Escolar Documentos
Profissional Documentos
Cultura Documentos
e Econômica
São Cristóvão/SE
2010
Botânica Sistemática e Econômica
Elaboração de Conteúdo
Ana Paula do Nascimento Prata
Hermeson Alves de Menezes
Diagramação
Nycolas Menezes Melo
Ilustração
Ana Paula do nascimento Prata
Gerri Sherlock Araújo
Revisão
Edvar Freire Caetano
CDU 582:581.6
Presidente da República Chefe de Gabinete
Luiz Inácio Lula da Silva Ednalva Freire Caetano
Vice-Reitor
Angelo Roberto Antoniolli
AULA 2
Gimnospermas .................................................................................. 41
AULA 3
Caracterização e origem das angiospermas ..................................... 71
AULA 4
Nomenclatura botânica...................................................................... 93
AULA 5
#$###6., ............................................................... 105
AULA 6
Angiospermas basais e magnoliídeas ............................................. 129
AULA 7
Monocotiledôneas ........................................................................... 157
AULA 8
Rosídeas ......................................................................................... 191
AULA 9
Asterídeas ....................................................................................... 205
AULA 10
Botânica econômica ........................................................................ 223
Aula
MÉTODOS E TÉCNICAS DE
1
COLETA VEGETAL EM
ANGIOSPERMAS
META
Apresentar as técnicas de coleta em Angiospermas, incluindo noções básicas de coleta, prensagem e
herborização de material botânico, dinâmica e funcionamento de Herbário, além de fornecer algumas
dicas para herborizar famílias botânicas que apresentam peculiaridades morfológicas (Bromeliaceae,
Cactaceae e Arecaceae.
OBJETIVOS
6#$%%&"(
possuir conhecimento básico sobre as técnicas de coleta em Angiospermas;
saber sobre a dinâmica de um Herbário e a importância de uma boa prensagem e herborização
para a aquisição de uma exsicata adequada.
PRÉ-REQUISITOS
O aluno deverá ter noções sobre o conceito de Angiospermas (Magnoliophyta).
.D?<I9oI@FI<}E<:FEALEKFJ;<<JGu:@D<JM<><K8@JJ<:FJ<GI<EJ8;FJ
OFJ<D=FC?8J
;< :8IKFC@E8 ;<M@;8D<EK< @;<EK@
:8;FJ :8K8CF>8;FJ < ;@JGFJKFJ J<>LE;F 8 :C8JJ@
:8tqF
9FKpE@:8
gLD8:FC<tqF;@EpD@:8;<GC8EK8JJ<:8JHL<G<ID@K<<OKI8@ILK@C@Q8I<8;@:@FE8I
@E=FID8t{<JJF9I<:8;8LD8;8J<JGu:@<J:FE?<:@;8J<JF9I<EFM8J<JGu:@<JM<><K8@J
FEK<?KKG>CF9DLE;
C<J
NFI;GI<JJ
:FD
Botânica Sistemática e Econômica
" #JH
D=8:<;8;@M<IJ@;8;<9FKpE@:88KL8C<;8E<:<JJ@;8;<;<J<GI<J<IM8I
<I<>@JKI8I8J<JGu:@<JG8I8<JKL;FJ=LKLIFJuE<:<JJoI@F<@E;@JG<EJoM<CF
:FE?<:@D<EKF;<Ku:E@:8J8;<HL8;8JG8I8:FC<K8ID8K<I@8C9FKpE@:F:FE
J@;<I8E;F8J;@=<I<Et8J<DJL8<JKILKLI8GFIK<:FEJ@JKvE:@8<K:
DI<C8tqF8FGFIK<<8F?o9@KF8JGC8EK8J<D><I8CGF;<DJ<I:C8J
J@
:8;8J<D?<I9o:<8J8I9LJK@M8J8I9yI<8JKI<G8;<@I8J8HLoK@:8J<K:
JK8
:C8JJ@
:8tqF<DI<C8tqF8FGFIK<?o9@KF;FM<><K8CuLDGFL:FGFCvD@:8
GF@J:<IKFJC@D@K<J8@E;8EqF<JKqF9<D<JK89<C<:@;FJ
(LKI8:C8JJ@
:8tqF
:FEJ@;<I8EqFJFD<EK<FGFIK<;8GC8EK8D8JK8D9uD8CF:8C@Q8tqF;8
><D8;<:I<J:@D<EKF8FCFE>F;F:@:CF;<M@;8;FFI>8E@JDF+"33"'"
+.'$"+
8GL; ('e%/,%(+'3"
J ?<I9o:<8J GFJJL<D LD 8CKF K<FI ;< o>L8 <D JL8 :FEJK@KL@tqF <
JqF <D ><I8C ;< G<HL<EF GFIK< :FDF GFI <O<DGCF 8J FIHLw;<8J < FJ
>@I8JJy@J
J8I9LJK@M8JJqFI8D@
:8;8J;<J;<8JL898J<<GFJJL<D:<IKF
K<FI;<C@>E@
:8tqF<8J8I9yI<8JJqF><I8CD<EK<C@>E@
:8;8JI8D@
:8;8J
<;<D8@FIGFIK<HL<8J8EK<I@FID<EK<:@K8;8J
CuD;8G8IK@:LC8I@;8;<<D
I<C8tqF8FGFIK<?o9@KFGF;<DFJ:@K8I<O<DGCFJ;<GC8EK8JHL<GFJJL<D
LD8Ku:E@:8;<:FC<K8;@=<I<E:@8;8:FDF8JIFD<C@8:<8<9IFDuC@8J>I8
M8KoJD8:8D9@I8J8J8:K8:<8<G8CD8JD8E;8:8ILJO@HL< O@HL<J8J
I<:8:<8<:FHL<@IFJC@:LI@Q<@IFJ;<E;<Q<@IFJ<K:
8C>LD8J<JGu:@<J;<
GC8EK8J9LC9FJ8JD8IPC@;8:<8<"I@;8:<8<CC@8:<8<8JGC8EK8J8HLoK@:8J
&8P8:8:<8<<!P;IF:8I@K8:<8<;<EKI<FLKI8J
8
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
@>LI8O<DGCFJ;<GC8EK8J:FDGFIK<;@M<IJF8G8CD<@I8J
9?<I9o:<8FDD<C@E8:<8<
:
8I9yI<8
;8:K8:<8<
<8HLoK@:8J
=8I9LJK@M8J
>IFD<C@8:<8<?<I9o:<8
/F:vJ89@8HL<<JK8JKu:E@:8J;@=<I<E:@8;8J;<:FC<K8JJ<;qFG<CF=8KF
;<JK8JGC8EK8JGFJJLwI<D8C>LD8J:8I8:K<IwJK@:8JG<:LC@8I<J
'8J IFD<C@8:<8< 8J =FC?8J JqF ;<EJ8D<EK< ;@JGFJK8J 8F I<;FI ;<
LD<@OF=FID8E;FLD8IFJ<K8FHL<;@
:LCK88:FC<K8<GI@E:@G8CD<EK<8
GI<EJ8><D<8?<I9FI@Q8tqF;FD8K<I@8C
J8:K8:<8<GFJJL<D<JG@E?FJ98JK8EK<8
8;FJ<;@M<IJ@
:8;FJ8CuD
;<GFJJLwI<D98JK8EK<o>L8<DJL8:FEJK@KL@tqFFHL<K8D9uD;@
:LCK8E8
?FI8;8GI<EJ8><D<8ID8Q<E8D<EKFGF@J<JK<8:}DLCF;<o>L8GF;<Io
GIFG@:@8IFGFJK<I@FI8G8I<:@D<EKF;<=LE>FJEFD8K<I@8C;<GFJ@K8;FEF
!<I9oI@F:8JFFD8K<I@8CEqFK<E?8J@;FGI<EJ8;F:FII<K8D<EK<
8JG8CD<@I8J)8I8=8Q<I8GI<EJ8><D:FII<K8;<HL8CHL<ID8K<I@8C
G8I8@E:CL@IEF!<I9oI@FGI<:@J8DFJ8:FE;@:@FE8I88DFJKI8<DLD8=FC?8
;<AFIE8C;F9I8;FHL<D<;<:D:FDGI@D<EKFO:DC8I>LI8/F:vJ
GF;<D@D8>@E8I:FDF@I<DFJ=8Q<I@JJF)8I8GC8EK8J?<I9o:<8JHL<JqF
<D><I8C;<G<HL<EFGFIK<u=o:@CD8J<:FDLD8=FC?8;<:FHL<@IF:FD
8HL<C8J;@D<EJ{<J
C8IFHL<<O@JK<LD8Ku:E@:8G8I8<JK8JG<:LC@8I@;8;<JGF@J8:FC<K8;<
LD8<JGu:@<?<I9o:<8;<G<HL<EFGFIK<J<Io;@=<I<E:@8;8<D8@JJ@DGC<J
;FHL<8:FC<K8;<LD8IFD<C@8:<8<HL<K8D9uDu?<I9o:<8ALJK8D<EK<
GFI<JK8J8GI<J<EK8I<D8C>LD8J;@=<I<Et8J
&8@J8;@8EK<@I<DFJ<OGC@:8I
8 Ku:E@:8 ;<K8C?8;8D<EK< D8J EF <EK8EKF F HL< MF:vJ GI<:@J8D C<M8I
<D:FEJ@;<I8tqFuHL<8CuD;<JK8JG<:LC@8I@;8;<J:@K8;8J8EK<I@FID<EK<
8C>LD8J=8DwC@8J;<GC8EK8J@IqF8GI<J<EK8I8C>LEJ:8I8:K<I<JK8OFEzD@:FJ
@DGFIK8EK<JHL<;<M<IqFJ<ID8EK@;FJ<<JK8I:FEK@;FJE88DFJKI8G8I8
HL< GFJJ8DFJ @;<EK@
:8I <JK< D8K<I@8C :FII<K8D<EK< EF =LKLIF
/8DFJ
