.
Arrenurus (Megaluracarus) bachmanni Rosso de Ferrads &
Bttger, 1997
Par; ln.
Arrenurus (Megaluracarus) bilaciniatus Viets, 1954
Viets, 1987: 62.
Amaz; ln.
Arrenurus (Megaluracarus) bipetiolatus Viets, 1954
Viets, 1987: 62.
Amaz; ln.
Arrenurus (Megaluracarus) bisinuosus Viets, 1954
Viets, 1987: 62.
Amaz; ln.
Arrrenurus (Megaluracarus) bituberculatus Viets, 1936
Viets, 1987: 62.
Cace; ln.
Arrenurus (Megaluracarus) bonettoi Rosso de Ferrads, 1973
Viets, 1987: 62.
Chaq; ln; lt.
Arrenurus (Megaluracarus) brevicaudatus Viets, 1936
Viets, 1987: 62.
Par; Amaz; Chaq; ln; lt.
Arrenurus (Megalurcarus) cachoerensis Lundblad, 1941
Viets, 1987: 63.
Par; ln.
Arrenurus (Megaluracarus) cataglyphus Lundblad, 1941
Viets, 1987: 64.
Par; ln.
Arrenurus (Megaluracarus) catharinensis Lundblad, 1937
Viets, 1987: 64.
Par; ln.
Arrenurus (Megaluracarus) consaguineus Viets, 1954
Viets, 1987: 65.
Amaz; ln; lt.
Subgnero Arrhenuropsis Viets, 1954

Arrenurus (Arrhenuropsis) clavilaminata (Viets, 1954)
Viets, 1987: 114.
Amaz; ln.
Arrenurus (Arrhenuropsis) curvipalpis (Viets, 1954)
Viets, 1987: 114.
Amaz; ln.
Subgnero Brevicaudaturus Smit, 1998
Arrenurus (Brevicaudaturus) imperator imperator Lundblad,
1944
Viets, 1987: 107; Smit, 1998: 253.
Par; ln.
Arrenurus (Brevicaudaturus) imperator goliath Lundblad, 1953
Viets, 1987: 107; Smit, 1998: 253.
Yup; ln.
Arrenurus (Brevicaudaturus) toriger Viets, 1940
Viets, 1987: 97; Smit: 253.
Cace; lt; ln.
Subgnero Dadayella Koenike, 1907
Arrenurus (Dadayella) clavipes (Lundblad, 1938)
Par; lt.
Arrenurus (Dadayella) guarani nomen novum (ver Comentarios)
Par; lt.
Arrenurus (Dadayella) guttifera (Viets, 1954)
Viets, 1987: 219; Cramer & Cook, 1992; 221.
Cace; ln.
Arrenurus (Dadayella) hirtipalpis (Besseling, 1949)
Viets, 1987: 220; Cramer & Cook, 1992; 221.
Amaz; ln.
Arrenurus (Dadayella) minima (Daday, 1905)
Par; Amaz; ln.
Arrenurus (Dadayella) projecta (Lundblad, 1938)
Par; Amaz; Chaq; ln; lt.
Arrenurus (Dadayella) rotunda rotunda (Daday, 1905)
Par; Cace; Chaq; lt; ln.
214
Arrenurus (Megaluracarus) cornicaudatus Viets, 1937
Viets, 1987: 65.
Amaz; Cace; ln.
Arrenurus (Megaluracarus) cornifrons Lundblad, 1938
Viets, 1987: 65.
Par; lt; ln.
Arrenurus (Megaluracarus) corniger Koenike, 1894
Viets, 1987: 65.
Par; Amaz; Cace; Chaq; ln; lt.
Arrenurus (Megaluracarus) corpulentus Viets, 1937
Viets, 1987: 65.
Cace; lt.
Arrenurus (Megaluracarus) correntinae Rosso de Ferrads &
Smit, 1998
Chaq; ln.
Arrenurus (Megaluracarus) crassus Lundblad, 1936
Viets, 1987: 65.
Par; Amaz; lt; ln.
Arrenurus (Megaluracarus) crenicauda Lundblad, 1938
Viets, 1987: 65.
Par; ln; lt.
Arrenurus (Megaluracrus) cultriger Viets, 1938
Viets, 1987: 65.
Amaz; Cace; ln; lt.
Arrenurus (Megaluracarus) curticaudatus Viets, 1938
Viets, 1987: 65.
Chaq; ln.
Arrenurus (Megaluracarus) davidcooki Rosso de Ferrads, 1998
Rosso de Ferrads, 1998: 332
Chaq; ln.
Arrenurus (Megaluracarus) deltensis Rosso de Ferrads, 1984
Viets, 1987: 986.
Chaq; ln.
Arrenurus (Megaluracarus) diabolus Lundblad, 1938
Viets, 1987: 66.
Par; Chaq; lt; ln.
Arrenurus (Megaluracarus) difiurus Rosso de Ferrads, 1989
Chaq; ln.
Arrenurus (Megaluracarus) diversisetus Viets, 1940
Viets, 1987: 66.
Cace; ln; lt.
Arrenurus (Megaluracarus) donfelipensis Rosso de Ferrads,
1989
Chaq; ln.
Arrenurus (Megaluracarus) einbergeri Mnchberg, 1960
Viets, 1987: 67.
Par; ln.
Arrenurus (Megaluracarus) epimerosus Marshall, 1919
Viets, 1987: 67.
Par; Amaz; Cace; Chaq; lt; ln.
Arrenurus (Megaluracarus) extensus Viets, 1959
Viets, 1987: 67.
Amaz; lt.
Arrenurus (Megaluracarus) formosus Viets, 1954
Viets, 1987: 68.
Amaz; ln.
Arrenurus (Megaluracarus) fuhrmanni Walter, 1912
Viets, 1987: 68.
Cv; Amaz; Chaq; ln; lt.

Arrenurus (Megaluracarus) funneliforme Rosso de Ferrads &
Bttger, 1997.
Par; ln.
Arrenurus (Megaluracarus) gibberifer Viets, 1933
Viets, 1987: 68.
Par; Chaq; lt.
Arrenurus (Megaluracarus) gibberimarginatus Viets, 1954
Viets, 1987: 68.
Amaz; ln; lt.
Arrenurus (Megaluracarus) gladiiferus Lundblad, 1941
Par; ln; lt.
Arrenurus (Megaluracarus) gracilipalpis Lundblad, 1937
Viets, 1987: 70.
Par; ln.
Arrenurus (Megaluracarus) hamulatus Viets, 1954
Viets, 1987: 71.
Amaz; ln.
Arrenurus (Megaluracarus) hansvietsi Viets, 1953
Viets, 1987: 71.
Pun; ln.
Arrenurus (Megaluracarus) holopygus Lundblad, 1938
Viets, 1987: 71.
Par; Chaq; ln; lt.
Arrenurus (Megaluracarus) illudens Lundblad, 1937
Par; Amaz; Cace; Chaq; ln; lt.
Arrenurus (Megaluracarus) immanis Viets, 1954
Viets, 1987: 71.
Amaz; ln.
Arrenurus (Megaluracarus) inchoatus Viets, 1954
Viets, 1987: 71.
Cace; ln; lt.
Arrenurus (Megaluracarus) infractus Viets, 1937
Viets, 1987: 72.
Cace; lt.
Arrenurus (Megaluracarus) inornatus Viets, 1937
Viets, 1987: 72.
Cace; ln.
Arrenurus (Megaluracarus) lamellatus Lundblad, 1937
Viets, 1987: 72.
Par; ln; lt.
Arrenurus (Megaluracarus) lateriangulatus Lundblad, 1944
Viets, 1987: 73.
Par; Cace; lt.
Arrenurus (Megaluracarus) longimaxillaris Lundblad, 1937
Viets, 1987: 74.
Par; ln.
Arrenurus (Megaluracarus) longisetus Lundblad, 1936
Viets, 1987: 74.
Par; ln.
Arrenurus (Megaluracarus) macrocercus Viets, 1954
Viets, 1987: 74.
Amaz; lt; ln.
Arrenurus (Megaluracarus) maderius Marshall, 1919
Viets, 1987: 74.
Amaz; lt.
Arrenurus (Megaluracarus) malleus Lundblad, 1938
Viets, 1987: 74.
Par; ln; lt.

