VII Congresso Latinoamericano de Paleontologia de Vertebrados programa e resumos

PROGRAMAÇÃO &
LIVRO DE RESUMOS
- programa e libro de resumenes - meeting program and abstract book -
1
VII Congresso Latinoamericano de Paleontologia de Vertebrados
19 a 22 de julho de 2022
Vitória da Conquista, Bahia, Brasil
EDITOR
Mário André Trindade Dantas
REALIZAÇÃO
Laboratório de Ecologia e Geociências
Instituto Multidisciplinar em Saúde
Universidade Federal da Bahia
CONTATO
viiclpv@gmail.com
SITE
www.even.com.br/viiclpv
ii
Evento Virtual – Vitória da Conquista, Bahia, Brasil
COMISSÃO ORGANIZADORA
Dr. Mário André Trindade Dantas (UFBA/IMS-CAT)
Dr. Hermínio Ismael de Araujo Jr (UERJ)
Dra. Carolina Saldanha Scherer (UFRB)
Dra. Simone Moraes (UFBA)
M.Sc. Lidiane de Asevedo Silva (UFS)
M.Sc. Thaís R. Pansani (UFSCAR)
M.Sc. Laís Alves Silva (UERJ)
M.Sc. André Vieira de Araujo (IFPE/UERJ)
COMISSÃO CIENTÍFICA
Dr. Agustín Martinelli
Dr. Alexander Wilhelm Armin Kellner
Dr. Alexandre Liparini
Dra. Ana Maria Ribeiro
Dra. Carolina Saldanha Scherer
Dr. Celso Lira Ximenes
Dr. Cesar Schultz
Dr. Douglas Riff
Dr. Édison Vicente Oliveira
MSc. Ednair Rodrigues Nascimento
Dr. Erwin González-Guarda
Dr. Estevan Eltink
Dr. Felipe Lima Pinheiro
Dr. Fellipe Muniz
Dr. Gustavo Darlim
Dr. Jose Luis Prado
Dr. Julio Marsola
MSc. Kamila L.N. Bandeira
Dr. Kleberson Porpino
Dr. Leonardo Lobo
Dra. Lucy Souza
Dra. Márcia Silva
Dra. Mariela Castro
Dr. Max Cardoso Langer
Dra. Pati Pérez
Dra. Rafaela Missagia
Dr. Renan Bantim
Dra. Renato Pereira Lopes
Dr. Rodrigo Parisi
MSc. Silvio Onari
Dra. Taissa Rodrigues Marques da Silva
Dr.Thiago Marinho
iii
SUMÁRIO
PROGRAMAÇÃO ........................................................................................................................ 7
SESSÃO MESOZÓICO .............................................................................................................. 14
UM ENIGMÁTICO NOVO ARCOSSAURO DE ARMADURA DO TRIÁSSICO SUL-BRASILEIRO
(CARNIANO, ZONA DE ASSOCIAÇÃO DE HYPERODAPEDON) ................................................ 14
REVIEWING THE DIVERSITY OF “SICKLE CLAWS” IN DROMAEOSAURIDAE (DINOSAURIA,
THEROPODA) ....................................................................................................................... 15
PATTERNS OF EARLY PAN-AVIAN DIVERSIFICATION ............................................................ 16
A HISTORICAL REVIEW OF COMPSOGNATHIDS IN THE 1800s ............................................. 17
NOVOS REGISTROS DE DINOSAURIA (SAUROPODOMORPHA) DA SEQUÊNCIA CANDELÁRIA
(TRIÁSSICO SUPERIOR, SUPERSEQUÊNCIA SANTA MARIA).................................................. 18
THE FOSSIL RECORD QUALITY OF BRAZILIAN SAUROPODOMORPHS (SAUROPODOMORPHA:
DINOSAURIA): A PRELIMINARY STUDY ................................................................................. 19
A PRELIMINARY OVERVIEW ON APPENDICULAR CHARACTERS WITHIN TITANOSAURIA
(SAUROPODA; SAURISCHIA) ................................................................................................. 20
AMAZONIAN TYRANT: REAPPRAISAL OF A THEROPOD TOOTH FROM THE CRETACEOUS
ALTER DO CHÃO FORMATION, BRAZIL ................................................................................ 21
PALEOECOLOGY AS A CORRELATION BETWEEN HABITAT USE AND FEEDING BEHAVIOR OF
Baurusuchus FROM THE LATE CRETACEOUS, BRAZIL ......................................................... 22
DESCOBERTA INÉDITA DE COPRÓLITOS DE VERTEBRADOS NA FORMAÇÃO AÇU, BACIA
POTIGUAR, (CRETÁCEO SUPERIOR) ..................................................................................... 23
ISOTOPIC NICHES (δ13C, δ18O) OF PREDATORS FROM THE LAJE DO CORINGA FLAGSTONE
(ALCANTARA FORMATION, CENOMANIAN, NORTHEASTERN BRAZIL) ................................... 24
PALEOHISTOLOGY OF AN AVIAN (ORNITHUROMORPHA) BONE FRAGMENT FROM THE
LÓPEZ DE BERTODANO FORMATION (MAASTRICHTIAN), ANTARCTICA ............................... 25
NEW PARAVIAN THEROPOD REMAINS FROM THE TURONIAN OF THE ARAÇATUBA
FORMATION (PARANÁ BASIN, BAURU GROUP) ..................................................................... 26
THERE AND BACK AGAIN: CHARACTERS, AND ONLY NOT TAXA MUST BE EXPLORED FOR A
CRITICAL REAPPRAISAL OF TITANOSAURIA SYSTEMATICS .................................................. 27