8GI<E;<ILDGFL:FJF9I<<JK8JKu:E@:8J;<:FC<K8GI<EJ8><DDFEK8><D
<:8I8:K<IwJK@:8J<JG<:w
:8J;<8C>LD8J=8DwC@8J
9
Botânica Sistemática e Econômica
:FC<K8;<D8K<I@8C9FKpE@:F;<M<GI@FI@Q8IJ<DGI<8HL8C@;8;<GIF
:LI8E;FI<GI<J<EK8IK8EKFHL8EKFGFJJwM<C8JM8I@8t{<JGFGLC8:@FE8@J<
J<IGI<:@J8E8J@E=FID8t{<J;@JGFE@9@C@Q8;8JHL8EKF8FCF:8C;8K8J<;8;FJ
I<=<I<EK<J8FD8K<I@8C<JG<:@8CD<EK<8HL<C<JHL<;<J8G8I<:<IqF;LI8EK<F
GIF:<JJF;<?<I9FI@Q8tqF:FI:?<@IF
)8I8=8:@C@K8IF<EK<E;@D<EKFM8DFJ@D8>@E8ILD8:FC<K8E8,<II8;<
"K898@8E8:?FHL<8D8@FI@8;<MF:vJAo:FE?<:<8,<II8<JL8J9<C<Q8J
E8KLI8@JKqF:@K8;8J
GI@D<@I8F9J<IM8tqFuHL<8:FC<K8;<D8K<I@8C9FKpE@:FC<M8IoF
;F9IFFLFKI@GCF;FK<DGF;FHL<J<=FIDFJ:FDG8I8I:FDLD8M@J@K8n
,<II8G8I8C8Q<I
)I<:@J8DFJ=8Q<IMoI@8JF9J<IM8t{<JGI<EJ8IFD8K<I@8C
:8II<>8I<HL@G8D<EKFJ<K:
/8DFJM<I@JJF:FDD8@J;<K8C?<J)8I8@E@
:@8IDFJ8EFJJ8:FC<K8GI<:@J8DFJEFJGI<G8I8IGC8E<A8IKF;FJFJ;<K8C?<J
<JJ<uFGI@D<@IFG8JJF
@>LI8O<DGCF;<G8IK<;8M<><K8tqF;8,<II8;<"K898@8E8,<I>@G<
FEK<'8J:@D<EKF #}E@FI
"!L!\"
(@Ew:@F
(F9A<K@MF;<J<I<8C@Q8ILD8:FC<K8u9LJ:8I<=FIE<:<ID8K<I@8CG8I8
G<JHL@J8J K8OFEzD@:8J FL 8EoC@J<J ;@M<IJ8J
LJ:8I < =FIE<:<I GFIHL< n
D<;@;8HL<8><EK<:FC<K8<@EJ<I<8J8DFJKI8JEF8:<IMF;<LD?<I9oI@F
@I<DFJGIFM<ID8K<I@8CG8I8G<JHL@J8J=LKLI8J
10
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
KI8Q<IGFIt{<J;<GC8EK8JHL<GF;<IqFJ<IM@I:FDF98J<G8I8MoI@8J89FI;8
><EJ;<EKIF;8oI<8;<FKpE@:8<oI<8J8
EJ
/8DFJM<I8C>LEJ<O<DGCFJ
8 DFJKI8J ;< GC8EK8J G8I8 GFJK<I@FI J<:8><D GI<=<I<E:@8CD<EK< :FD
1
;LGC@:8K8J
@>LI8DFJKI8J;<GC8EK8J
LI8EK<8:FC<K8
<GF@J;FGIF:<JJF;<?<I9FI@Q8tqF<
DFEK8><D
FEK<
'8J:@D<EKF #}E@FI
&8KFJ
9CFI<J=ILKFJ<G8IK<JM<><K8K@M8JGI<J<IM8;8J<D
FLoC:FFC8
G8I8<JKL;FJ8E8KzD@:FJ
@>LI8O<DGCF;<Ku:E@:8J;<GI<G8I8tqF;<CpD@E8J?@JKFCy>@:8J8G8IK@I;<=ILKFJ
O<DGCFJ;<D8K<I@8CLK@C@Q8;F
ILKFC@DqF
)C8EK8HL<=F@LJ8;8
G8I8I<8C@Q8IF:FIK<8E8KzD@:F
FIK<KI8EJM<IJ8C
FEK<FKpE@:F
4
O8;FIG8I88EoC@J<8E8KzD@:8=FIDFCoC:FFC<o:@;F
8:uK@:F6
11
Botânica Sistemática e Econômica
:&8K<I@8CG8I8<JKL;FJ;<:IFDFJJFDFJ
;ILKFJG8I88:8IGFK<:8
@>LI88IGFK<:8;F!<I9oI@F1@E>y
FEK<8IM8C?F
<DFJKI8J;<D8;<@I8JG8I8<JKL;FJ8E8KzD@:FJ
@>LI8FC<tqF;<8DFJKI8J;<D8;<@I8;81@CFK<:8;F#8I;@DFKpE@:F-IFG@:8C
FEK<FEK<?KKGNNN
@@:K
GK
=&8K<I@8CG8I8<JKL;FJHLwD@:FJ
>&8K<I@8CG8I88EoC@J<DFC<:LC8I<
?&L;8J<JK8:8J<J<D<EK<JG8I8GIFG8>8tqF
>FI8HL<AoJ89<DFJHL<J<GC8E<A8IDFJ9<DLD8J@DGC<J:FC<K8
GF;<I<DFJKI8Q<ID8K<I@8CG8I8MoI@8J89FI;8><EJ
8w<D;@8EK<uJy;<
:@;@IHL8@JG8IK<J;8GC8EK8@I<DFJ:FC<K8I<:FD<t8IDFJ8G<EJ8I<DHL8@J
D8K<I@8@J@I<DFJC<M8IG8I8I<8C@Q8IDFJLD8:FC<K88;<HL8;8
12
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
" !
8&8K<I@8@J;<LJFG<JJF8CK8@J:FDF:?8GuLy:LCFJ;<JFCGIFK<KFIJFC8I
1
I<G<C<EK<9FK8DF:?@C89@Ey:LCFJ:8@O8;<GI@D<@IFJJF:FIIFJ<K:
9)I<EJ8;<:8DGF8CKwD<KIF9}JJFC8D8G8FL ),
@>LI8O<DGCFJ;<D8K<I@8@JLK@C@Q8;FJG8I88:FC<K8<GI<EJ89FKpE@:8
+<K@I8;F;<"% (
('('"
13
Botânica Sistemática e Econômica
: -<JFLI8 ;< GF;8 =8:qF GF;qF <JG<:w
:F <HL@G8D<EKF ;< 8J:<EJqF
<FLKI8J=<II8D<EK8J8GIFGI@8;8J8F:FIK<;<;<K<ID@E8;8G8IK<;<LD8
GC8EK8C<E?FJ8FLG8I8I<DFM<ILD8GC8EK8;FJFCFJ8:FJGCoJK@:FJ<GFK<J
?<IDuK@:FJKI<E89@Ey:LCFDoHL@E8=FKF>Io
:8Io;@FJ:FDLE@:8;FI<J<K:
;#FIE8@JFLG8G<CG8I;F;<<D9ILC?F:D:FDGI@D<EKFO:DC8I
>LI8;F9I8;FEFJ<EK@;FCFE>@KL;@E8C
<8;<IEF;<:8DGFFL9CF:F;<
:?8J;<:FC<K8
=%oG@J:FDLDFL:8E<K8;<K@EK8@E;<CuM<C
! $JQ!
LI8EK<8:FC<K88C>LD8J@E=FID8t{<JGI<:@J8DJ<I8EFK8;8J8@E;8<D
:8DGFGF@J<C8J><I8CD<EK<JqFG<I;@;8JEFGIF:<JJF;<?<I9FI@Q8tqFK8@J
:FDF8:FI;8JFI<J;FJ=ILKFJF;FI<J<K:
g@EK<I<JJ8EK<=FKF>I8=8IKF;F
FD8K<I@8CGI@E:@G8CD<EK<8HL<C<HL<<JKo<DCF:8C;<;@=w:@C8:<JJFG8I88
:FC<K8
EFK8I8=I<HLvE:@8;8GC8EK8EF8D9@<EK<K8D9uDu@DGFIK8EK<
9<D:FDFI<8C@Q8IF9J<IM8t{<J<DI<C8tqF88C>LEJ8JG<:KFJHL<J<IqF
;<K8C?8;FJ8J<>L@I
JH
,FCF
<G<E;<E;F;FCF:8C<J:FC?@;FG8I88:FC<K8FJFCFM8@8GI<J<EK8I
8C>LD8J;@=<I<Et8J@DGFIK8EK<JHL<GI<:@J8DJ<I8EFK8;8J
FEJK@KL@tqFF9J<IM8IJ<FJFCFu8I>@CFJF8I<EFJF?LDw=<IFIF:?FJF
G<;I<>FJF:8C:oI<F<K:
'8KLI<Q8u@DGFIK8EK<I<>@JKI8IJ<FJFCFuJ<:F}D@;F9I<AFJF@ELE
;8;FFL@ELE;oM<CJ8C@EF8C:8C@EFo:@;F<K:
14
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
1
@>LI8O<DGCFJ;<:FC<K8<D;@=<I<EK<JCF:8C@;8;<J;<,<I>@G<
8E@E;u;<,qFI8E:@J:F
)@I8D9L
,<II8;8+@9<@I8
FEK<
,@CM8
-@GFM<><K8:@FE8C
88K@E>8 <II8;F +<JK@E>8 &8K8 (D9Iy
C8 &8K8 J<D@ ;<:w;L8
:8GF<@I8oI<8:LCK@M8;8<K:
"E;@:8E;F88CK@KL;<;FGFEKF;<:FC<K8<D
I<C8tqF8FEwM<C;FD8ID
J
D
@>LI8
O<DGCFJ;<K@GFJM<><K8:@FE8@J
+<JK@E>8
&8K8KCpEK@:8
88K@E>8
FEK<'8J:@D<EKF #}E@FI
,L9JKI8KF=FID8;<M@;8
,<>L<8JGI@E:@G8@J=FID8J;<M@;8J<>LE;F8;<
E@tqF;<FEK*L<I
-<IIw:FC8J/<><K8@JHL<M@M<DE8K<II8
,8Ow:FC8J/<><K8@JHL<:I<J:<D<EKI<IF:?8J
+LGw:FC8J/<><K8@JHL<:I<J:<D<DG<E?8J:FJ
Gw
K8J/<><K8@JHL<M@M<DJF9I<FLKIFJM<><K8@JJ<DK@I8IFJEL
KI@<EK<J;F?FJG<;<@IF
,8GIy
K8J /<><K8C ?<K<IyKIF=F HL< J< ELKI< ;< 8E@D8@J < M<><K8@J
DFIKFJ<;<HL8CHL<II<JKF;<FI>pE@:F<D;<:FDGFJ@tqF
HLoK@:FJ/<><K8CHL<:I<J:<:FDGC<K8FLG8I:@8CD<EK<JL9D<IJF
<D:LIJFJ;o>L8
%LD@EFJ@;8;<
.D9Iy
C8J(D9Iy
C8J)C8EK8JHL<GI<=<I<DJFD9I8
15
Botânica Sistemática e Econômica
JHB"
)FIK<?o9@KF
IM88I9LJKFFLJL98I9LJKF8IMFI<K8oIMFI<KI<G8;<@I8<K:
@E;@:8E;F
8JD<;@;8J8GIFO@D8;8JGI@E:@G8CD<EK<88CKLI8
jI>qFJM<><K8K@MFJ
+8@QEqF?oI<=<IvE:@88EqFJ<ILDK@GF<JG<:@8C
8LC<K<OKLI8;8:8J:8:FI8IFD8HL8E;F=FI:8I8:K<IwJK@:F
FC?8F9J<IM8I8:FEJ@JKvE:@8:FI?<K<IF C@8C8K<J:vE:@88IFD8
@>LI8
O<DGCFJ;<yI>qFJM<><K8K@MFJ
8LC<
FC?8J
+8@Q
FEK<'8J:@D<EKF
#}E@FI
jI>qFJI<GIF;LK@MFJ
CFI:FI;8J;@=<I<EK<JG8IK<JF;FIJ<OF
"EFI<J:vE:@8
ILKF:FEJ@JKvE:@8K@GF:FI>I8L;<D8KLI@;8;<
16
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
KEF9FKpE@:8
'FD<GFGLC8ILK@C@;8;<G8I88:FDLE@;8;<CF:8C
1
"!"