Arrenurus (Megaluracarus) mediopunctatus Viets, 1936
Viets, 1987: 75.
Par; Cace; ln.
Arrenurus (Megaluracarus) merrilli Marshall, 1919
Viets, 1987: 76.
Amaz; ln.
Arrenurus (Megaluracarus) mirabilis Viets, 1938
Viets, 1987: 76.
Cace; ln.
Arrenurus (Megaluracarus) montanus Lundblad, 1941
Viets, 1987: 76.
Yup; ln.
Arrenurus (Megaluracarus) neuquenensis Rosso de Ferrads,
1987
Aap; ln.
Arrenurus (Megaluracarus) nitidus Viets, 1937
Viets, 1987: 77.
Cace; ln.
Arrenurus (Megaluracarus) nudicaudatus Viets, 1954
Viets, 1987: 77.
Amaz; Chaq; ln; lt.
Arrenurus (Megaluracarus) parvicaudatus parvicaudatus
Lundblad, 1937
Viets, 1987: 78.
Par; ln.
Arrenurus (Megaluracarus) parvicaudatus sinusoides Lundblad,
1944
Viets, 1987: 78.
Par; ln.
Arrenurus (Megaluracarus) paucisetus Lundblad, 1941
Viets, 1987: 78.
Par; ln.
Arrenurus (Megaluracarus) pepefernandezi nomen novum
(Ver Comentarios)
Viets, 1987: 64.
Par; ln.
Arrenurus (Megaluracarus) perlongus Viets, 1936
Viets, 1987: 79.
Amaz; Cace; ln; lt.
Arrenurus (Megaluracarus) productus Viets, 1936
Viets, 1987: 79.
Cace; Chaq; ln; lt.
Arrenurus (Megaluracarus) pugiunculatus Viets, 1954
Viets, 1987: 80.
Amaz; ln.
Arrenurus (Megaluracarus) quadricornicus Marshall, 1919
Viets, 1987: 80.
Amaz; lt.
Arrenurus (Megaluracarus) quadrituberculatus Viets, 1937
Viets, 1987: 80.
Cace; Chaq; ln; lt.
Arrenurus (Megaluracarus) recifensis Viets, 1954
Viets, 1987: 80.
Cace; ln.
Arrenurus (Megaluracarus) ribagai Lundblad, 1936
Viets, 1987: 80.
Par; ln.
Arrenurus (Megaluracarus) santafesinus Rosso de Ferrads, 1973
Viets, 1987: 81.
Chaq; ln.
Arrenurus (Megaluracarus) scopularis Lundblad, 1938
Viets, 1987: 81.
Par; ln; lt.
215
Arrenurus (Megaluracarus) sextuberculatus Viets, 1937
Viets, 1987: 82.
Cace; ln.
Arrenurus (Megaluracarus) solitarius Viets, 1968
Viets, 1987: 82; Cook, 1988: 189.
Csc; ln.
Arrenurus (Megaluracarus) spathulifer Lundblad, 1936
Viets, 1987: 82.
Par; Chaq; ln.
Arrenurus (Megaluracarus) spinicalcar Lundblad, 1937
Viets, 1987: 82.
Par; ln.
Arrenurus (Megaluracarus) tamboensis Lundblad, 1953
Viets, 1987: 83.
Yup; ln.
Arrenurus (Megaluracarus) taurus Viets, 1954
Viets, 1987: 83.
Amaz; ln.
Arrenurus (Megaluracarus) tenuicollis tenuicollis Viets, 1936
Viets, 1987: 83.
Cace; lt.
Arrenurus (Megaluracarus) tenuicollis acornutus Viets, 1968
Viets, 1987: 83.
Csc; ln.
Arrenurus (Megaluracarus) tenuicollis gibberipalpis Lundblad,
1944
Viets, 1987: 83.
Par; Cace; ln.
Arrenurus (Megaluracarus) tenuicollis megacercus Viets, 1954
Viets, 1987: 83.
Cace; Chaq; ln; lt.
Arrenurus (Megaluracarus) tenuicollis megaluroides Lundblad,
1937
Viets, 1987: 83.
Par; Cace; ln.
Arrenurus (Megaluracarus) tenuicollis orthocercus Lundblad,
1944
Viets, 1987: 83.
Par; Cace; ln.
Arrenurus (Megaluracarus) triconicus Marshall, 1919
Viets, 1987: 83.
Amaz; Cace; Par; Cv; Chaq; ln; lt.
Arrenurus (Megaluracarus) tumidus Lundblad, 1938
Viets, 1987: 84.
Par; ln; lt.
Arrenurus (Megaluracarus) uncatus Daday, 1905
Viets, 1987: 84.
Par; Cace; ln; lt.
Arrenurus (Megaluracarus) undulatus Lundblad, 1937
Viets, 1987: 84.
Par; ln.
Arrenurus (Megaluracarus) undulicauda Lundblad, 1941
Viets, 1987: 84.
Par; lt; ln.
Arrenurus (Megaluracarus) unisetiger Viets, 1959
Viets, 1987: 84.
Amaz; lt.
Arrenurus (Megaluracarus) verrucifer Viets, 1954
Viets, 1987: 84.
Amaz; ln.
Arrenurus (Megaluracarus) willinki Rosso de Ferrads, 2000
Chaq; lt.
216
Subgnero Truncaturus Thor, 1901
AGRADECIMIENTOS
Arrenurus (Truncaturus) ivani Rosso de Ferrads, 1998

Par; lt.
Arrenurus (Truncaturus) masonus Cook, 1980
Viets, 1987: 100.
Yup; lt.
Arrenurus (Truncaturus) tucumanensis Cook, 1980
Viets, 1987: 103.
Yup; lt.
Arrrenurus (Truncaturus) yartesus Cook, 1980
Viets, 1987: 103.
Yup; lt.
Al Dr. A. Valdecasas por alentarnos a finalizar esta tarea y

brindndonos la posibilidad de publicar en una revista espaola. A los Dres. R. Gerecke, P. Martin, T. Goldschmidt y M.
Cimpean por sus rpidas respuestas a nuestras consultas y
pedidos. A los Dres. E. Lavilla y L. Acosta por su ayuda con
algunas dudas nomenclaturales. A la Dra. H. Proctor por su
ayuda con el ingls y su aliento en esta empresa. Al CONICET
por financiar nuestras investigaciones. A Jos (Pepe)
Fernndez por su paciencia en las tareas de edicin.
GNERO Micruracaropsis Viets, 1939

Micruracaropsis phytotelmaticola (Viets, 1939)
Viets, 1987: 512.
Amaz; fit.
GNERO Thoracaphoracarus Viets, 1914
Subgnero Thoracaphoracarus Viets, 1914
Thoracaphoracarus (Thoracaphoracarus) simplex Cook, 1988
Cook, 1988: 186.
Csc; lt.
Subgnero Xenthoracaphorus Cook, 1988
Thoracaphoracarus (Xenthoracaphorus) chilensis Cook, 1988
Cook, 1988: 187.
Csc; lt.
Comentarios
HOMONIMIAS HALLADAS EN EL PRESENTE ELENCO DE

ESPECIES. Cramer y Cook (1992) realizaron un
estudio de las especies de Dadayella y como conclusin del mismo, cambiaron el estatus del gnero
Dadayella a subgnero de Arrenurus. Al efectuar
este cambio nomenclatural, dos especies de
Arrenurus quedaron en homonimia con dos de
Dadayella: ellas son, Arrenurus boettgeri Viets,
1968 y A. (Megaluracarus) clavipes Lundblad,
1941. Esta situacin fue advertida en la elaboracin
de este trabajo por D. Jos Fernndez. Por tal motivo como A. (Dadayella) boettgeri Viets & Bttger,
1986, es un homnimo primario de A. boettgeri,
proponemos el nombre de Arrenurus (Dadayella)
guarani para reemplazarla.
Por otra parte, A. (Megaluracarus) clavipes
Lundblad es un homnimo primario de A.
(Dadayella) clavipes (Lundblad, 1938) proponindose el nombre de A. (Megaluracarus) pepefernandezi para reemplazarlo.
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Crdoba (Argentina). III. Revista de la Sociedad
Entomolgica Argentina, 58(3-4): 109-127.
ROSSO DE FERRADS, B. & SMIT, H., 1998. Nuevos
hidracridos (Acari: Hydrachnidia) relacionados con
la cuenca del Paran, Argentina. Revista de la
SCHWOERBEL, J., 1986. Rheophile Wassermilben aus
Chile 2. Die Gattungen Rhynchaturus Besch und
Paraspidiobates nov. gen. (Acari: Hygrobadidae).
Archiv fr Hidrobiologie, 106: 233-244.

