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Chapter 30
Defining Plants
The kingdom Viridiplantae includes land plants and green algae -Red and brown algae are excluded All green plants arose from a single species of freshwater algae The green algae split into two major clades -Chlorophytes Never made it to land -Charophytes Did!
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Defining Plants
Defining Plants
Land plants have two major features 1. Protected embryos 2. Multicellular haploid and diploid phases
Defining Plants
Adaptations to terrestrial life 1. Protection from desiccation by a waxy cuticle and stomata 2. Evolution of leaves which increase photosynthetic surface area 3. Shift to a dominant vertical diploid generation
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Plants have a haplodiplontic life cycle -Multicellular diploid stage = Sporophyte -Multicellular haploid stage = Gametophyte
Chlorophytes
Green algae have two distinct lineages -Chlorophytes Gave rise to aquatic algae -Streptophytes Gave rise to land plants Chlamydomonas -Unicellular chlorophyte with two flagella -Have eyespots to direct swimming -Reproduces asexually as well as sexually
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Chlorophytes
Volvox -Colonial chlorophyte -Hollow sphere of a single layer of 500-60,000 cells -A few cells are specialized for reproduction
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Chlorophytes
Ulva -Multicellular chlorophyte
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Charophytes
Charophytes are green algae related to land plants
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Charophytes
Charales (300 species) -Macroscopic -Plant-like plasmodesmata -Sister clade to land plants Choleocaetales (30 species) -Microscopic -Plant-like mitosis -Next closest plant relatives
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Charophytes (Cont.)
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Bryophytes
Bryophytes are the closest living descendants of the first land plants -Called nontracheophytes because they lack tracheids (specialized transport cells) Simple, but highly adapted to diverse terrestrial environments Non-photosynthetic sporophyte is nutritionally dependent on the gametophyte
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Bryophytes
Liverworts (phylum Hepaticophyta) -Have flattened gametophytes with liver-like lobes -Form gametangia in umbrellashaped structures -Also undergo asexual reproduction
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Bryophytes
Hornworts (phylum Anthocerotophyta) -Sporophyte has stomata -Sporophyte is photosynthetic -Cells have a single large chloroplast
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Bryophytes
Mosses (phylum Bryophyta) -Gametophytes consist of small, leaflike structures around a stemlike axis -Anchored to substrate by rhizoids -Multicellular gametangia form at the tips of gametophytes -Archegonia Female gametangia -Antheridia Male gametangia Mosses withstand drought, but not air pollution
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Lycophytes
Club mosses are the earliest vascular plants -They lack seeds -Superficially resemble true mosses but they are not related -Homosporous or heterosporous
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Pterophytes
The phylogenetic relationships among ferns and their relatives is still being sorted out
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Pterophytes
Whisk ferns -Saprophyte consists of evenly forking green stems without leaves or roots
Pterophytes
Horsetails -All 15 living species are homosporous -Constitute a single species, Equisetum
-Consist of ribbed, jointed photosynthetic stems that arise from branching rhizomes
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Pterophytes
Ferns -The most abundant group of seedless vascular plants with about 11,000 species -The conspicuous sporophyte and much smaller gametophyte are both photosynthetic
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Pterophytes
The fern life cycle differs from that of a moss -Much greater development, independence and dominance of the ferns sporophyte Fern morphology -Sporophytes have rhizomes -Fronds (leaves) develop at the tip of the rhizome as tightly rolled-up coils -They unroll and expand
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Pterophytes
Fern reproduction -Most fern are homosporous -Produce distinctive sporangia in clusters called sori on the back of the fronds -Diploid spore mother cells in sporangia produce haploid spores by meiosis -At maturity, the spores are catapulted by snapping action
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Gymnosperms
Gymnosperms are plants with naked seeds -Ovule is exposed on a scale at pollination There are four living groups -Coniferophytes -Cycadophytes -Gnetophytes -Ginkgophytes All lack flowers and fruits of angiosperms
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Gymnosperms
Conifers (phylum Coniferophyta) are the largest gymnosperm phylum -Include: -Pines, spruces, firs, cedars and others -Coastal redwood Tallest tree -Bristlecone pine Oldest living tree Conifers are sources of important products -Timber, paper, resin and taxol (anti-cancer)
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Gymnosperms
Pines
-More than 100 species, all in the Northern hemisphere -Produce tough needlelike leaves in clusters -Leaves have: 1. Thick cuticle and recessed stomata 2. Canals into which cells secrete resin
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Gymnosperms
Pine reproduction -Male gametophytes (pollen grains) develop from microspores in male cones by meiosis -Female pine cones form on the upper branches of the same tree -Female cones are larger, and have woody scales -Two ovules develop on each scale
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Gymnosperms
Pine reproduction -Each ovule contains a megasporangium called the nucellus -Surrounded by the integument -Opening Micropyle -One layer becomes the seed coat -While scales of female cone are open, pollen grains drift down between them -Are drawn to top of nucellus
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Gymnosperms
Pine reproduction -While female gametophyte is developing, a pollen tube emerges from the pollen grain -It digests its way to the archegonium -Fifteen months after pollination, pollen tube reaches archegonium and delivers its sperm
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Gymnosperms
Cycads (phylum Cycadophyta) -Slow-growing gymnosperms of tropical and subtropical regions
-Sporophytes resemble palm trees -Have largest sperm cells of all organisms!
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Gymnosperms
Gnetophytes (phylum Gnetophyta) -Only gymnosperms with vessels in their xylem -Contain three (unusual) genera -Welwitschia -Ephedra -Gnetum
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Gymnosperms
Ginkgophytes (phylum Ginkgophyta) -Only one living species remains -Ginkgo biloba
Angiosperms
Angiosperms are the flowering plants -Ovules are enclosed in diploid tissue at the time of pollination
-The carpel, a modified leaf that covers seeds, develops into fruit
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Angiosperms
Angiosperm origins are a mystery -The oldest known angiosperm in the fossil record is Archaefructus -The closest living relative to the original angiosperm is Amborella
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Angiosperms
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Angiosperms
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Angiosperms
Flower morphology -Primordium develops into a bud at the end of a stalk called the pedicel -Pedicel expands at the tip to form a receptacle, to which other parts attach
Angiosperms
Flower morphology -Outermost whorl = Sepals -Second whorl = Petals -Third whorl = Stamens (androecium) -Each stamen has a pollen-bearing anther and a filament (stalk) -Innermost whorl = Gynoecium -Consists of one or more carpels that house the female gametophyte
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Angiosperms
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Angiosperms
Carpel structure -Three major regions -Ovary = Swollen base containing ovules -Later develops into a fruit -Stigma = Tip -Style = Neck or stalk
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