g@E;@JG<EJoM<C:FC<K8IFJ<JGu:@D<JKqF:FDGC<KFJHL8EKFGFJJwM<C
)8I88<
:@vE:@8;8@;<EK@
:8tqF8FIu89JFCLK8D<EK<E<:<JJoI@88JJ@D
:FDFFJ=ILKFJHL8E;F?FLM<IGF@JJ<D<JJ<J<C<D<EKFJuHL8J<@D
GFJJwM<CI<:FII<I8HL8@JHL<IJ@JK<D8J;<:C8JJ@
:8tqF<O@JK<EK<J<DHL<8
FI
>LI8:FDFFyI>qFD8@J@DGFIK8EK<
(D8K<I@8C:FC<K8;F;<M<J<IGI<=<I<E:@8CD<EK<GI<EJ8;F<;<GFJ@K8;F
E8 <JKL=8 EF D<JDF ;@8 ;8 :FC<K8
'F DFD<EKF ;8 :FC<K8 C<D9I8I ;<
KFD8IEFK8JF9I<8C>LEJ8JG<:KFJ;8GC8EK8:FDF:FI;8JFI<J<=ILKFJ
:?<@IFGI<J<Et8;<GvCFJEFJyI>qFJ;8GC8EK88CKLI8;8GC8EK8<FLKI8J
:8I8:K<IwJK@:8JHL<GF;<DJ<II<C<M8EK<J
JJ<J;<K8C?<JJqF;<<OKI<D8@D
GFIKpE:@8EFGIF:<JJF;<@;<EK@
:8tqF9FKpE@:8<DL@KFJ;<C<JJ<G<I;<D
8GyJ8GI<EJ8><DJ<:8><D;FD8K<I@8C:FE=FID<=F@:@K8;F8EK<I@FID<EK<
CuD;8J:8I8:K<IwJK@:8J:@K8;8JFLKIFJ;<K8C?<J;<M<DJ<I8EFK8;FJ
.D8;<J:I@tqF;<K8C?8;8;<JK<J;8;FJ<E:FEKI8 J<EFG8E<KF;F!<I9oI@F
;8.E@M<IJ@;8;<<;<I8C;<,<I>@G<,;@JGFEwM<CE8@EK<IE<KEFJ<>L@EK<
<E;<I<tF?KKGNNN
L=J
9IDLJ<LJ?<I98I@FFLE8GC8K8=FID8DFF;C<
-I8K8E;F J<;<GC8EK8J?<I9o:<8Ju:FEM<E@<EK<:FC<K8IF<O<DGC8I
@EK<@IFGF@J8;@JGFJ@tqF;8JI8wQ<J;<M<J<IF9J<IM8;8G8I88:C8JJ@
:8tqF
@>LI8
O<DGCFJ;<:FC<K8;<GC8EK8J?<I9o:<8J
FEK<
,@CM8
17
Botânica Sistemática e Econômica
#oG8I88JGC8EK8J;<D8@FIGFIK<:FC<K8II8DFJFI@;FJ;<8GIFO@
D8;8D<EK< :D
,<8J=FC?8J;<LD8GC8EK8=FI<DDL@KF>I8E;<J
GF;<IqFJ<I;F9I8;8J;<DF;FHL<J<:FEJ<IM<D@EK<@I8J
_IMFI<J<8I
9LJKFJGF;<IqFJ<ID8I:8;FJ<D:8DGFG8I8J<I<D:FC<K8;FJ<DFLKI8J
F:8J@{<J<8JJ@DGF;<I J< oF9K<IFI<JFL=ILKFJ;<uGF:8J;@=<I<EK<J
@>LI8
O<DGCF;<LK@C@Q8tqF;FGF;qFG8I8:FC<K8ID8K<I@8C=uIK@C;<oIMFI<J
FEK<
&8KFJ
(D8K<I@8C;<oIMFI<GF;<IoM@I8:FDG8E?8;F:FD8DFJKI8J;8I<JG<:K@M8
Dióica :8J:8
,<?FLM<I;@=<I<Et8JD8I:8EK<JEF8JG<:KF;8J=FC?8J;898J<;8:FG8<
Planta – Diz-se da ;FoG@:<8DFJKI8J;<8D98J;<M<IqFJ<I:FC<K8;8J
*L8E;FFJI8DFJ=uIK<@J
estritamente mas-
;<LD8;<K<ID@E8;8<JGu:@<8GI<J<EK8I<D:8I8:K<I<JDFI=FCy>@:FJ;@=<I<EK<J
culinas e estrita- ;FJ<JKuI<@J8D9FJ;<M<DJ<I:FC<K8;FJ
*L8E;F8GC8EK8u ,1Y1+);<M< J<
mente femininas :FC<K8ID8K<I@8C;<8D9FJFJJ<OFJ
18
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
@>LI8 O<DGCF;<=ILKF>I8E;<J8GL:8@8
FEK<!KKG
9G
9CF>JGFK
:FD
JGC8EK8J;<JK@E8;8J8LD88EoC@J<HLwD@:8GF;<DJ<I:FC<K8;8J<J
KuI<@J<FG<JF;FD8K<I@8C;<:8;8<JGu:@<;<M<IoJ<IJLG<I@FI8
$>
g
@DGI<J:@E;wM<CHL<LD88DFJKI8=uIK@C;FD<JDFD8K<I@8C<DHL<=F@=<@K8
88EoC@J<J<A8@E:FIGFI8;88F8:<IMF;F!<I9oI@F
<M< J<:FC<K8I;<88DFJKI8J;<:8;8<JGu:@<;<DF;FHL<J<GFJJ8
LJ8I8C>LD8G8I8<O8D<;<K8C?8;FFLKIFJ:FDFK@GFJ8J<I<D:FEJ<IM8;FJ
FL8@E;8G8I8G<IDLK8;<;LGC@:8K8J:FDFLKI8J@EJK@KL@t{<J:@<EKw
:8J
!o
:8JFJ<DHL<FE}D<IF;<;LGC@:8K8J;<M<J<ID<EFIFLD<JDFEqF?8M<I
;LGC@:8K8JHL8E;FJ<KI8K8I;<LD8<JGu:@<I8I8FL8D<8t8;8;<<OK@EtqF
9-5:)/-4
(@;<8CJ<I@8?<I9FI@Q8I8GC8EK8@D<;@8K8D<EK<CF>F8GyJ8:FC<K8FL
J<A8GI<EJo C8<J<:o C8G8I8D<C?FIGI<J<IM8IJL8J:8I8:K<IwJK@:8J
*L8E;F
@JJFEqF=FIGFJJwM<C;<M< J<C<M8I8GC8EK88@E;8=I<J:8G8I88@;<EK@
:8tqFD8EK<E;F 8<DJ8:FJGCoJK@:FJ=<:?8;FJG8I8HL<JL8J=FC?8JEqF
J<HL<D<E<D<EIL>L<D?<I9FI@Q8E;F 8GFJK<I@FID<EK<
'8GI<EJ8><D
:8;88DFJKI8;<GC8EK8u:FCF:8;8<EKI<=FC?8J;<AFIE8@J8CK<IE8;8J:FD
=FC?8J;<G8G<CqF
(:FEALEKF;8J8DFJKI8Ju:FCF:8;F<EKI<;L8JGI<EJ8J
;<D8;<@I8<8D8II8;F=FIK<D<EK<:FD:FI;{<JI<J@JK<EK<J
19
Botânica Sistemática e Econômica
@>LI8 O<DGCF;<GI<EJ8><D;<D8K<I@8C9FKpE@:F
)I<EJ8E;F
gDL@KF@DGFIK8EK<8EFK8IFEFD<<E}D<IF;<:FC<K8E8=FC?8;<AFIE8C
HL<:FEKuD8GC8EK8
;<@O<FK8D8E?F;88DFJKI8D<EFI;FHL<F;FAFIE8CGF@J8JG8IK<JHL<
JF9I<JJ8wI<DJ<IqF;8E@
:8;8J8FGI<EJ8I
J<8J8DFJKI8JEqF=FI<DGI<EJ8;8J8@E;8<D:8DGFu@DGFIK8EK<<M@K8I
8G<I;8;< o>L8DLI:?8
JK8G<I;8 GF;<J<I;@D@ELw;8 :FCF:8E;F8J
8DFJKI8JE8JJ8:FC8JGCoJK@:8J:FDo>L8GF;<E;F8;@:@FE8IK8D9uDG8G<C
DFC?8;F<;<@Oo C8JnJFD9I8
G8I8;@D<EJ@FE8ID<C?FIFJI8DFJ>I8E;<JHL<9I< FJ<D=FID8;</
)8I88JKI<G8;<@I8J:LIMo C8J<D=FID8;<.