SCHWOERBEL, J., 1987. Rheophile Wassermilbe (Acari:
Hydrachnellae) aus Chile 3. Arten aus
Thermalgewssern. Archiv fr Hydrobiologie, 110:
399-407.
SMIT, H., 1997. Australian water mites of the genus
Arrenurus, with the description of twelve new species from northern and western Australia (Acari:
Hydrachnellae: Arrenuridae). Records of the Western
Australian Museum, 18: 233-261.
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Argentina, with the description of one new genus and
three new species (Acari: Hydrachnidia). Zootaxa,
103: 1-23.
SMITH, I. M. & COOK, D. R., 1994. North American species of Neomamarsinae Lundblad (Acari: Hydrachnida: Limnesiidae). The Canadian Entomologist,
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Recibido, 12-I-2005
Aceptado, 12-XI-2005
Publicado, 31-XII-2005
219
NDICE ALFABTICO DE NOMBRES CIENTFICOS
Las categoras supragenricas se indican en maysculas, los gneros y subgneros en negrita y las especies y subespecies en cursiva.
aberratus .................................. 198
abnormipes .............................. 208
acanthifera ................................ 204
Acantholimnesia ...................... 193
acanthoscelus .......................... 193
acornutus .................................. 215
Actinacarus .............................. 195
actinophora .............................. 211
aculeata, Anisitsiella ................ 188
aculeata, Neoaxona .................. 208
aculeatipalpis .......................... 206
acuta ........................................ 202
acutidens, Corticacarus .......... 197
acutidens, Koenikea ................ 204
acutidens, Piona ...................... 206
acutidens, Unionicola .............. 201
acutidentatus ............................ 211
acutiepimeratus ........................ 212
acutipes .................................... 206
acutirostris ................................ 189
adulterinus ................................ 213
aequalipilis .............................. 189
affinis, Actinacarus .................. 195
affinis, Axonopsella .................. 207
affinis, Corticacarus ................ 196
affinis, Koenikea ...................... 202
affinis, Mideopsis .................... 210
agilis ........................................ 184
alatus ........................................ 208
Albia ........................................ 209
ALBIINAE .............................. 209
alienus ...................................... 186
allarka ...................................... 189
allasa ........................................ 189
Allolimnesia ............................ 193
alphus ...................................... 200
alta ............................................ 189
alza .......................................... 209
amabilis .................................... 205
Amazonella .............................. 202
amazonica, Geayia .................. 211
amazonica, Limnesia ................ 192
amazonica, Nilotonia .............. 189
amazonica, Piona .................... 206
amazonicola .............................. 193
amazonicus .............................. 213
ampliatus .................................. 194
amplipalpis .............................. 194
amplipenicillatus ...................... 212
amplipora ................................ 192
ampullariae .............................. 201
Ampullariatax .......................... 201
anchista .................................... 209
Anchistalbia ............................ 209
anchistus .................................. 197
andalienensis ............................ 190

Andesobates ............................ 200
andiana .................................... 187
andina, Axonopsella ................ 207
andina, Limnesia ...................... 192
andina, Rhyncholimnochares .... 185
andinus, Aturus ........................ 207
andinus, Sperchon .................... 188
andinus, Thermacarus .............. 188
andrewfieldi .............................. 213
angelescui ................................ 197
angelieri, Limnesia .................. 192
angelieri, Rutacarus ................ 189
angulata .................................... 192
angulicauda .............................. 213
angulipes .................................. 205
angulosus .................................. 213
angustipalpis, Koenikea .......... 202
angustipalpis, Limnesia ............ 193
angustipalpis, Unionicola ........ 201
angustiporus ............................ 195
angustus .................................... 198
anisitsi, Arrenurus .................... 212
anisitsi, Eylais .......................... 185
anisitsi, Piona .......................... 206
Anisitsiella ................................ 188
ANISITSIELLIDAE ................ 188
Anisitsiellides .......................... 188
ANISITSIELLINAE ................ 188
ANKELOTHYADINAE .......... 187
Ankelothyas .............................. 187
Apeltosperchon ........................ 188
APELTOSPERCHONTINAE .. 188
apertulus .................................. 194
apertus, Arrenurus .................... 212
apertus, Hygrobates ................ 194
aphanoraphtha ........................ 189
apophylonga ............................ 190
aranda ...................................... 191
archangelskii ............................ 213
arcuata ...................................... 199
argentinensis, Axonopsella ...... 208
argentinensis, Corticacarus .... 197
argentinensis, Lethaxonella ...... 208
argentinensis,
Tetrahygrobatella ................ 200
armata ...................................... 185
armipes .................................... 204
arpodos .................................... 204
ARRENURIDAE .................... 211
arrenuripalpis, Notoaturus ...... 209
arrenuripalpis,
Phreatomideopsis ................ 210
arrenuripalpis, Plaumannia .... 211
ARRENUROIDEA .................. 209
Arrenurus ........................ 211, 212

Arrhenuropsides ...................... 213
Arrhenuropsis .......................... 213
aspida ...................................... 191
Aspidiobates ............................ 195
assimilis .................................... 203
asterifera, Clathrosperchonella .. 187
asterifera, Koenikea ................ 203
Atacella .................................... 201
ATHIENEMANNIIDAE .......... 211
Atractidella .............................. 195
atractidellides .......................... 195
Atractides ................................ 195
atrorubra .................................. 206
ATURIDAE .............................. 206
Aturides .................................... 207
ATURINAE .............................. 207
Aturus ...................................... 207
Australatax .............................. 201
Australiobatella ........................ 195
Australiobates .......................... 196
australis, Anisitsiellides ............ 188
australis, Tyrrellia .................... 190
avinotata .................................. 211
Axonopsella .............................. 207
AXONOPSINAE ...................... 207
Axonopsis ................................ 208
aymara ...................................... 192
bachmanni ................................ 213
balteata .................................... 205
barbifera .................................. 204
barri .......................................... 207
beschi, Mixolimnesia ................ 191
beschi, Notoaturus .................... 209
beschi, Notosperchonopsis ...... 188
beschi, Rhynchaturus .............. 200
besselingi .................................. 192
bicornis .................................... 202
bidens ........................................ 191
bidentata, Limnesia .................. 193
bidentata, Neoalbia .................. 208
bifida ........................................ 208
bilaciniatus .............................. 213
bipapillata ................................ 202
bipetiolatus .............................. 213
biscutata, Atractidella .............. 195
biscutata, Koenikea .................. 202
bisinuosus ................................ 213
bituberculata ............................ 204
bituberculatus .......................... 213
biverrucata .............................. 210
bodus ........................................ 198
boettgeri, Arrenurus ................ 212
boettgeri, Arrenurus
(Dadayella) ...................... 216
Bolivartonia ............................ 189