@>LI8
+<K@I8;8;<%",'++
@<C;K<:?E@HL<JLJ<;
9P&@JJFLI@FK8E@:8C 8I;<E
20
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
J<8GC8EK8uG<HL<E8:FCF:8IMoI@8JE8D<JD8=FC?8 1
@>LI8
+<K@I8;8;<%",'++
@<C;K<:?E@HL<JLJ<;9P
&@JJFLI@FK8E@:8C 8I;<E
8IILD<8JGC8EK8J;<=FID8HL<8J;L8JJLG<I=w:@<J;8J=FC?8J<;8J<J
KILKLI8JI<GIF;LK@M8JGFJJ8DJ<IF9J<IM8;8J
GI<EJ<8C>LD8JFI<J89<IK8JFLKI8J=<:?8;8J<FLKI8J:FD8J<JKILKLI8J
I<GIF;LK@M8JnDFJKI8
:8JF?8A8JFD<EK<LD8}E@:8=FC?8>I8E;<;@JGFEwM<C;F9Io C8;<=FID8
HL<8J;L8J=8:<JGFJJ8DJ<IF9J<IM8;8J
@>LI8
+<K@I8;8;<%",'++
@<C;K<:?E@HL<JLJ<;9P&@JJFLI@FK8E@:8C 8I;<E
EqF :L9I8 8J <JKILKLI8J I<GIF;LK@M8J ;< JL8 8DFJKI8 :FD 8J =FC?8J
FCFHL< 8J8:@D8;8J=FC?8J
@>LI8
+<K@I8;8;<%",'++
@<C;K<:?E@HL<JLJ<;9P&@JJFLI@FK8E@
:8C 8I;<E
21
Botânica Sistemática e Econômica
I<K@I<F<O:<JJF;<=FC?8J;88DFJKI8
'F<EK8EKF;<@O<LDG<;8tF;F
G<:wFCFnDFJKI8G8I88F9J<IM8tqF;8;@JGFJ@tqF=FC@8I
@>LI8 +<K@I8;8;<%",'++
@<C;K<:?E@HL<JLJ<;9P&@JJFLI@FK8E@:8C 8I;<E
K<ID@E8;88DFEK8><D;8GI<EJ8:FCF:8 J<FD8K<I@8CE8<JKL=88V
GFILDK<DGFDu;@F;<;@8J;<G<E;<E;F;FK@GF;<D8K<I@8C
GyJ
<JK<J:@E:F;@8J89I@I8GI<EJ8G8I8:?<:8IFHL<Ao<JKoJ<:FGIFEKFG8I8
J<II<K@I8;F
HL<C<D8K<I@8CHL<8@E;8<JK@M<I}D@;F;<@O8IE8<JKL=88Ku
HL<<JK<A8:FDGC<K8D<EK<J<:F
GyJ8J8w;8;8<JKL=8FD8K<I@8C;<M<IoJ<I@;<EK@
:8;F;<M@;8D<EK<
<K@HL<K8;F<@E:FIGFI8;F8F8:<IMF
(!<I9oI@F,K<DJL8JGIyGI@8J
I<>I8J ;< @E:FIGFI8tqF ;< D8K<I@8C HL< <JKqF ;@JGFEwM<@J EF J<>L@EK<
<E;<I<tF?KKGNNN
L=J
9IDLJ<LJ?<I98I@F
@>LI8
O<DGCFJ;<GI<EJ8><D;<D8K<I@8C
)I<EJ8JG8G<C{<J:FEIIL>8;FJ
FEK<
&8KFJ
'8J:@D<EKF #}E@FI
22
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
,<:8><D
+<:FD<E;8 J<8LK@C@Q8tqF;<<JKL=8JG8I8HL<8J8DFJKI8JJ<HL<DD8@J
I8G@;8D<EK<<;<D8E<@I8LE@=FID<
FD@JJF<M@K8 J<8:FEK8D@E8tqFGFI
1
=LE>FJ<C@D@E8E;F J<K8D9uDGFJJwM<@JGI8>8JHL<8:FDG8E?<D8JGC8EK8J
FDI<C8tqFnJ<JKL=8JGF;<DJ<In98J<;<I<J@JKvE:@8CpDG8;8JFL
8>oJ
'8=8CK8;<JJ<JI<:LIJFJLD8J<:8><DE8KLI8CGF;<J<I@DGIFM@J8;8
FD8K<I@8CGI<EJ8;F;<M<
:8I<DCF:8C<EJFC8I8;FM<EK@C8;FFLGIyO@DF
;<LD8=FEK<;<:8CFI
(JAFIE8@J}D@;FJ;<M<DJ<IKIF:8;FJ;@8I@8D<EK<
<8GI<EJ8EqF;<M<
:8IJF9FJ<I<EFG8I8<M@K8IHL8CHL<I:FEK8D@E8tqF
@>LI8O<DGCFJ;<<JKL=8;<J<:8><D;<D8K<I@8C9FKpE@:F
JKL=8@E;LJKI@8C
<JKL=8
;<:8DGF
!<I9FI@Q8tqF
(DF;F;<GI<G8I8I8JGC8EK8JJ<:8E;F 8J<EKI<8JGI<EJ8JuJ@DGC<J<
8G<E8JI<HL<ILDGFL:F;<G8:@vE:@8
:FE;@:@FE8 J<:8;88DFJKI8ELD8
=FC?8;<AFIE8C;F9I8;F8J=FC?8JJ<IqF8D8II8;8JE8GI<EJ8<<JK8C<M8;8
G8I8LD8<JKL=8
@>LI8O<DGCFJ;<GI<EJ8><D?<I9FI@Q8tqF;<D8K<I@8C9FKpE@:F
23
Botânica Sistemática e Econômica
D<O:LIJ{<JIoG@;8JFD8K<I@8CGF;<IoJ<IKI898C?8;FEFC89FI8KyI@F
HL8E;FCFE>8JEF
D;F;@8FLEFDFD<EKF;8:FC<K89FII@=8IFD8K<I@8C
:FD98JK8EK<oC:FFCDFC?8E;F@E:CLJ@M<FG8G<C<D9ILC?8I:FDGCoJK@:F
M<;8E;FFD<C?FIGFJJwM<CG8I8<M@K8I8<M8GFI8tqF;FoC:FFCFD8K<I@8C
O8;FGF;<IoJ<I>L8I;8;FJ<DJ<:8IGFIMoI@FJ;@8J
JJ<GIF:<JJF;<M<Io
J<I@E;@:8;FEFJIyKLCFJ;FD8K<I@8CDFEK8;F
JGC8EK8JHL<I<HL<I<DD8@FI<JKI898C?FJE8?<I9FI@Q8tqF<D><I8C
JqF8JJL:LC<EK8J8J9LC9FJ8J<8J8HLoK@:8J
,L:LC<EK8J
@>LI8O<DGCF;<GC8EK8JL:LC<EK8&8E;8:8IL
8:K8:<8<
Cladódio
Caule fotossinteti-
zante, caracter- 8:K8:<8<
ístico das plantas JK8=8DwC@8u:FEJ@;<I8;8LD8;8JD8@J@EK<I<JJ8EK<J;<EKI<8JE>@FJG<I
áfilas em clima
D8JGFI8GI<J<EK8ILD8:C8I8<JG<:@8C@Q8tqF88D9@<EK<JJ<:FJ,FLQ8
mais árido. Pos-
sui crescimento %FI<EQ@
,qF<IM8J><I8CD<EK<JL:LC<EK8J:FD:8LC<J<>D<EK8;F<D
indeterminado e :C8;y;@FJ<=FC?8JDF;@
:8;8J<D<JG@E?FJ
co das Cactaceae
'8
>LI8;F<O<DGC8I<D:8DGFGF;< J<M@JL8C@Q8IFCF:8C;<@E:@JqF
e Euphorbiaceae.
Cladódios achata- ;F:C8;y;@F
GyJFGI@D<@IF:FIK<JqF=<@K8JJ<t{<JCFE>@KL;@E8@JG8I8I<K@I8I
dos são chamados FG8IvEHL@D8<J<:t{<JKI8EJM<IJ8@JKFD8E;FF:L@;8;F;<GI<J<IM8I
de filocládios. 8J:FJK<C8J@EK8:K8J
(E}D<IF;<:FJK<C8Ju:FEJ@;<I8;F:8IoK<IK8OFEzD@:F
renzi 2007).
24
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
@>LI8O<DGCF;8:FC<K8;<8:K8:<8<
!o9@KF
,<tqFCFE>@KL;@E8C;F:C8;y;@F
CFI
DFJKI8:FDJ<tqFCFE>@KL;@E8C;F:C8;y;@F<;8FI<J<tqFKI8EJM<IJ8C;F=ILKF
,<tqF
CFE>@KL;@E8C;8FI
,<tqFKI8EJM<IJ8C;F:C8;y;@F
FEK<"
%<DFJ#I
)8I88C>LD8JJL:LC<EK8J;<>I8E;<GFIK<uI<:FD<E;8;FD<I>LC?o
C8J<Do>L8=<IM<E;FGFI8C>LEJD@ELKFJ<I<K@I8IFG8IvEHL@D8;<GF@J
;<=<IM@;8J
*L8E;F 8 GC8EK8 8 J<:8I GFJJL@ <JG@E?FJ FL 8:}C<FJ KFIE8 J< E<
:<JJoI@F;<@K8I J<FJGI@D<@IFJGFID<@F;<LD8=8:8FLLD8<JGoKLC8;<
D8;<@I8FJJ<>LE;FJ@JKFuFJ8:}C<FJ;<M<DJ<II<K@I8;FJ8HL<C<JHL<
:8D:FEKI8FG8G<C:FEJ<IM8E;F J<8G<E8JFJC8K<I8@JFLJ<A8FJHL<
EqFGI<AL;@:8D8GI<EJ8><D
IFD<C@8:<8<
<M@;Fn;@JGFJ@tqF;<JL8J=FC?8J8J9IFDuC@8J8:LDLC8Do>L8EF
J<L@EK<I@FI=8KFIHL<8KI8@MoI@FJFI>8E@JDFJ:FDF8E=w9@FJ8IKIyGF;<J
IuGK<@J;<EKI<DL@KFJFLKIFJ
)FI@JJF;<M< J<K<IDL@K8:8LK<C88F:FC<K8I
9IFDuC@8JG8I8HL<8:@;<EK<JJ<A8D<M@K8;FJ
)8I8I<K@I8IF@E;@Mw;LF;F
JFCFuE<:<JJoI@888AL;8;<LD=8:qF
FI;8JG8I8GLO8IF@E;@Mw;LFJqF
98JK8EK<}K<@JGI@E:@G8CD<EK<HL8E;FJqF<JGu:@<J;<>I8E;<GFIK<
F
I<K@I8I88DFJKI8;8I8C>LD8J98K@;8J:FDF=8:qFG8I8HL<FJ8E@D8@J
J8@8D<DJ<>L@;8I<K@I8 J<KF;8J8J=FC?8JJ<C<:@FE8E;F J<8D8@FI<8
D<EFIG8I88GI<EJ8><DALEK8D<EK<:FD8@EFI<J:vE:@8
.D@DGFIK8EK<
:8IoK<IK8OFEzD@:F;<JK8=8DwC@8JqF8J:FI<J;8J9Io:K<8J<;8J98J<J;8J
=FC?8J98@E?8J
@:8J;<FC<K8;<IFD<C@8:<8<F==8E@ 'LE<J
JGu:@D<J:FDIFJ<K88GyJJ<II<K@I8;F;FJL9JKI8KF;<M<8M@Io C8
G8I8I<K@I8IKF;88o>L8
25
Botânica Sistemática e Econômica
:?8:F8C?8I<98K<IE8IFJ<K8G8I8;<IIL98I8D8KuI@8FI>pE@:88:LDLC8;8
<GFJJwM<@J8E@D8@JHL<<JK<A8DEFJ<L@EK<I@FI
<JGu:@D<J;<G<HL<EFGFIK<8Ku:<I:8;< :DFL:FDGFL:8J
=FC?8JD<EFJ;<
E8IFJ<K8GF;<GI<EJ8IKL;F
<JGu:@D<J;<>I8E;<GFIK<8:@D8;<:DFLDL@K8J=FC?8JD8@J;<
8<JDFEK8I8IFJ<K89<J:FC?<IFL=FC?8J;8I<>@qF@EK<ID<;@oI@8
;8IFJ<K8=FC?8J9<D;<J<EMFCM@;8J<:FD9FD<JK8;FJ<DDL@KFJD8
:?L:8;FJ(,'qF:FIK<8=FC?8<C8;<M<<JK8I@EK<@I8GFIK8EKF:FD
898@E?8<CpD@E8=FC@8I:F<J:8GF;8@EFI<J:vE:@8K8D9uD;<M<
:8I
@EK<@IFG8I8GI<EJo CF
@EFI<J:vE:@8DL@KF>I8E;<:FD<J:8GFIF9LJKFD8@FIHL<:D;<
C8I>LI88E<JJ<J:8JFJGF;<;@M@;@I8@EFI<J:vE:@8<F<J:8GF8FCFE>F
;FJ<L<@OFCFE>@KL;@E8C
"&)(+-'-'8;}M@;8;<:FDFGI<EJ8IFD8K<I@8C;<M<<JK8I
GI<J<EK<EFDwE@DFF<J:8GF<8@EFI<J:vE:@8@E=ILK<J:vE:@8@EK<@IFJ<
FL=FC?8J<D9FD<JK8;FK8D9uD@EK<@IFJ
@>LI8O<DGCFJ;<:FC<K8;<IFD<C@8:<8<
FEK<
,@CM8
'8J:@D<EKF
#}E@FI
26
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
I<:8:<8<G8CD8<8JG8CD<@I8J
/8DFJ8>FI8MFCK8IEFJJ88K<EtqFG8I88JG8CD<@I8J(HL<><I8CD<EK<
EFJ:?8D888K<EtqF<DGC8EK8J;<JK8=8DwC@8"D8>@E<LD:FHL<@I8C
,qF
1
GC8EK8J<D><I8C:FD:8LC<;@=<I<E:@8;F;FK@GF<JK@G<HL<GFJJL<D:I<J:@
D<EKF><I8CD<EK<GI@DoI@F<J<DI8D@
:8t{<J
J=FC?8J<D><I8C
:8DI<
LE@;8JEFoG@:<;8GC8EK8ALEK8D<EK<:FD8J@EFI<J:vE:@8J<@E=ILK<J:vE:@8J
(D8@J@DGFIK8EK<uJ89<IHL<JqFGC8EK8J><I8CD<EK<:FDMoI@FJD<KIFJ;<
:FDGI@D<EKF<HL<8J=FC?8JJqF8DGC8J
8DFJKI8HL<GI<:@J8DFJ:FC<K8I
G8I8;<GFJ@K8IEF!<I9oI@FFLG8I8=8Q<IG<JHL@J8J;<M<J<IG8;IFE@Q8;8<
:89<I<DEFJJ8=FC?8;<AFIE8C;F9I8;F:<I:8;<:D:FDGI@D<EKFO
:DC8I>LI8
8>FI8FDFM8DFJ=8Q<I@JJF*L8CG8IK<;<M<I<DFJ:FC<K8I
/F:vJAoEFK8I8DHL<8C>LD8JG8CD<@I8J8GI<J<EK8D<JG@E?FJ*L<
FLKI8JGFJJL<DI8D@
:8t{<J<DJ<LJ:8LC<J-:;17-:HL<<O@JK<D8C
Estipe
>LD8J<JGu:@<J8I9LJK@M8JFDFM8DFJ=8Q<I8EFJJ88DFJKI8 Termo especial-
JK8=8DwC@8uLK@CwJJ@D8G8I8F?FD<D
FIE<:<D8K<I@8C;<:FEJKILtqF mente usado para
M@>8J:FCLE8J<=FC?8JLJ8;8J:FDFK<C?8;F<;@M@JyI@8J
9I8>IFJJ<@I8G8I8 caules de palmeira
:FE=<:tqF ;< LK<EJwC@FJ I<;<J G<E<@I8J :<JKFJ <JK<@I8J :?8GuLJ DyM<@J – Caule ereto, mais
:FI;8JM8JJFLI8J
9I8
E8G8I8K<:<IIFLG8J:FIK@E8J<K8G<K<JGC8EK8J;< ou menos cilíndri-
;<:FI8tqF<G8@J8>@JDF:<I8J:8IE8}98yC<FJ;<E;v:F:F=ILKFJKpD8I8J
onde as folhas con-
8t8w:F:F9LK@oGIFK<wE8J8t}:8II8E>8J8:?8I@=<I8G8CD@KFJLK<IG< centram-se apenas
JJG
M@E?FJ;<J<@M8<;FJ=ILKFJ=<ID<EK8;FJ=8I@E?8J<FLKIFJGIF;LKFJ no ápice (Gonçalves
8C@D<EK8I<J:FDFFJ8>L&<KIFO@CLDILDG?@@
,<IM<D;<=FII8><DG8I8 & Lonrenzi 2007).