bonariensis .............................. 202
bondi ........................................ 193
bonettoi .................................... 213
boopis ...................................... 194
bordoni .................................... 208
boutata ...................................... 208
bovala ...................................... 200
Brachypodopsis ........................ 208
brasiliana .................................. 193
brasiliensis, Arrenurus ............ 212
brasiliensis, Atractides ............ 195
brasiliensis, Eylais .................. 185
brasiliensis, Koenikea .............. 204
brasiliensis, Sperchon .............. 188
brasiliensis, Unionicola .......... 201
brassanus .................................. 197
brauni ...................................... 192
braytola .................................... 192
Brevaturus ................................ 196
Brevicaudaturus ...................... 213
brevicaudatus .......................... 213
brevipalpis ................................ 185
brevipetiolatus .......................... 212
brevirostris ................................ 187
brevis, Recifella ........................ 205
brevis, Torrenticola .................. 190
breviscuta ................................ 185
breviseta .................................. 204
brevisuturata ............................ 201
brogavus .................................. 197
brundini .................................... 197
brunnea .................................... 202
cachoerensis ............................ 213
Callumobates .......................... 196
Camposea ................................ 196
cancellatus ................................ 200
caquetiorum .............................. 212
cardioporus .............................. 195
caripense .................................. 208
cataglyphus .............................. 213
cataphracta .............................. 188
catharinensis, Arrenurus .......... 213
catharinensis, Geayia .............. 211
caudatus .................................... 207
cautinensis ................................ 200
caviventris ................................ 210
cekalovici .................................. 196
Centrolimnesia ........................ 193
chechoi ...................................... 186
Chilaxonopsella ...................... 207
chilensis, Anisitsiellides ............ 188
chilensis, Axonopsella .............. 207
chilensis, Brevaturus ................ 196
chilensis, Corticacarus ............ 197
220
chilensis, Frontipodopsis .......... 206
chilensis, Hydrachna ................ 184
chilensis, Koenikea .................. 204
chilensis, Limnesia .................. 192
chilensis, Oxus .......................... 189
chilensis, Piona ........................ 206
chilensis, Tetrahygrobatella .... 200
chilensis, Thoracaphoracarus .. 216
chillanensis .............................. 198
chiloensis .................................. 209
choconensis .............................. 209
chorreronica ............................ 191
ciliata ........................................ 189
circularis, Illiesiella ................ 188
circularis, Rhynchaturus .......... 200
circumcincta, Koenikea ............ 204
circumcincta, Limnesia ............ 192
circummarginata ...................... 204
clanopus .................................... 200
clathrata .................................. 201
Clathrosperchon ...................... 187
Clathrosperchonella ................ 187
CLATHROSPERCHONTINAE
............................................ 187
clavata ...................................... 205
clavigera .................................. 203
clavilaminata ............................ 213
clavipes, Arrenurus
(Dadayella) .......................... 213
clavipes, Arrenurus
(Megaluracarus) .................. 216
clavipes, Hydrodroma .............. 187
clavipes, Mideopsis .................. 209
clavipes, Torrenticola .............. 190
clevamus .................................. 194
coarctipalpis ............................ 196
Coaxonopsella .......................... 207
coeruleocruciata ...................... 211
coeruleolineata ........................ 211
coerulescens ............................ 202
coeruleus .................................. 196
coldomus .................................. 197
colpophthalma .......................... 185
columbiana, Recifella .............. 205
columbiana, Torrenticola ........ 190
columbianus, Atractides .......... 195
columbianus, Corticacarus ...... 196
columbianus, Paraschizobates .. 199
columbicola .............................. 208
columbiensis ............................ 185
cometes .................................... 212
condiscipulorum ...................... 199
confertus .................................. 212
confinis .................................... 206
conimaxillaris .......................... 211
conirostris ................................ 190
conjuncta .................................. 201
consaguineus ............................ 213
consimilis .................................. 204
conspicuos ................................ 212

constricta .................................. 184
convexa, Koenikea .................... 202
convexa, Krendowskia .............. 210
copinae .................................... 197
coriacea, Atractidella .............. 195
coriacea, Hygrobatella
(Hygrobatella) ...................... 198
coriacea, Hygrobatella
(Hygrobatella) puberula ...... 199
cornicaudatus .......................... 214
corniculata ................................ 202
cornifrons ................................ 214
corniger .................................... 214
corpulentus .............................. 214
correntinae .............................. 214
Corticacarellus ........................ 197
Corticacarides .......................... 197
Corticacarus ............................ 196
coscaroni .................................. 186
Crameratax .............................. 201
crassa ........................................ 204
crassipalpis, Amazonella .......... 202
crassipalpis, Clathrosperchon .. 187
crassipalpis, Corticacarus ...... 196
crassipalpis, Koenikea ............ 203
crassipalpis,
Notosperchonopsis .............. 188
crassipalpis, Schadeella .......... 205
crassipes, Oxus ........................ 189
crassipes, Atractides ................ 195
crassipes, Hygrobates .............. 194
crassipes, Neotorrenticola ........ 191
crassipes, Notomomonia .......... 209
crassipes, Submiraxona ............ 209
crassiseta, Hydrodroma ............ 187
crassiseta, Koenikea ................ 204
crassiseta, Limnesia ................ 192
crassoalata .............................. 203
crassus ...................................... 214
crawfordi .................................. 185
crenicauda ................................ 214
Crenohygrobates ...................... 198
Crenolimnesia .......................... 191
crenophila, Plaumannia .......... 211
crenophila, Tyrrellia ................ 190
critus ........................................ 200
cruciata, Recifella .................... 205
cruciata, Roqueella .................. 211
cultriger .................................... 214
curicola .................................... 204
Curryatax ................................ 201
curticaudatus ............................ 214
curtipalpis ................................ 196
curvimarginata ........................ 202
curvipalpis, Arrenurus .............. 213
curvipalpis, Hygrobates .......... 194
curvipalpis, Koenikea .............. 204
curvipes, Koenikea .................. 204
curvipes, Mideopsis .................. 210
curvipes, Neumania .................. 205
curvirostris, Corticacarus ........ 197

curvirostris, Koenikea .............. 203
curvitarsis ................................ 201
cyanozona ................................ 203
Dadayella ................................ 213
davecooki .................................. 196
davidcooki, Arrenurus .............. 214
davidcooki, Piona .................... 206
Decaroqueella .......................... 211
Decussobates ............................ 198
deformis .................................... 206
dejecta ...................................... 206
deltensis .................................... 214
dentata ...................................... 202
dentatus .................................... 207
dentifera, Mideopsis ................ 209
dentifera, Polyaxonopsella ...... 208
dentipalpis, Arrenurus .............. 212
dentipalpis, Corticacarus ........ 196
dentipalpis, Limnesia .............. 192
dentipes .................................... 207
desajunos .................................. 186
despiciens ................................ 187
diabolica .................................. 204
diabolus .................................... 214
diamphida ................................ 190
Diamphidaxona ...................... 198
dictyoderma .............................. 200
difiurus ...................................... 214
digitatus .................................... 200
dipersiai .................................... 185
Diplohydrachna ...................... 184
Diplokoenikea .......................... 203
diplothoracica .......................... 205
discrepans ................................ 194
disjunctus .................................. 200
dissipata .................................... 206
distendens, Eylais .................... 185
distendens, Hygrobates ............ 195
distorta ...................................... 192
diversidentatus .......................... 194
diversisetus .............................. 214
dividuus .................................... 187
divisus, Corticacarus ................ 197
divisus, Corticacarus
multiscutus .......................... 197
Dodecabates ............................ 198
dodecaporus ............................ 198
dolichopalpis ............................ 189
doloma ...................................... 207
donfelipensis ............................ 214
dorsalis .................................... 192
dorsolineata .............................. 210
dorsoscutata ............................ 205
Dubiobates .............................. 198
dubiosa .................................... 192
duricoria .................................. 193
edentulus .................................. 196
einbergeri ................................ 214
elata .......................................... 199
elegans ...................................... 204

elegantula ................................ 199
elliptica .................................... 205
elongata .................................... 210
emydoides ................................ 187
entrerrianensis .......................... 201
Eocorticacarus ........................ 198
Eorecifella ................................ 205
Eorutacarus ............................ 188
epimeratus ................................ 191
epimerosus ................................ 214
erectipetiolatus ........................ 212
eremita ...................................... 191
erratica .................................... 206
Eupatrella ................................ 186
EUPATRELLINAE .................. 186
Euwandesia .............................. 187
evolus ........................................ 198
excavata .................................... 205
excavatoides ............................ 205
exigua ...................................... 192
exilis .......................................... 196
expansipes ................................ 209
expansiseta .............................. 185
extensus .................................... 214
EYLAIDAE .............................. 185
Eylais ........................................ 185
EYLOIDEA .............................. 185
faceta ........................................ 190
facetus ...................................... 200
falcaria .................................... 202
falcipalpis ................................ 190
falsificata .................................. 192
ferradasae, Omartacarus ........ 194
ferradasae, Rutacarus .............. 188
Ferradasatax ............................ 201
Ferradasia ................................ 200
FERRADASIIDAE .................. 200
filunguis .................................... 208
fissa .......................................... 206
fissipalpis .................................. 201
fissipes ...................................... 201
flabellifera ................................ 211
Flabellifrontipoda .................... 189
flabellum .................................. 199
flagellaris .................................. 203
flagellatus ................................ 212
flagellifera ................................ 211
flosculipora .............................. 192
fluctuata .................................... 202
forficipalpis .............................. 210
formosus, Arrenurus ................ 214
formosus, Neoatractides .......... 189
frijolito ...................................... 186
frontalis .................................... 210
Frontipoda .............................. 189
FRONTIPODOPSINAE .......... 206
Frontipodopsis ........................ 206
fuhrmanni, Arrenurus .............. 214
fuhrmanni, Limnesia ................ 192

funneliforme .............................. 214
furcaticornis ............................ 204
furcatimaculata ........................ 203
gaethgensi ................................ 205
Geayella .................................... 211
Geayia ...................................... 211
Geayidea .................................. 211
geijskesi .................................... 194
geniculata ................................ 194
genitalis .................................... 210
gennadus .................................. 199
geometricus .............................. 196
gereckei .................................... 191
gessneri .................................... 192
gibberifer .................................. 214
gibberimarginatus .................... 214
gibberipalpis, Arrenurus .......... 215
gibberipalpis, Mideopsis .......... 210
gibberipalpis, Recifella ............ 205
gigantea .................................... 201
gladiiferus ................................ 214
Gledhillatax .............................. 201
gledhilli, Brevaturus ................ 196
gledhilli, Zabobates .................. 200
glieschi ...................................... 185
globipalpis ................................ 207
godeti ........................................ 186
goliath ...................................... 213
gomorus .................................... 196
gonseri ...................................... 196
gracilidens ................................ 194
gracilipalpis, Arrenurus .......... 214
gracilipalpis, Oxus .................. 189
gracilipes .................................. 201
gracilirostris ............................ 185
gradatipalpis ............................ 210
granuligera .............................. 192
grossa ...................................... 203
guacharensis ............................ 198
guarani, Arrenurus .......... 213, 216
guarani, Centrolimnesia .......... 194
guttifera .................................... 213
hamata, Atractidella ................ 195
hamata, Axonopsella ................ 208
hamatoides ................................ 208
hamulata .................................. 204
hamulatus ................................ 214
hansvietsi .................................. 214
harveyi ...................................... 195
hastata ...................................... 203
henrechi .................................... 207
hesperia, Limnesia .................. 192
hesperia, Torrenticola .............. 190
Heteratractides ........................ 190
heterophthalma ........................ 184
hexagona .................................. 192
hexaporus ................................ 200
Hexaroqueella .......................... 211
hiporreica ................................ 198
hirsutipalpis .............................. 212
hirtipalpis ................................ 213

Hispidosperchon ...................... 188
holopygus .................................. 214
honoratus .................................. 212
hopkinsi .................................... 209
horrida ...................................... 204
Humboldtaxonopsella .............. 207
Hydrachna .............................. 184
HYDRACHNIDAE .................. 184
HYDRACHNOIDEA .............. 184
Hydrodroma ............................ 187
HYDRODROMIDAE .............. 187
Hydryphantes .......................... 185
HYDRYPHANTIDAE ............ 185
HYDRYPHANTINAE ............ 185
HYDRYPHANTOIDEA .......... 185
Hygrobatella ............................ 198
Hygrobates .............................. 194
HYGROBATIDAE .................. 194
Hygrobatides ............................ 195
HYGROBATINAE .................. 194
HYGROBATOIDEA ................ 190
hygropetrica ............................ 189
icotus ........................................ 197
iheringi .................................... 200
Illiesiella .................................. 188
illudens .................................... 214
imitata ...................................... 204
immanis .................................... 214
imperator .................................. 213
inachus ...................................... 190
inchoata .................................... 211
inchoatus .................................. 214
incisa ........................................ 200
inconspicua .............................. 206
incurvatus ................................ 196
inermis ...................................... 201
inflatipalpis .............................. 212
infractipalpis ............................ 210
infractus .................................... 214
inka .......................................... 205
inornatus .................................. 214
intercursor ................................ 212
interpositus .............................. 194
interstitialis .............................. 191
ischiotricha .............................. 188
ivani .......................................... 216
jemobus .................................... 196
jujuyensis, Axonopsella ............ 207
jujuyensis, Hydryphantes ........ 185
juncta ........................................ 206
Jurucuia .................................. 203
karinae ...................................... 199
karlvietsi .................................. 208
keldomus .................................. 186
klaasseni .................................. 196
Koenikea .................................. 202
Koenikella ................................ 203
kolus .......................................... 198
Kongsbergia ............................ 207
221
Krendowskia ............................ 210
Krendowskiella ........................ 210
KRENDOWSKIIDAE .............. 210
kronestedti ................................ 207
kurtvietsi .................................. 196
Kyphohygrobatella .................. 199
Kyphohygrobates .................... 199
labialis ...................................... 197
laceripalpis .............................. 210
laeta .......................................... 192
lamellatus ................................ 214
Lamellaxona ............................ 208
lamellipalpis ............................ 185
laminipes .................................. 205
Lasalleatax .............................. 201
lata ............................................ 203
lateriangulatus .......................... 214
lateribarbata ............................ 205
latidens .................................... 197
latigenitalis .............................. 192
latipes ...................................... 205
latirostris .................................. 211
LEBERTIOIDEA ...................... 188
lemba ........................................ 191
lembus ...................................... 196
lenzi .......................................... 211
leptoglyphus .............................. 209
lethaea ...................................... 186
Lethaxonella ............................ 208
levipalpis .................................. 201
Limnesia .................................. 192
Limnesides .............................. 191
Limnesiella .............................. 193
Limnesiellula .......................... 193
LIMNESIIDAE ........................ 190
LIMNESIINAE ........................ 192
Limnesiopsides ........................ 193
LIMNOCHARIDAE ................ 185
linaresi ...................................... 207
lineata ...................................... 210
litatus ........................................ 196
loaensis .................................... 188
lobatus ...................................... 212
longa ........................................ 191
longidens, Atractidella ............ 195
longidens, Mideopsis ................ 210
longidens, Unionicola .............. 202
longigenitalis ............................ 199
longimanus .............................. 195
longimaxilaris .......................... 211
longimaxillaris, Arrenurus ...... 214
longimaxillaris, Megapella ...... 199
longimaxillaris, Neocalonyx .... 186
longipalpis, Andesobates .......... 200
longipalpis, Neocalonyx .......... 186
longipalpis,
Rhyncholimnochares ............ 185
longipalpis, Stylohygrobates .... 200
longipalpis, Tetrahygrobatella .. 200
longipes, Hygrobates ................ 195
longipes, Koenikea .................. 203