FJ8E@D8@J;FDuJK@:FJ?FJG<;8D<8C@D<EK8DLD8>I8E;<M8I@<;8;<;<=FI
D8J;<M@;8J@CM<JKI<
(JLJFJEqFJqF8G<E8J8IK<J8E8@J
:<I8;<:8IE8}98
GFI<O<DGCFK<D8GC@:8t{<J8<IFEoLK@:8JDL@KF<JG<:@8@J<uGI<:@FJ8G8I8
I<JK8LI8;FI<J;<F9I8J;<8IK<
:FDFM8DFJ:FC<K8I<O<DGC8I<JG<IK<E:<EK<J8<JK8=8DwC@8/F:vJAo
;<M<DK<I@D8>@E8;FHL<EqFGI<:@J8DFJ:FC<K8IF<JK@G<
+<JK8D8J=FC?8J<@E
FI<J:vE:@8J@E=ILK<J:vE:@8JHL<JqF><I8CD<EK<>I8E;<J<;<;@=w:@CGI<EJ8><D
DI<C8tqFnJ=FC?8J:FIK8 J<:8;8LD8<DKIvJG8IK<J<JG<:@
:8E;F898J<:FD
8C<KI8FD<@F:FD8C<KI8&<FoG@:<:FD8C<KI8
,<8@EFI<J:vE:@8<
FL@E=ILK<J:vE:@8=FI<D>I8E;<JGF;< J<;F9Io C8J<D;L8JFLKIvJG8IK<JG8I8
:FEJK@KL@I88DFJKI8
9Io:K<8HL<GIFK<><<JK8J<JKILKLI8JK8D9uDu98JK8EK<
@DGFIK8EK<<;<M<J<ID8EK@;8E88DFJKI8
<D8@J@E=FID8t{<JJF9I<8CKLI8
GI<J<Et88LJvE:@8;<<JG@E?FJ;<M<DJ<I8EFK8;8JEF:8;<IEF;<:8DGF
@>LI8O<DGCFJ;<I<:8:<8<
FHL<@IF
<E;<Q<@IF
)8CD@K<@IF
FEK<"EK<IE<K
,@K<J<D+<=<IvE:@8J@9C@F>Io :8J
27
Botânica Sistemática e Econômica
LC9FJ8J
'8JGC8EK8J9LC9FJ8JuE<:<JJoI@FnJM<Q<JJ<::@FE8IFJ9LC9FJ<D
FLG8IK<J
@>LI8 O<DGC8I<J9LC9FJFJ;8=8DwC@8D8IPCC@;8:<8<
FEK<"EK<IE<K
,@K<J<D+<=<IvE:@8J
@9C@F>Io
:8J
HLoK@:8J
J8HLoK@:8JDL@KF=Io><@JJqF;@JK<E;@;8JJF9I<LD8=FC?8;<G8G<C
8@E;8HL8E;F;<EKIF;FI<:@G@<EK<:FDo>L8J<:8D8F8IJF9I<FGIyGI@F
G8G<CFE;<=FI8D<JK<E;@;8J
@>LI8O<DGCFJ;<GC8EK8J8HLoK@:8J
FEK<
)
)I8K8
'8J:@D<EKF #}E@FI
28
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
CFI<J
-I8K8E;F J<;<FI<JuD8@J8:<IK8;F;<@Oo C8J<DoC:FFCGFI?FI8J
FLLDGFL:FD8@J
EK<J;<GI<EJo C8J:FEMuD@JFC8IK8EKFHL8EKFGFJ
1
JwM<C8JGuK8C8JLD8J;8JFLKI8JG8I8<M@K8IHL<D8E:?<D
D
!FA< F F9A<K@MF ;8 FI>8E@Q8tqF ;< LD ?<I9oI@F EqF u D8@J F ;<
J@DGC<JD<EK< :FC<:@FE8I GC8EK8J G8I8 J<I<D @;<EK@
:8;8J < :8K8CF>8;8J
,<LF9A<K@MFuDL@KFD8@J8DGCFGF@JM@J8K8D9uD=FIE<:<I;8;FJG8I8
=LKLIFJ <JKL;FJ <:FCy>@:FJ <
KF><F>Io
:FJ
FD @JJF F ?<I9FI@Q8;FI
8;HL@I<FLKI8JF9I@>8t{<J8CuD;<:FC<K8I<JGu:@D<J
,L8K8I<=8GI@E:@G8C
u8;<J<ILDF9J<IM8;FI:FEJ:@<E:@FJF<<O8KF;<:LA8J<O:LIJ{<J8CuD
;8:FEKI@9L@tqFD8K<I@8C<DGC8EK8JJ<:8JI<JLCK<D<JKL;FJ<F9J<IM8t{<J
8LKvEK@:8J<:@<EKw
:8JJF9I<8M@;8M<><K8CJL8J:FE;@t{<J
J@:F 9@FCy>@:8J
<8JI<C8t{<J<EKI<FD<@F<8FI8E8I<>@qFG<I:FII@;8<D:8;8LD8;8J
<O:LIJ{<J
)8I88JJ@DJ<IuGI<:@JF<DGI@D<@IFCL>8IF9J<IM8I<<DJ<>L@;8
8EFK8I<JJ8JF9J<IM8t{<J;<GI<=<IvE:@8<DLD:8;<IEFJy8@JJF;<JK@E8;F
<ELE:8;<@O8IG8I8D8@JK8I;<FLG8I88MFCK8<JJ8JEFK8J
(LKI8J@E=FID8t{<J:FDFEFD<GFGLC8I<LK@C@;8;<J;8GC8EK8G8I88
:FDLE@;8;<CF:8C=I<HLvE:@8EF8D9@<EK<J<M@M<D@JFC8;8JFL<D:FDL
E@;8;<J:FEJK@KL<D;8;FJI<C<M8EK<J
E
DKF;8<HL8CHL<IEFK8JF9I<8J
G8IK@:LC8I@;8;<J;8GC8EK8uJ<DGI<M8C@FJ8G8I8<JKL;FJGFJK<I@FI<J;<J;<
HL<EqFJ<A8GFJJwM<CF9J<IMo CFJE8GC8EK8J<:8
29
Botânica Sistemática e Econômica
(HL<uLD?<I9oI@F
.D?<I9oI@FuLD8:FC<tqF:@<EKw
:8;<GC8EK8JGI<J<IM8;8JGIFM<E@
<EK<J;<;@M<IJ8JI<>@{<J><F>Io
:8J
@>LI8O<DGCFJ;8<JKILKLI8=wJ@:8;<8C>LEJ!<I9oI@FJ
)8I@J )
.E@M<IJ@;8;<;<,qF
)8LCF ,)
D9I8G8I8JwC@8'
!<I9oI@F;F'<N2FIBFK8E@:8C 8I;<E'2
FC<t{<J:@<EKw
:8JJqF@DGFIK8EK<JGFI
,<I<DFI<>@JKIFG<ID8E<EK<;<9@F;@M<IJ@;8;<
,<I<D898J<G8I88G<JHL@J8<DDL@K8J;@J:@GC@E8J:@<EKw
:8J
)I<J<IM8I<D8J <JGu:@<J ;<=FID8 8 G<ID@K@I8M<I@
:8tqF ;8 M8C@;8;<
K8OFEzD@:8
CuD;<:FEK<I<D8?@JKyI@8E8KLI8C:FD98J<<DJ<LJ8:<IMFJGF;<
J<:I@8IDF;<CFJ;<GI<M@JqFHL<GF;<DJ<ILJ8;FJG8I8;<:@J{<JGFCwK@:8J
FLEFGC8E<A8D<EKF;<G<JHL@J8J=LKLI8J
KI8MuJ;8J:FC<t{<JGF;< J<K<I8:<JJF8@E=FID8t{<JJF9I<F8LD<EKF
<D89LE;pE:@8<8;@JKI@9L@tqF;<;<K<ID@E8;8<JGu:@<FLJF9I<F;<:CwE@F
;<FLKI89<D:FDF<JK89<C<:<IG8;I{<J9@F><F>Io
:FJ?@JKyI@:FJG8I88J
<JGu:@<J
,qFK8D9uD@DGFIK8EK<J@EJKILD<EKFJ;<<;L:8tqF8D9@<EK8C
30
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
@E=FID8E;F;@I<K8FL@E;@I<K8D<EK<FJ<LG}9C@:FJF9I<FG8G<C;8=8LE8
FI8<D@:IF9@FK8EFJ<:FJJ@JK<D8J<8E<:<JJ@;8;<;<JL8GI<J<IM8tqF<J
K@DLC8E;FF@EK<I<JJ<G<CFDLE;FE8KLI8C
,qF@>L8CD<EK<@DGI<J:@E;wM<@J
1
G8I8F;<C@E<8D<EKF;<D<K8J<<JKI8Ku>@8J;<:FEJ<IM8tqF<D8E<AFJLJK<E
K8;F;89@F;@M<IJ@;8;<
FDFu8;@EpD@:8;<LD?<I9oI@F
";<EK@
:8tqFG<ID8E<EK<;<GC8EK8J
)IF:<JJ8D<EKF;8:FC<tqF9FKpE@:8
&8ELK<EtqF;8:FC<tqF
EI@HL<:@D<EKFFL:I<J:@D<EKF;8:FC<tqFGFID<@F;<GIF>I8D8J;<
:FC<K8<@EK<I:pD9@F:FDFLKIFJ?<I9oI@FJ
KL8C@Q8tqF;8@;<EK@
:8tqF;8J<OJ@:8K8J
<LDDF;F><I8CKF;88;@EpD@:8HL<<EMFCM<LD?<I9oI@F:8I8:
K<I@Q8 F:FDFLD8LKvEK@:FC89FI8KyI@FGIoK@:F<;@;oK@:FG8I888HL@J@tqF
;FJ89<I<EMFCM<E;FLOFJ:FEKwELFJ<G<ID8E<EK<J;<:FE?<:@D<EKF
8FG<ID@K@IF8:<JJF<8F9J<IM8tqF;88DFJKI89FKpE@:8
<JK8=FID8
KFIE8 J<@EJKILD<EKF9oJ@:F;<<EJ@EF@EK@D8D<EK<8JJF:@8;Fn=FID8tqF
;<EFMFJG<JHL@J8;FI<J<JG<:@8C@Q8;FJE8J;@=<I<EK<JoI<8J;89FKpE@:8
F=<I<:<E;FI<KFIEF@>L8CD<EK<nJF:@<;8;<:FDF:<EKIF;<;@MLC>8tqF<
<;L:8tqF8D9@<EK8C<;<DFEJKI8E;FJL8<
:o:@8<D8t{<J:FEJ<IM8:@FE@JK8J
J;@D<EJ{<J;<LD?<I9oI@F<FK@GF;<KI898C?F;<J<EMFCM@;F@IqF