longipes, Tyrrellia .................... 190
longipes, Unionicola ................ 201
longipora .................................. 192
longirostris, Corticacarus ........ 197
longirostris, Limnesia .............. 192
longiscuta ................................ 185
longiseta .................................. 187
longisetus .................................. 214
longiuscula .............................. 202
longivalvata .............................. 192
loretoensis ................................ 193
loricata .................................... 205
Luciaxonopsella ...................... 207
Lundbladacarus ...................... 197
Lundbladatax .......................... 201
lundbladi, Anisitsiellides .......... 188
lundbladi, Centrolimnesia ........ 194
lundbladi, Hygrobates .............. 195
lundbladi, Recifella .................. 205
lundbladi, Roqueella ................ 211
lundbladi, Szalayella ................ 200
lunipes ...................................... 204
maceripalpis ............................ 188
macrocercus .............................. 214
macronyx .................................. 187
maculata, Mideopsis ................ 210
maculata, Roqueella ................ 211
maderius .................................. 214
magna ...................................... 195
magniacetabulae ...................... 195
magnifica .................................. 191
magnipora ................................ 193
malacoderma ............................ 193
malleifer .................................... 196
malleus ...................................... 214
Mamersellides .......................... 188
Mamersonia ............................ 189
mamillata .................................. 208
mandibularis ............................ 207
Mapuchacarus ........................ 199
masonus .................................... 216
mediolineata ............................ 211
medioposita .............................. 203
mediopunctatus ........................ 215
megacercus .............................. 215
megalopsis ................................ 210
Megaluracarus ........................ 213
megaluroides ............................ 215
Megapella ................................ 199
megapoides .............................. 199
melini ........................................ 205
Meramecia .............................. 190
merrilli ...................................... 215
mexicana .................................. 190
microdactylus .......................... 210
microdon .................................. 193
micronycha .............................. 211
micronyx .................................. 187
Micruracaropsis ...................... 216
222
Mideopsella .............................. 210
MIDEOPSIDAE ...................... 209
Mideopsides .............................. 210
Mideopsis ................................ 209
miliaria .................................... 184
millepora .................................. 193
minima ...................................... 213
minor ........................................ 187
minuscula .................................. 192
minuta ...................................... 199
minutidentata ............................ 192
minutissimus ............................ 197
minutus .................................... 198
mirabilis .................................... 215
Miraxonidella .......................... 208
Miraxonides ............................ 208
misionum .................................. 207
Mitchellatax ............................ 201
Mixolimnesia .......................... 191
MIXOLIMNESIINAE .............. 191
Mixosperchon .......................... 188
Momoniella .............................. 209
MOMONIIDAE ...................... 209
Monatractides .......................... 190
monochroa ................................ 211
Monokoenikea ........................ 205
montana, Eylais ........................ 185
montana, Hygrobatella ............ 199
montanus .................................. 215
monticolus ................................ 188
motasi, Corticacarus ................ 197
motasi, Rheolimnesia .............. 191
motasi, Sperchon ...................... 188
motasi, Unionicola .................. 201
Motasia .................................... 199
multangulus .............................. 212
multiacetabulata,
Hygrobatella ........................ 199
multiacetabulata,
Tubophorella ........................ 194
multipapillatus .......................... 212
multiporus, Corticacarus ........ 198
multiporus, Crenohygrobates .... 198
multiporus, Neoaturus .............. 207
multiscutata .............................. 188
multiscutus ................................ 197
multiseta .................................. 204
multispina ................................ 185
musicola .................................... 200
mystrophorus ............................ 212
nana .......................................... 208
Neoalbia .................................. 208
Neoalbiella .............................. 208
Neoatractides .......................... 189
NEOATRACTIDINAE ............ 189
Neoaturus ................................ 207
Neoaxona ................................ 208
Neoaxonopsella ........................ 208
Neoaxonopsis .......................... 208
Neocalonyx .............................. 186

Neocorticacarus ...................... 199
Neokoenikea ............................ 204
Neokrendowskia ...................... 211
Neolimnochares ...................... 185
Neomamersa ............................ 190
NEOMAMERSINAE .............. 190
Neoneumania .......................... 204
neopuberula .............................. 199
Neoroqueella ............................ 211
Neorutacarus .......................... 189
Neotorrenticola ........................ 191
NEOTORRENTICOLINAE .... 191
neotropica ................................ 189
neotropicus .............................. 188
Neotyrrellia .............................. 190
Neoxystonotus .......................... 210
Neumania ................................ 204
neuquenensis, Arrenurus .......... 215
neuquenensis,
Notosperchonopsis .............. 188
nevadensis ................................ 188
Nilotonia .................................. 189
NILOTONIINAE .................... 189
nilsoni ...................................... 197
nitidus ...................................... 215
nobilis ...................................... 209
Noesaturus .............................. 209
nonlamellata ............................ 184
noodti, Corticacarus ................ 197
noodti, Tyrrellia ........................ 190
nordenskioldi ............................ 204
Notaxona .................................. 209
NOTOATURINAE .................. 209
Notoaturus .............................. 209
Notomideopsis .......................... 204
Notomomonia .......................... 209
Notopanisus ............................ 186
Notosperchonopsis .................. 188
novus, Atractides ...................... 195
novus, Neoaturus ...................... 207
nudicaudatus ............................ 215
obliquua .................................... 185
oblonga .................................... 208
oblongata .................................. 203
obscura .................................... 204
obtusidens, Atractidella ............ 195
obtusidens, Hygrobates ............ 194
ochiepus .................................... 209
octopapillata ............................ 203
octopora .................................... 208
octoporus .................................. 200
odontogaster ............................ 208
ogavus ...................................... 196
OMARTACARIDAE .............. 194
Omartacarus ............................ 194
onophora .................................. 192
opimipalpis .............................. 205
ornatula .................................... 185
orthocercus .............................. 215
orthophthalma .......................... 185
Osornobates ............................ 199

otendus ...................................... 209
Otongacarus ............................ 186
oviformis .................................. 210
ovum, Arrenurus ...................... 212
ovum, Mideopsis ...................... 210
OXIDAE .................................. 189
Oxus ........................................ 189
oxyrhynchus .............................. 194
oxyura ...................................... 206
oxyurus .................................... 212
pachydermis .............................. 195
pachyscelus .............................. 202
pacrodactylus .......................... 210
pallida, Koenikea .................... 203
pallida, Mixolimnesia .............. 191
pampaensis .............................. 191
panniculata .............................. 206
papillata, Hygrobatella ............ 199
papillata, Koenikea .................. 203
papillata, Neotorrenticola ........ 191
papillata, Roquella .................. 211
papilliger .................................. 212
Papilloporus ............................ 186
papillosus .................................. 186
paprzyckii ................................ 210
Paracalonyx ............................ 186
Paracorticacarus ...................... 197
paraelongatus .......................... 194
paraensis .................................. 195
paraguayensis .......................... 203
Parakoenikea .......................... 204
Paralimnesia ............................ 193
Paralimnochares ...................... 185
Parameramecia ........................ 190
Pararoqueella .......................... 211
Paraschizobates ...................... 199
Paraspidiobates ........................ 196
Paraxonopsella ........................ 208
Partnuniella ............................ 186
parva, Frontipoda .................... 189
parva, Limnesia ........................ 192
parvicaudatus .......................... 215
parvipons .................................. 185
parviscutatus ............................ 213
parvula ...................................... 199
patagonica, Frontipoda ............ 189
patagonica, Limnesia .............. 192
patagonicus .............................. 196
paucidens .................................. 207
paucipora, Limnesia ................ 192
paucipora, Neotyrrellia ............ 190
paucipora, Recifella ................ 205
pauciscutata .............................. 188
pauciseta, Koenikea ................ 202
pauciseta, Limnesia .................. 192
paucisetus ................................ 215
paupercula ................................ 205
pectinata .................................. 203
pectinifera ................................ 203
pectunguis ................................ 186

pectungulata ............................ 193
penai, Corticacarus .................. 197
penai, Dubiobates .................... 198
penai, Frontipoda .................... 189
penai, Neocalonyx .................... 186
pennapodus .............................. 212
Pentalbia .................................. 209
pepefernandezi ................ 215, 216
peregrina .................................. 187
pereirai .................................... 198
perforata .................................. 201
perincisa .................................. 185
perlongus .................................. 215
perpusilla, Limnesia ................ 193
perpusilla, Unionicola .............. 201
petricola .................................... 190
petrophila, Neolimnochares .... 185
petrophila, Tyrrellia .................. 190
phreaticola ................................ 196
phreaticus ................................ 209
Phreatomideopsis .................... 210
phytotelmaticola ...................... 216
Pilolimnesia ............................ 193
pina .......................................... 186
pinguipalpis .............................. 186
Piona ........................................ 206
PIONATACINAE .................... 202
PIONIDAE .............................. 206
PIONINAE .............................. 206
Pirapama .................................. 211
placibilis .................................. 191
placoderma .............................. 199
placophora, Crenolimnesia ...... 191
placophora, Hygrobatella ........ 199
placophora, Neolimnochares .... 185
placophorus, Corticacarus ...... 196
placophorus, Neocalonyx ........ 186
planus ...................................... 198
platano ...................................... 186
plaumanni, Arrenurus .............. 212
plaumanni, Atractides .............. 195
plaumanni, Axonopsella .......... 207
plaumanni, Corticacarus .......... 197
plaumanni, Hygrobates ............ 194
plaumanni, Koenikea ................ 202
plaumanni, Limnesia ................ 193
plaumanni, Mideopsis .............. 209
plaumanni, Neoaturus .............. 207
plaumanni, Neoaxona .............. 208
plaumanni, Piona .................... 206
plaumanni, Unionicola ............ 202
Plaumannia .............................. 211
plebejus .................................... 194
plenipalpis ................................ 206
plicata ...................................... 203
plicatus .................................... 194
plumipes .................................... 191
Polyatacides ............................ 202
Polyaxonopsella ...................... 208

Polycorticacarellus .................. 198
polygramma .............................. 199
Polyhydryphantes .................... 186
Polyhygrobatella ...................... 199
polyplacophora ........................ 189
polypora, Amazonella .............. 202
polypora, Limnesia .................. 193
polypora, Neotyrrellia .............. 190
polypora, Polyaxonopsella ...... 208
polypora, Polyhygrobatella ...... 199
polytricha .................................. 205
porophora ................................ 195
porosus, Atractides .................. 195
porosus, Hygrobates ................ 194
portigera .................................. 184
porulosa .................................... 192
Poundsatax .............................. 202
praedentata .............................. 193
praegracilis .............................. 206
pravanus .................................. 197
prehendens ................................ 201
procursa .................................... 203
procursus .................................. 194
procurvipes .............................. 201
productus, Arrenurus ................ 215
productus, Hygrobates .............. 194
projecta .................................... 213
projectus .................................. 207
promacrus ................................ 212
prominens ................................ 202
propinquus ................................ 212
protendens ................................ 185
Protolimnesella ........................ 191
Protolimnesia .......................... 191
PROTOLIMNESIINAE .......... 191
Protzia ...................................... 186
PROTZIINAE .......................... 186
proyectus .................................. 200
Pseudokoenikea ...................... 204
Pseudowandesia ...................... 187
puberula .................................... 199
pugionipalpis ............................ 201
pugiunculata ............................ 185
pugiunculatus .......................... 215
pulchra ...................................... 205
pumilus .................................... 186
punctatissima ............................ 206
punctatus .................................. 187
pusilla, Hydrachna .................. 184
pusilla, Limnesia ...................... 193
pusilla, Recifella ...................... 205
pusuligera ................................ 201
pyriformis ................................ 188
quadricornicus .......................... 215
quadricornuta .......................... 203
quadrimaculata ........................ 210
quadripustulata ........................ 211
quadrisetosus ............................ 212
quadrituberculatus .................. 215
quinquemaculata ...................... 203
quiponensis .............................. 198

radilofus .................................... 195
ramosa ...................................... 203
ramosus .................................... 186
Recifella .................................. 205
recifensis .................................. 215
rectangularis ............................ 210
rectangulata .............................. 203
rectimarginata .......................... 206
rectipes .................................... 204
rectipetiolatus .......................... 212
rectirostris, Hydrachna ............ 184
rectirostris, Koenikea .............. 203
recurva ...................................... 190
recurvus .................................... 198
reducta ...................................... 193
reductus .................................... 197
reflexidens ................................ 206
remotipora, Limnesia .............. 193
remotipora, Mideopsis .............. 210
retractidens, Koenikea .............. 203
retractidens, Unionicola .......... 202
retrocornuta .............................. 203
Rhabdohydrachna .................. 184
Rheolimnesia .......................... 191
RHEOLIMNESIINAE ............ 191
RHYNCHATURINAE ............ 200
Rhynchaturus .......................... 200
RHYNCHOHYDRACARIDAE
............................................ 187
RHYNCHOHYDRACARINAE
............................................ 187
Rhynchohydracarus ................ 187
Rhyncholimnochares .............. 185
ribagai, Amazonella ................ 202
ribagai, Arrenurus .................... 215
ribagai, Limnesia .................... 193
ribagai, Mixolimnesia .............. 191
rimorus .................................... 197
ringueleti .................................. 212
rionegrensis .............................. 213
ripiphorus ................................ 212
robusta ...................................... 187
robustipalpis ............................ 206
Roqueella ................................ 211
rosea ........................................ 202
roseni ........................................ 191
Rostaxonopsella ...................... 208
rostrata, Limnesia .................... 193
rostrata, Mideopsis .................. 210
rostratus, Atractides ................ 195
rostratus, Corticacarus ............ 197
rotunda, Arrenurus .................. 213
rotunda, Piona .......................... 206
rotundiformis ............................ 206
rubicunda .................................. 210
rufolineata ................................ 211
rufus .......................................... 195
rugosa ...................................... 201
Rutacarus ................................ 188
223
rutae, Clathrosperchonella ...... 187
rutae, Koenikea ........................ 204
rutae, Submiraxona .................. 209
salsa .......................................... 186
saltensis .................................... 190
santafesinus .............................. 215
Schadeella ................................ 205
schadei, Atractides .................. 195
schadei, Centrolimnesia .......... 194
schadei, Corticacarus .............. 196
schadei, Neumania .................. 205
schadei, Recifella .................... 205
schindowskii ............................ 186
Schizobates .............................. 200
schmitzi .................................... 188
Schubartella ............................ 195
schubarti, Centrolimnesia ........ 194
schubarti, Geayia .................... 211
schubarti, Koenikea .................. 202
schubarti, Oxus ........................ 189
schubarti, Unionicola .............. 201
schusteri .................................... 197
schwoerbeli, Brevaturus .......... 196
schwoerbeli, Corticacarus ...... 197
schwoerbeli, Dubiobates .......... 198
schwoerbeli, Limnesia .............. 193
Schwoerbelobatella .................. 199
scopularis ................................ 215
scutatus .................................... 198
scutellum .................................. 204
scutica ...................................... 203
Scutohydrachna ...................... 184
securipes .................................. 198
sellatus ...................................... 212
semireticulata .......................... 193
sensitiva .................................... 187
separata .................................... 198
Seppiella .................................. 193
serratipalpis .............................. 199
serratipes .................................. 193
serratirostris ............................ 190
serratiseta ................................ 206
setifera ...................................... 191
setipes, Corticacarus ................ 198
setipes, Neoaturus .................... 207
setipes, Piona .......................... 206
sextuberculatus ........................ 215
sica, Mideopsis ........................ 210
sica, Unionicola ...................... 202
sicaria ...................................... 206
Sigthoriella .............................. 189
silvestrii .................................... 184
sima .......................................... 191
similis, Brevaturus .................... 196
similis, Dubiobates .................. 198
similis, Schizobates .................. 200
simoni ...................................... 189
simplex ...................................... 216
simplicipes ................................ 201
simulans .................................... 203
singularis .................................. 209

sinuata ...................................... 201
sinuatipes, Atractides .............. 195
sinuatipes, Recifella ................ 205
sinuosa ...................................... 193
sinusoides ................................ 215
siolii, Eocorticacarus .............. 198
siolii, Koenikea ........................ 203
siolii, Unionicola ...................... 201
slanopa .................................... 193
smithi ........................................ 198
sneiderni .................................. 189
solitarius .................................. 215
solivaga .................................... 190
soona ........................................ 204
sorpresa .................................... 191
sovus ........................................ 197
spathulifer ................................ 215
Sperchon .................................. 188
SPERCHONTIDAE ................ 188
SPERCHONTINAE ................ 188
spinaxa ...................................... 198
spinicalcar ................................ 215
spinigera .................................. 207
spinosa ...................................... 204
staheli ...................................... 207
stalagmophila, Hydrodroma .... 187
stalagmophila, Wandesia .......... 187
stellatus .................................... 196
sterrodermus ............................ 194
striata ........................................ 211
stukabus .................................... 197
Stygalbiella .............................. 209
Stylohygrobates ........................ 200
Subcorticacarus ...................... 200
Submeramecia ........................ 190
Submiraxona ............................ 209
subrecta .................................... 201
subrotundata ............................ 213
subulifer .................................... 212
suchezae .................................... 198
sudamericana, Momoniella ...... 209
sudamericana, Piona ................ 206
surinamensis ............................ 193
sursumhians .............................. 185
sutava ........................................ 193
Szalayella ................................ 200
Tamboella ................................ 204
tamboensis, Arrenurus .............. 215
tamboensis, Hygrobates .......... 194
tanda ........................................ 207
taurus ........................................ 215
tenuicollis ................................ 215
tenuicoxalis .............................. 193
tenuipalpis ................................ 206
tenuirostris ................................ 186
testudo, Corticacarus .............. 198
testudo, Rhynchohydracarus .... 187
Tetracorticacarellus ................ 198
Tetrahygrobatella .................... 200
224
Tetralimnesia .......................... 193
Tetraneumania ........................ 204
thermacalis .............................. 186
THERMACARIDAE .............. 188
Thermacarus ............................ 188
thermalis .................................. 187
thermophila .............................. 195
Thoracaphoracarus ................ 216
thoracica .................................. 206
Thoracohygrobates .................. 200
THYADINAE .......................... 186
tigrina ...................................... 204
toriger ...................................... 213
tornquistensis ............................ 207
Torrenticola .............................. 190
TORRENTICOLIDAE ............ 189
TORRENTICOLINAE ............ 190
tortus ........................................ 212
tranasa ...................................... 189
transversa ................................ 203
transversalis ............................ 195
transversus ................................ 187
triangularis, Brevaturus .......... 196
triangularis, Hygrobates .......... 194
triangularis, Koenikea .............. 203
trichophorus ............................ 212
tricolor ...................................... 210
triconicus .................................. 215
tricornis .................................... 200
trifida ........................................ 208