;<K<ID@E8IFJDuKF;FJ;<8II8EAF<:LI8;FI@8E<:<JJoI@FJG8I88JL8D8
ELK<EtqF
.D?<I9oI@FGF;<K<ILD<E=FHL<><I8C:FD:FC<t{<JI<GI<
J<EK8K@M8J;<;@M<IJ8JG8IK<J;FDLE;F
<I8CD<EK<JqFFJD8@J8EK@>FJ
<89I@>8D:<EK<E8J;<D@C?8I<JFLD@C?{<J;<<JGu:@D<J9<D:FDFLD8
<OK<EJ8:FC<tqF;<K@GFJEFD<E:C8KLI8@J
(LKIFJ?<I9oI@FJI<<K<D8JG<:
KFJK<DoK@:FJFL?@JKyI@:FJE<JK<:8JF8J:FC<t{<JGF;<DJ<I;@I<:@FE8;8J
8LD8=FID8tqFM<><K8C<JG<:w
:88LD8CF:8C@;8;<8LD:FC<KFI8LD
>ILGF<K:
!oLD8M8I@<;8;<;<?<I9oI@FJEFGC8E<K8
<EKI<FJD8@FI<J<JKqF
F;F&LJ<L;<!@JKyI@8'8KLI8C<D)8I@J):FD
<O<DGC8I<J
< F ;F +FP8C FK8E@: 8I;<EJ $<N $ :FD
;< <O<DGC8I<J
'FI8J@CFJD8@FI<JJqFFJ;F&LJ<L'8:@FE8C;F+@F;<#8E<@IF+
:FD
<F;F#8I;@D9FKpE@:F;F+@F;<#8E<@IF+:FD
8DFJKI8J;<GC8EK8J
(I8J@CK<D ?<I9oI@FJ8K@MFJ<J<D@E=FID8t{<J8KL8C@Q8;8J
31
Botânica Sistemática e Econômica
8:FDG8E?8E;FF@E>I<JJFGFI:FC<K8J<FLG<IDLK8<;F8tqF<
>
8I
9FJ8)<@OFKF
(J?<I9oI@FJ9I8J@C<@IFJ<JKqFC@>8;FJ8DLJ<LJFL
LE@M<IJ@;8;<J < =8:LC;8;<J < =LE:@FE8D =I<HL<EK<D<EK< :FDF :<EKIFJ
8K@MFJ;<KI<@E8D<EKF<=FID8tqF;<I<:LIJFJ?LD8EFJ<DFKpE@:8<J
G<:@8CD<EK<<D,@JK<DoK@:8
JJ@DJ<DGI<?oLDMFCLD<:FEJ@;<IoM<C;<
D8K<I@8C;8E;F<EKI8;8EF8:<IMF
)FIuD8J@DGC<J8K@M@;8;<;<I<>@JKIF<
:8K8CF>8tqF;FJ<JGu:@D<J@E>I<JJ8E;F:FJKLD8<JK8IDL@KF;<=8J8;8GFI
=8CK8;<G<JJF8CKu:E@:F@DGFJJ@9@C@K8E;FFGIFEKF8:<JJF8<C<J
'F:8JF;<LDG8wJ;<K<EKFI;<LD8;8JFI8JD8@JI@:8J;F>CF9F
:FD LD8 8CK8 ;@M<IJ@;8;< <JG<:w
:8 <JK@D8 J< LD MLCKF ;8 FI;<D ;<
<JGu:@<J8G<E8J;<8E>@FJG<ID8JG8I8FI8J@C8@DGFIKpE:@8;F
@E:I<D<EKF;8J:FC<t{<Ju@EHL<JK@FEoM<C
FJ89@C?{<J;<8DFJKI8J
HL<J<<JK@D8<O@JK@I;<GFJ@K8;8J<D:FC<t{<J9@FCy>@:8J;FDLE;FKF;F
FJ8:<IMFJ9I8J@C<@IFJ;<KvD8G<E8J:<I:8;<D@C?{<J;<8DFJKI8J:FI
I<JGFE;<E;F8
;FKFK8C
DI<C8tqFnJ:FC<t{<J9FKpE@:8JFKFK8C;<
<JGu:@D<J>L8I;8;FJ<DKF;FJFJ?<I9oI@FJ9I8J@C<@IFJG<I=8QGFL:FD8@J
;<D@C?{<J
"JJFuDL@KFGFL:FI<GI<J<EK8K@MFJ<=FI<D:FDG8I8;FJFJ
DFEK8EK<J8EK<I@FI<J:FD8D<>89@F;@M<IJ@;8;<;FG8wJKFIE8E;Fy9M@8
8E<:<JJ@;8;<;<I<8C@Q8tqF;<D8@J:FC<K8J@EK<EJ@M8J<<OK<EJ@M8J
)FL:F
J<J89<JF9I<8:FDGFJ@tqFFIwJK@:8;<M8JK8J<OK<EJ{<J;FEFJJFK<I
I@KyI@F<K8D9uDF:FE?<:@D<EKFJF9I<FJMoI@FJ8JG<:KFJ;8J@JK<DoK@:8
<
CF><E@8;8D8@FI@8;FJ>ILGFJ;<GC8EK8JE<FKIFG@:8@J<JKo8@E;8DL@KF
8HLuD;F@;<8CJF9I<KL;FD<;@8EK<8J:I<J:<EK<JK8O8J;<;<JKIL@tqF;FJ
?89@K8KJE8KLI8@J
J@E;@:8t{<J<@E=FID8t{<JJF9I<FJ?<I9oI@FJDLE;@8@J<JKqFGI<
J<EK<JEFJ<>L@EK<<E;<I<tF?KKGJ:@N<9
EP9>
FI>J:@<E:<"E;<O!<I
98I@FILD
8JG
(HL<uLD8<OJ@:8K8
OJ@:8K8JJqF8DFJKI8J;<GC8EK8J;<J@;I8K8;8JJ<:8J:FC8;8J<DLD
G<;8tF;<:8IKFC@E89I8E:8;<K8D8E?FG8;IFE@Q8;F<D:D:FDGI@
D<EKFO:DC8I>LI8
:FDG8E?8D8<OJ@:8K8LD8
:?8;<@;<EK@
:8tqF
FE}D<IF;<<EKI8;8EF!<I9oI@FF:8I@D9F:FD8J@>C8;<JJ<!<I9oI@F
<LDG<HL<EF<EM<CFG<G8I8>L8I;8IG8IK<J;8GC8EK8HL<GFJJ8DJ<;<J
GI<E;<I:FDFG8JJ8I;FK<DGF
32
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
@>LI8
OJ@:8K8J;F!<I9oI@F;8.E@M<IJ@;8;<<;<I8C;<,<I>@G<,
FEK<
&8KFJ
(J?<I9oI@FJ><I8CD<EK<JqF89I@>8;FJGFI@EJK@KLKFJ;<G<JHL@J8<
FL <EJ@EF
( <JG8tF E<:<JJoI@F G8I8 F 8:FE;@:@FE8D<EKF ;8J :FC<t{<J
:FEJK@KL@ J<E8D8@FI@8;8JM<Q<JEFGI@E:@G8CGIF9C<D8<E=I<EK8;FG<C8J
:LI8;FI@8J<><IvE:@8J
(<JG8tF=wJ@:F;<M<JLGFIK8I=LKLI8J<OG8EJ{<J
AoHL<8J:FC<t{<JJqF;@EpD@:8J<K<E;<D8:I<J:<I8CuD;8J8K@M@;8;<J
;< :LI8;FI@8 GI<G8I8tqF < :FEJ<IM8tqF ;< 8DFJKI8J
GyJ 8 :FC<K8 <D
:8DGF8KuFD8K<I@8CJ<I@E:CLw;FEF8:<IMF;F?<I9oI@FFD8K<I@8CG8JJ8
GFI8C>LD8J<K8G8J;<J:I@K8J8J<>L@I
!<I9FI@Q8tqF
FE=FID<Ao=F@M@JKF8EK<I@FID<EK<uFDF;F;<GI<G8I8I8JGC8EK8J
J<:8E;F 8J<EKI<8JGI<EJ8J
JK<DuKF;FuJ@DGC<J<8G<E8JI<HL<ILD
GFL:F ;< G8:@vE:@8 8:FE;@:@FE8 J< :8;8 8DFJKI8 ELD8 =FC?8 ;< AFIE8C
;F9I8;F8J=FC?8JJ<IqF8D8II8;8JE8GI<EJ8<;<M<IqFJ<IKIF:8;8J;@8
I@8D<EK<GFIFLKI8J:FDGC<K8D<EK<J<:8J8KuJ<IGFJJwM<CC<M8I8GI<EJ8
G8I8LD8<JKL=8
K@HL<K8><D
#LEKF8:8;8D8K<I@8C?<I9FI@Q8;F@E:CL@ J<LD8<K@HL<K8;<G8G<C
O
:D:FDFCF:8C<O8KF;8:FC<K8EFD<;F:FC<KFIE}D<IF;8:FC<K8;8K8
<F9J<IM8t{<J
-8@J@E=FID8t{<JJqFK8D9uDI<>@JKI8;8J<DLD8:8;<IE<K8n
G8IK<FE;<F:FC<KFII<>@JKI8KF;8J8JJL8J:FC<K8J<DELD<I8tqFJ<HL<E:@8C
33
Botânica Sistemática e Econômica
&FEK8><D
<GF@J;<?<I9FI@Q8;88GC8EK8uDFEK8;8JF9I<G8G<C:8IKqF:8D@J8
<D><I8C:FJKLI8;8FLGI<J8:FD
K88;<J@M8<:FCF:8;8;<EKIF;<LD8
=FC?8 ;LGC8 J<A8 ;< :8IKFC@E8 FL FLKIF K@GF ;< G8G<C J8@8 F :FEALEKF
=FID8LD8<OJ@:8K8
8K8CF>8tqF<+<>@JKIF
8;8<OJ@:8K8I<:<9<LDE}D<IF;<I<>@JKIF;F!<I9oI@F
:?8;F
:FC<KFIHL<8:FDG8E?88GC8EK8uKI8EJ:I@K8G8I8LD8
:?8;<?<I9oI@F
:FDKF;FJFJ;8;FJFI@>@E8@J8J<I8
O8;8n<OJ@:8K8<FLKI8
:?8@;vEK@:8
u=<@K8G8I8F
:?oI@F;<:FEJLCK8
JJ<
:?oI@FuFI>8E@Q8;F;<=FID8
J@JKvD@:8GFI=8DwC@8J>vE<IFJ<<JGu:@<J
KL8CD<EK<<O@JK<D98E:FJ;<
;8;FJ;@>@K8@JFE;<8J@E=FID8t{<JJF9I<FJ<JGu:@D<JJqF8DGC8D<EK<
;@MLC>8;8J8KI8MuJ;8@EK<IE<K
'FC@MIF;<I<>@JKIF8EFK8 J<GFIFI;<DELDuI@:8FE}D<IF;<I<>@JKIF
;8GC8EK8
(D8K<I@8Cu<EKqF@E:CLw;FEF!<I9oI@F><I8C;<8:FI;F:FD8
JL8@;<EK@
:8tqF