trifoliata .................................... 210
trilobata .................................... 210
trispinosa .................................. 189
trispinosus ................................ 200
tronchonii ................................ 191
truncata .................................... 205
Truncaturus ............................ 216
Tubophorella ............................ 194
tucumanensis, Arrenurus .......... 216
tucumanensis, Limnesia .......... 193
tucumanensis, Omartacarus .... 194
tucumanensis, Stygalbiella ...... 209
tumidipalpis .............................. 201
tumidus .................................... 215
tumulosus .................................. 212
Tyrrellia .................................... 190
TYRRELLINAE ...................... 190
uberifer .................................... 212
umadus ...................................... 197
uncatus ...................................... 215
uncipalpis ................................ 190
undulata .................................... 205
undulatifrons ............................ 186
undulatus .................................. 215
undulicauda .............................. 215
unguiculata, Koenikea .............. 203
unguiculata, Protolimnesia ...... 191
ungularis .................................. 206
ungulata .................................... 187
unidens, Piona .......................... 206
unidens, Unionicola ................ 201

Unionicola ................................ 200
Unionicolella ............................ 202
UNIONICOLIDAE .................. 200
Unionicolides .......................... 202
UNIONICOLINAE .................. 200
unisetiger .................................. 215
unoka ........................................ 189
vaginalis .................................. 207
valdiviensis, Arrenurus ............ 212
valdiviensis, Dubiobates .......... 198
valdiviensis, Hygrobatella ........ 199
valdiviensis, Oxus .................... 189
valencius .................................. 213
valida ........................................ 192
validipalpis, Atractides ............ 195
validipalpis, Hygrobatella ........ 199
validipalpis, Neocorticacarus .... 199
validipalpis, Recifella .............. 205
vantermus ................................ 196
venezuelae ................................ 211
venezuelensis ............................ 206
ventrifissa ................................ 184
ventriperforatus ........................ 188
venusta ...................................... 209
verrucifer, Arrenurus ................ 215
verrucifer, Kyphohygrobates .... 199
vicina ........................................ 189
vicinipora .................................. 210
viciniseta .................................. 202
vicinus ...................................... 213

victoriola .................................. 203
vietsi, Australiobatella .............. 195
vietsi, Axonopsella .................... 207
vietsi, Centrolimnesia .............. 194
vietsi, Corticacarus .................. 197
vietsi, Koenikea ........................ 202
violacea, Mideopsis .................. 209
violacea, Neoalbia .................... 208
violacea, Neotorrenticola ........ 191
virescens .................................. 189
walteri, Arrenurus .................... 213
walteri, Neotorrenticola .......... 191
walteri, Submiraxona .............. 209
walteri, Wandesia .................... 187
Wandesia .................................. 186
WANDESIINAE ...................... 186
weberi ...................................... 188
wetzeli ...................................... 186
wetzlari .................................... 196
willinki, Arrenurus .................... 215
willinki, Neomamersa .............. 190
Wilsonatax .............................. 202
Xenthoracaphorus .................. 216
xiphophora ................................ 204
xystus ........................................ 197
yartesus .................................... 216
yungasa .................................... 198
Zabobates ................................ 200
zoldomus .................................. 195

Elenco y Biogeografía de Los Ácaros Acuáticos de Sudamerica

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Elenco y Biogeografía de Los Ácaros Acuáticos de Sudamerica

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Graellsia, 61(2): 181-224 (2005)

ELENCO Y BIOGEOGRAFA DE LOS CAROS ACUTICOS (ACARI,

B. Rosso de Ferrads1 y H. R. Fernndez2

ROSSO DE FERRADS & FERNNDEZ

los aos 60, siendo la mayora de las colecciones

CAROS ACUTICOS DE SUDAMRICA

ROSSO DE FERRADS & FERNNDEZ

trabajos anteriores a esta fecha y no incluidos en el

Andes Australes + Patagonia de Argentina.

Esta informacin sobre la distribucin y hbitats

CAROS ACUTICOS DE SUDAMRICA

GNERO Eylais Latreille, 1796

Hydryphantes (Hydryphantes) jujuyensis Nordenskild, 1904

ROSSO DE FERRADS & FERNNDEZ

Hydryphantes (Hydryphantes) pumilus (Viets, 1954)

Neocalonyx (Neocalonyx) godeti Walter, 1919

SUBFAMILIA PROTZIINAE Koenike, 1909

Eupatrella platano Goldschmidt & Gerecke, 2003

GNERO Neocalonyx Walter, 1919

GNERO Wandesia Schechtel, 1912

Neocalonyx (Neocalonyx) desajunos Goldschmidt & Gerecke,

Wandesia (Partnuniella) chechoi Cook, 1988

CAROS ACUTICOS DE SUDAMRICA

FAMILIA RHYNCHOHYDRACARIDAE Lundblad, 1936

GNERO Euwandesia Andr & Naudo, 1962

Euwandesia sensitiva Andr & Naudo, 1962

GNERO Rhynchohydracarus Lundblad, 1936

SUBFAMILIA CLATHROSPERCHONTINAE Lundblad, 1936

GNERO Clathrosperchon Lundblad, 1936

Clathrosperchon crassipalpis Lundblad, 1936

ROSSO DE FERRADS & FERNNDEZ

GNERO Thermacarus Sokolow, 1927

Subgnero Mixosperchon Viets, 1926

GNERO Sperchon Kramer, 1877

GNERO Rutacarus Lundblad, 1937

Sperchon loaensis Nom. nud.

Rutacarus (Rutacarus) pyriformis Lundblad, 1937

Subgnero Hispidosperchon Thor, 1901

Subgnero Eorutacarus Cook, 1980

Sperchon (Hispidosperchon) brasiliensis Lundblad, 1941

Rutacarus (Eorutacarus) ferradasae Cook, 1980

CAROS ACUTICOS DE SUDAMRICA

ROSSO DE FERRADS & FERNNDEZ

Neoatractides inachus Lundblad, 1941

Neomamersa uncipalpis Lundblad, 1953

GNERO Torrenticola Piersig, 1896

GNERO Meramecia Cook, 1963

Torrenticola (Torrenticola) columbiana (Lundblad, 1941)

Meramecia (Parameramecia) saltensis Fernndez, 2001

Subgnero Heteratractides Lundblad, 1941

Subgnero Submeramecia Cook, 1980

SUBFAMILIA TYRRELLINAE Koenike, 1910

Subgnero Monatractides Viets, 1926

GNERO Tyrrellia Koenike,1895

Torrenticola (Monatractides) brevis brevis (Lundblad, 1941)

Tyrrellia australis Besch, 1962

SUPERFAMILIA HYGROBATOIDEA Koch, 1842

GNERO Neotyrrellia Lundblad, 1938

CAROS ACUTICOS DE SUDAMRICA

SUBFAMILIA RHEOLIMNESIINAE Goldschmidt, 2004

SUBFAMILIA MIXOLIMNESIINAE Cook, 1988

ROSSO DE FERRADS & FERNNDEZ