FEJ<IM8tqF
J <OJ@:8K8J JqF :FEJ<IM8;8J ;<EKIF ;< 8IDoI@FJ ?<ID<K@:8D<EK<
=<:?8;FJFL<DC8K8J:FD98JK8EK<E8=K8C@E8<:pE=FI8FLFLKIFJGIF;LKFJ
)C8EK8J?<I9FI@Q8;8JJqFDL@KFJLJ:<KwM<@J8J<I<D8K8:8;8JGFI@EJ<KFJ
D<JDFHL8E;F>L8I;8;8J;<EKIF;8JC8K8J8IDoI@FJ
-F;F:L@;8;F;<M<
J<I;@JG<EJ8;FG8I88JL8:FEJ<IM8tqF
34
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
+<=<IvE:@8J
""D&! 1
<GF@JHL<FD8K<I@8CG8JJ8GFIKF;FFGIF:<JJF8:@D8;<J:I@KF:?<>FL
8?FI8J<AoEqF=F@=<@KFEFDFD<EKF;8:FC<K8;<I<8C@Q8I8@;<EK@
:8tqF
;FD8K<I@8CFL;<:FE
ID8I8HL<C<EFD<8EFK8;F;LI8EK<8:FC<K8
)8I8
I<8C@Q8IDFJ8@;<EK@
:8tqF;FD8K<I@8CGI<:@J8DFJK<I9@9C@F>I8
8<JG<
:@8C@Q8;8HL<8LO@C@<E8@;<EK@
:8tqF<DEwM<C;<=8DwC@8>vE<IF<<JGu:@<
)8I8I<8C@Q8I@;<EK@
:8t{<J<DEwM<C;<=8DwC@8I<:FD<E;8 J<FC@MIF;<
,FLQ8%FI<EQ@HL<=FIE<:<:?8M<J8KL8C@Q8;8JHL<EFJG<ID@K<D
=8Q<I<JJ8@;<EK@
:8tqF
)8I8@;<EK@
:8IDFJ>vE<IFJ<<JGu:@<JKFIE8 J<E<
:<JJoI@888HL@J@tqF;<9@9C@F>I8
8J<JG<:w
:8J:FDFI<M@J{<JK8OFEzD@:8J
K<J<J8IK@>FJ;@M<IJFJJF9I<FJ>vE<IFJ<DHL<JKqF
C>LD8J=8DwC@8JGFJJL<DK<ID@EFCF>@898JK8EK<<JG<:w
:8K8@J:FDF
)F8:<8<<PG<I8:<8<E<:<JJ@K8E;F;<LD:FE?<:@D<EKFD8@J8:LI8;F;8J
<JKILKLI8J<K<ID@EFCF>@8G8I8I<8C@Q8I8@;<EK@
:8tqF
,qF=8DwC@8J:FDLD8
I<;LtqF<M8I@8tqFDFI=FCy>@:8DL@KF>I8E;<
/8C<nG<E8C<D9I8I8E<:<JJ@;8;<;<I<>@JKI8ILDD8K<I@8C K<JK<DLE?F
EF?<I9oI@FJ<DGI<HL<@E@:@8ILD88EoC@J<8E8KzD@:8HLwD@:8=8ID8:FCy>@
:8FLHL8CHL<IKI898C?F:@<EK@
:FHL<<EMFCM88@;<EK@
:8tqF;8<JGu:@<
.D?<I9oI@F><I8CD<EK<GFJJL@<DJ<L8:<IMFD8K<I@8CG8JJwM<C;<J<
I<8C@Q8I:FDG8I8t{<JJF9I<<JGu:@D<J;8FI8CF:8C<9@9C@F>I8
8;@JGFEwM<C
G8I88LO@C@8IE8@;<EK@
:8tqF;FJ<JGu:@D<J
CuD;@JJF8:LI8;FI@8;F
?<I9oI@FI<8C@Q8G<I@F;@:8D<EK<G<IDLK8;<D8K<I@8C:FDFLKIFJ?<I9oI@FJ
G8I8GF;<I8DGC@8I8J@E=FID8t{<JJF9I<;<K<ID@E8;8J<JGu:@<J;@JKI@9L@tqF
><F>Io
:8;8JD<JD8J<8DGC@KL;<;<89I8E>vE:@8;F8:<IMF
FEJLCK88FK8OFEFD@JK8;F!<I9oI@F<nJI<=<IvE:@8J9@9C@F>Io
:8J
JqF<JJ<E:@8@JG8I88LO@C@8IE8@;<EK@
:8tqF:FII<K8;FD8K<I@8C<DHL<JKqF
'80
)8I8I<8C@Q8I8:FE
ID8tqF;8>I8
8;<8C>LEJEFD<J<G8I88LO@C@8IEF
:FE?<:@D<EKFJF9I<8;@JKI@9L@tqF><F>Io
:8;<8C>LD8J<JGu:@<JGF;<DFJ
LK@C@Q8I8C>LEJI<:LIJFJ;@JGFEwM<@JE80
,GC@EB )IFA<KF:FD;8;FJGI@DoI@FJ;<:FC<t{<J?KKGJGC@EB
:I@8
FI>
9I
"E;<O?<I98I@FILD?KKGNNN
EP9>
FI>9J:@@?@?
?KDC:FEK@EL
8D<EK<8KL8C@Q8;F
-+()"(, ?KKGDF9FK
DF9FK
FI>0-,<8I:?M8JK
?KDC
GIFMvEFD<J8:<@KFJ8LKFI<JGL9C@:8t{<J;<KoOFEJ
?KDC@EK<>I8FJwE;@:<J;F+ $<N!8IM8I;.E@M<IJ@KP<LJKI8C@8E
'8K@FE8C!<I98I@LDKI8Q<E;FEFD<J<89@9C@F>I8
89oJ@:88JJF:@8;8;<
KF;8J8J<JG<ID8Ky
K8J
E>@FJG<ID)?PCF><EP0<9J@K<?KKGNNN
DF9FK
FI>&((-
35
Botânica Sistemática e Econômica
+<J<8I:?)N<9<D:FEJK8EK<I<=FIDLC8tqF8GI<J<EK88JD8@JI<:<EK<J
<M@;vE:@8J;<I<8C@E?8D<EKF<D
CF><E@8;<8E>@FJG<ID8J<8C@K<I8KLI8
8JJF:@8;8
?KDKI8Q CF><E@8JI<:<EK<J;<>I8E;<J>ILGFJ;<GC8EK8J<8E@D8@J
'FK<II@KyI@FE8:@FE8C8Go>@E8-8OFEFD@8EFI8J@C?KKGNNN
L=I>J
9IK8OFEFD@8M@E:LC8;8n,F:@<;8;<FKpE@:8;FI8J@C8GI<J<EK8
;8;FJ8KL8C@Q8;FJ;<KF;FJFJ?<I9oI@FJ9I8J@C<@IFJ<;FJG<JHL@J8;FI<J
<JG<:@8C@Q8;FJ<D:8;8>ILGFK8OFEzD@:F
#!H
(I8J@C89I@>88FI8D8@JI@:8;FDLE;FFHL<:<IK8D<EK<<JKoI<C8
:@FE8;FnJL8<OK<EJqFK<II@KFI@8C8JJF:@8;8n<EFID<;@M<IJ@;8;<<;o
:8
:C@DoK@:8<><FDFI=FCy>@:8C<M8E;F8LD88DGC8>8D8;<K@GFJM<><K8:@F
E8@J ".%"--"<K8C
D=8:<;8;@M<IJ@;8;<9FKpE@:8<O@JK<EK<
8KL8CD<EK<<;8I8G@;<Q:FDHL<8C>LEJ8D9@<EK<J9@FD8J<JKqFJ<E;F
8CK<I8;FJ;<>I8;8;FJD<;@8EK<88tqF8EKIyG@:8FG8G<C;FJ?<I9oI@FJu
;<=LE;8D<EK8C@DGFIKpE:@8G8I8>8I8EK@InJ=LKLI8J><I8t{<JF:FE?<:@
D<EKFJF9I<8JGC8EK8J;<;<K<ID@E8;8I<>@qF
(G8G<C;FK8OFEFD@JK8
J<Io=FIE<:<IFEFD<:@<EKw
:F:FII<KF;8JGC8EK8JG8I8HL<<C8JGFJJ8D
J<I<JKL;8;8J<J<KFIE8I<DF9A<KF;<MoI@8J89FI;8><EJJ<=FIF:8JFK8@J
:FDF8EoC@J<J8E8KzD@:8JHLwD@:8JDFC<:LC8I<J;<EKI<FLKI8J<F:FE?<:@
D<EKFJF9I<8HL<C8<JGu:@<GFJJ8J<I8DGC8D<EK<;@MLC>8;F
<M<DFJC<M8I
<D:FEJ@;<I8tqFHL<LD8}E@:8GC8EK8GF;<K<IMoI@FJEFD<JGFGLC8I<J8
;<G<E;<I;8I<>@qF;<F:FIIvE:@8<;<M<DFJKFD8IJ<DGI<F:L@;8;F;<
LK@C@Q8IFEFD<:@<EKw
:FG8I8J<I<=<I@IFLEFD<8I8J<JGu:@<J
<M<DFJ
:FE?<:<I8J=8DwC@8J9FKpE@:8JHL<8GI<J<EK8DG<:LC@8I@;8;<J<DI<C8tqF
8GI<EJ8><D<?<I9FI@Q8tqF9<D:FDFFJ:8I8:K<I<J@DGFIK8EK<JG8I88
JL8@;<EK@
:8tqFG8I8HL<GFJJ8DFJGI<J<IMo CFJ;LI8EK<8:FE=<:tqF;F
D8K<I@8C9FKpE@:F8J<I@E:CLw;FEF?<I9oI@F<OJ@:8K8MFL:?<IG8I8HL<
;<JJ8=FID8FJK8OFEFD@JK8J<JG<:@8C@JK8JGFJJ8D@;<EK@
:8IFD8K<I@8C
8;<HL8;8D<EK<
36
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
!# 1
/@DFJE<JK<:8GwKLCFHL<<O@JK<DKu:E@:8JG8I8:FC<K8I<JGu:@<J;<G<E
;<E;F;<J<LGFIK<<HL<8:FC<K8;<LD8<JGu:@<?<I9o:<8G<HL<E8J<Io
;@=<I<E:@8;8<D8@JJ@DGC<J;FHL<8:FC<K8;<LD8IFD<C@8:<8<HL<K8D
9uDu?<I9o:<8ALJK8D<EK<GFI<JGu:@<J;<JK8=8DwC@88GI<J<EK8I8C>LD8J
;@=<I<Et8JG<:LC@8I<J
CuD;@JJFFLKI8J=8DwC@8J9FKpE@:8J@IqF8GI<J<EK8I
8C>LEJ :8I8:K<I<J K8OFEzD@:FJ @DGFIK8EK<J HL< ;<M<IqF J<I D8EK@;FJ <
<JK8I:FEK@;FJE88DFJKI8G8I8HL<GFJJ8DFJ@;<EK@
:8I<JK<D8K<I@8C:FI
I<K8D<EK<EF=LKLIF
(GC8E<A8D<EKFG8I88:FC<K8uDL@KF@DGFIK8EK<
g
F@Ew:@F;<KL;F
)I<:@J8DFJ=8Q<IMoI@8JF9J<IM8t{<JGI<EJ8IFD8K<I@8C
:8II<>8I<HL@G8D<EKFJ<K:
DLD8J@DGC<J:FC<K8GF;<DFJKI8Q<ID8K<
I@8CG8I8MoI@8J89FI;8><EJ8J89<I88DFJKI8J;<GC8EK8JG8I8GFJK<I@FI
J<:8><DGI<=<I<E:@8CD<EK<:FD;LGC@:8K8J9FI<J=ILKFJ<G8IK<JM<>
<K8K@M8JGI<J<IM8;8J<D
FLoC:FFC8G8I8<JKL;FJ8E8KzD@:FJ
:D8K<I@8CG8I8<JKL;FJ;<:IFDFJJFDFJ;=ILKFJG8I88:8IGFK<:8<
DFJKI8J;<D8;<@I8JG8I8<JKL;FJ8E8KzD@:FJ=D8K<I@8CG8I8<JKL;FJ
HLwD@:FJ>D8K<I@8CG8I88EoC@J<DFC<:LC8I<?DL;8J<JK8:8J<J<D<EK<J
G8I8 GIFG8>8tqF
/@DFJ K8D9uD HL8@J D8K<I@8@J GI<:@J8DFJ C<M8I G8I8
I<8C@Q8I8:FC<K8;<EKI<<C<JD8K<I@8@JG<JJF8@J<D8K<I@8@JG8I8EFJ8LO@C
@8I8I<K@I8I8GC8EK8;FJFCF<:FIK8IFJI8DFJD8K<I@8CG8I8GI<EJ8I<K:
/@DFJHL<;<M<DFJ=8Q<IMoI@8JF9J<IM8t{<J<D:8DGFGI@E:@G8CD<EK<
;8J<JKILKLI8JFL@E=FID8t{<JHL<GFJJ8DJ<IG<I;@;8J;LI8EK<8J<:8><D
:FDF:FI<F;FI
GyJ8:FC<K8M<D8GI<EJ8><DJ<:8><D?<I9FI@Q8tqF
;FD8K<I@8CHL<u8G8IK<;FGI<G8IF;8JGC8EK8JG8I8@E:CL@IEF8:<IMF;F
!<I9oI@FHL<uLD8:FC<tqF:@<EKw
:8;<GC8EK8JGI<J<IM8;8JGIFM<E@<EK<J
;<;@M<IJ8JI<>@{<J><F>Io
:8J
<LDDF;F><I8CKF;88;@EpD@:8HL<
<EMFCM<LD?<I9oI@F:8I8:K<I@Q8 F:FDFLD8LKvEK@:FC89FI8KyI@FGIoK@:F
<;@;oK@:FG8I888HL@J@tqF;FJ89<I<EMFCM<E;FLOFJ:FEKwELFJ<G<ID8
E<EK<J;<:FE?<:@D<EKF8FG<ID@K@IF8:<JJF<8F9J<IM8tqF;88DFJKI8
9FKpE@:8
<JK8=FID8KFIE8 J<@EJKILD<EKF9oJ@:F;<<EJ@EF@EK@D8D<EK<
8JJF:@8;Fn=FID8tqF;<EFMFJG<JHL@J8;FI<J<JG<:@8C@Q8;FJE8J;@=<I<EK<J
oI<8J;89FKpE@:8F=<I<:<E;FI<KFIEF@>L8CD<EK<nJF:@<;8;<:FDF:<EKIF
;<;@MLC>8tqF<<;L:8tqF8D9@<EK8C<;<DFEJKI8E;FJL8<
:o:@8<D8t{<J
:FEJ<IM8:@FE@JK8J
J;@D<EJ{<J;<LD?<I9oI@F<FK@GF;<KI898C?F;<
J<EMFCM@;F@IqF;<K<ID@E8IFJDuKF;FJ;<8II8EAF<:LI8;FI@8E<:<JJoI@FJ
G8I88JL8D8ELK<EtqF
!oLD8M8I@<;8;<;<?<I9oI@FJEFGC8E<K8
<EKI<
FJD8@FI<J<JKqFF;F&LJ<L;<!@JKyI@8'8KLI8C<D)8I@J):FD
<O<DGC8I<J<F;F+FP8CFK8E@: 8I;<EJ$<N$:FD
;<
<O<DGC8I<J
'FI8J@CFJD8@FI<JJqFFJ;F&LJ<L'8:@FE8C;F+@F;<
#8E<@IF+:FD
<F;F#8I;@D9FKpE@:F;F+@F;<#8E<@IF+
:FD
8DFJKI8J;<GC8EK8J
<GF@JHL<FD8K<I@8CG8JJ8GFIKF;FF
GIF:<JJF:@K8;F8EK<I@FID<EK<:?<>FL8?FI8;<I<8C@Q8I8JL8@;<EK@
:8tqF
37
Botânica Sistemática e Econômica
FL;<:FE
ID8I8HL<C<EFD<8EFK8;F;LI8EK<8:FC<K8
)8I8I<8C@Q8IDFJ
8 @;<EK@
:8tqF ;F D8K<I@8C GI<:@J8DFJ K<I 9@9C@F>I8
8 <JG<:@8C@Q8;8 HL<
8LO@C@<E8@;<EK@
:8tqF<DEwM<C;<=8DwC@8>vE<IF<<JGu:@<
KL8CD<EK<
K<DFJ8:<JJF8F8:<IMF;<MoI@FJ?<I9oI@FJ8KI8MuJ;8@EK<IE<KG8I8HL<
<JKL;@FJFJ<<JG<:@8C@JK8JGFJJ8D8E8C@J8I
"$!
8t8LDI<JLDFJF9I<8JGI@E:@G8@JKu:E@:8J;<:FC<K8<D&8>EFC@FG?PK8
%<@88D8KuI@8GL9C@:8;8G<C8=FC?8;<,qF)8LCF<D<;@J:LK8
8@DGFIKpE:@8;FJ?<I9oI@FJ<;FJK8OFEFD@JK8JG8I88;<J:F9<IK8;<EF
M8JGC8EK8J
?KKGNNN
=FC?8
LFC
:FD
9I=FC?8:@<E:@8LCKL
J?KDC
)<JHL@J<E8@EK<IE<KFLEFC@MIF;<,FLQ8<%FI<EQ@<=8t8LDI<
JLDFJF9I<FJ:8I8:K<I<J;@8>EyJK@:FJ;8J=8DwC@8JIFD<C@8:<8<8:K8:<8<
<I<:8:<8<)8CD8<
8I8:K<I<J;@8>EyJK@:FJJqF8HL<C<JHL<EFJ8LO@C@8D
8@;<EK@
:8II<:FE?<:<I8J<JGu:@<J>vE<IFJ=8DwC@8J
v<O<DGCFJ;<
KIvJ<JGu:@<J;<:8;8=8DwC@8JLGI8:@K8;8
/<I@
HL< 8 HL8EK@;8;< ;< <O<DGC8I<J ;F !<I9oI@F ;8 .E@M<IJ@;8;<
<;<I8C;<,<I>@G<,
O&#
'8GIyO@D88LC8@I<DFJ<JKL;8I8J @DEFJG<ID8JLD>ILGF;<GC8EK8J
:FDJ<D<EK<JEL8JEqF<E:<II8;8J<DFMoI@FJ
M!
&(+",
<K8C
)5<)3,-4)5-26,60-9*S916.)5-96/T41+6
8
<;
<EKIF;<)<JHL@J8J;F8:8L"C?uLJ
".%"--"
&
+)"'"
'+&
#
*."+(3%
)
,"%/#
&
(I>J
3)5;):9)9):,69):13
<CF!FI@QFEK<
FEJ<IM8tqF"EK<IE8:@FE8C;FI8J@C
G
('e%/,
%(+'3"!
69.636/1)$-/-;)3(I
>8EF>I8
8<@:@FEoI@F"CLJKI8;F;<&FI=FCF>@8;8J)C8EK8J/8J:LC8I<J
'FM8(;<JJ8,)"EJK@KLKF)C8EK8ILD
"% ((
('('"/
%
+
"W+51+):,- +63-;) 79-:-9=)VU6
-0-9*691A)VU6,-4);-91)3*6;T51+6
,qF)8LCF"EJK@KLKF;<FKpE@:8
38
Métodos e Técnicas de Coleta Vegetal em Angiospermas
Aula
('-*.+)
1++165)916,-*6;S51+)
8I:<CFE8;@KFI@8CC89FI
#(%2
FKpE@:8 5;96,<VU6R;)?65641)=-/-;)C
<;
,qF)8LCF
1
FDG8E?@8;@KFI8'8:@FE8C
%",'++
1-3,;-+0518<-:<:-,*@1::6<916;)51+)3)9,-5
"'?KKGEJ;C
FI>I<JFLI:<
-
)"%%+/
-:+91VU6,-+64<51,),-:=-/-;)1:
.+ ,<
G8IK8D<EKF;<FKpE@:8
@JGFEwM<C<D?KKG<:FHL8
<:FCF>@8
L=I>J
9I
C8JJ@
:8tqFM<><K8C<FJGI@E:@G8@JKoOFEJ
@JGFEwM<C<D?KKGNNN
L==
9IC89<J=8ID8:@8;@;8K@:F:C8JM<>
?KD
+/')
!
/+-+
"!!(+',
1636/1)=-/-;)3
<;
+@F;<#8E<@IF+# L8E898I8$FF>8E
+"33"'"
-
"9);),6,-_;6/-6/9)_),69):138JG<:KFJ<:FCF>@:FJ
JF:@FCF>@:F<FI@JK@:FJ
+@F;<#8E<@IF+#D9@KF:LCKLI8C
,(.3/
%(+'3"!
6;T51+):1:;-4S;1+)>L@8@CLJKI8;FG8I8
@;<EK@
:8tqF;8J=8DwC@8J;<8E>@FJG<ID8J;8FI89I8J@C<@I898J<8;F<D
) ""
'FM8(;<JJ8,)"EJK@KLKF)C8EK8ILD
,"-,(',.%-(,
?KKG@D8><J
>FF>C<
:FD
9I@D8><J>9M?CGK +H;<E;<Q<@IF
J8'JK8IKE;JG
?KKGNNN
A8I;@D;<FI<J
:FD
9IFI<J<=FC?8J#) ,@K?<879CL
D<E8M@8
AG>
?KKG
9G
9CF>JGFK
:FD
?KKG8IMFI<J;<J8FG8LCF
C<J
NFI;GI<JJ
:FD
?KKG@D8><J
>FF>C<
:FD
9I@D>I<J@D>LIC?KKGNNN
GFIK8CJ8F=I8E
:@J:F
:FD
9I
39