ATHLETIC CAPABILITY IN PTEROSAURS .............................................................................. 28
SYNONYMY BETWEEN Brasileodactylus araripensis AND Cearadactylus atrox (PTEROSAURIA,
ANHANGUERIDAE; APTIAN, ROMUALDO FORMATION) ......................................................... 29
RESISTÊNCIA DA COROA DENTÁRIA EM TERÓPODES NÃO-AVIANOS (DINOSAURIA)........... 30
SYSTEMATIC REVIEW ON NON-AVIAN DINOSAUR TEETH FROM CRETACEOUS BRAZIL ...... 31
A CAVIDADE CEREBRAL DE Clevosaurus brasiliensis (LEPIDOSAURIA, RHYNCHOCEPHALIA)
DO TRIÁSSICO SUPERIOR DO SUL DO BRASIL .................................................................... 32
JUVENILE SPECIMEN OF Notosuchus terrestris FROM BAJO DE LA CARPA FORMATION
(UPPER CRETACEOUS), NEUQUÉN, ARGENTINA .................................................................. 33
WHAT DID THEY EAT? USING STABLE ISOTOPES TO STUDY THE DIET OF A TRIASSIC
VERTEBRATE FAUNA FROM BRAZIL .................................................................................... 34
iv
TEETH OF Priohybodus arambourgi (HYBODONTIFORM) FROM THE TUCANO BASIN (ALIANÇA

FORMATION - JURASSIC): MORPHOLOGY, MORPHOMETRY AND HISTOLOGY ..................... 35
PALEOHISTOLOGIA DE ESCAMAS DE PEIXES TELEÓSTEOS DO GRUPO SANTANA
(CRETÁCEO INFERIOR), BACIA DO ARARIPE ........................................................................ 36
A NEW PLIOSAUR FROM THE PAJA FORMATION OF COLOMBIA SUPPORTS HIGH TOP
PREDATOR DIVERSITY DURING THE EARLY CRETACEOUS ................................................. 37
NEW RECORD OF Brasilosuchus mendesi (CROCODYLIA: TOMISTOMINAE?) IN CAJUEIRO
LOCALITY, BOCA DO ACRE-AM ............................................................................................ 38
FIRST FOSSIL RECORD OF Melanosuchus cf. Melanosuchus niger (CAIMANIDAE:
CROCODYLIA) ON CAJUEIRO SITE, BOCA DO ACRE, AMAZONAS ........................................ 39
ESTUDO COMPARATIVO DE ESPINHOS FÓSSEIS DE CTENACANTIFORMES (FORMAÇÃO
PEDRA DE FOGO, ESTADO DO TOCANTINS) ........................................................................ 40
SIMPÓSIO EVOLUÇÃO DOS ROEDORES ................................................................................. 41
EL REGISTRO FÓSIL DE LOS PAN-CHINCHILLIDAE (RODENTIA, CAVIOMORPHA): SU
IMPORTANCIA BIOESTRATIGRÁFICA Y PALEOAMBIENTAL ................................................... 41
LATE MIOCENE DINOMYIDAE, NEOEPIBLEMAE, AND HYDROCHOERINAE CAVIOMORPHS
(RODENTIA: HYSTRICOGNATHI) FROM ENVIRA RIVER, NORTHERN BRAZIL ........................ 42
ADVANCES IN THE STUDY OF NEOEPIBLEMIDAE RODENTS (HYSTRICOGNATHI:
CAVIOMORPHA) FROM THE LATE MIOCENE OF BRAZIL ...................................................... 43
GIGANTISM EVOLUTION IN CAVIOMORPHS: ASSESSING THE CRANIAL ENDOCAST
ANATOMY AND BRAIN SIZE OF Josephoartigasia monesi (RODENTIA: DINOMYIDAE) ............ 44
REDESCRIPTION OF Neoreomys huilensis FIELD (RODENTIA, HYSTRICOGNATHI) FROM THE
MIDDLE MIOCENE OF LA VENTA (VILLAVIEJA, HUILA, COLOMBIA) ..................................... 45
ANÁLISIS PRELIMINAR DE LA DIVERSIDAD DE CAVIDOS DEL MIOCENO-PLEISTOCENO EN
ARGENTINA .......................................................................................................................... 46
THE AUDITORY REGION OF Neoreomys australis AMEGHINO, 1887 (HYSTRICOGNATHI,
CAVIOIDEA) FROM THE EARLY MIOCENE OF PATAGONIA ................................................... 47
DESCRIPTION OF THE SKULL OF Acarechimys (CAVIOMORPHA, PAN-OCTODONTOIDEA) AND
EARLY EVOLUTIONARY CONSIDERATIONS .......................................................................... 48
SIMPÓSIO TAFONOMIA & PALEOECOLOGIA .......................................................................... 49
NEW OCCURRENCE AND TAPHONOMY OF Myrmecophaga tridactyla LINNAEUS, 1758 IN A
CAVE IN NORTHEASTERN BRAZIL ........................................................................................ 49
BIOLOGICAL AND TAPHONOMIC BIAS OF LISSAMPHIBIAN RECORDS IN SOUTH AMERICA . 50
ESTUDO TAFONÔMICO DE VERTEBRADOS TERRESTRES FÓSSEIS DA FORMAÇÃO AÇU
(?APTIANO-CENOMANIANO DA BACIA POTIGUAR), NORDESTE DO BRASIL.......................... 51
REGISTRO E ASPECTOS TAFONÔMICOS DE UM PHASIANIDAE INDETERMINADO (AVES) EM
DEPÓSITO CÁRSTICO DA BAHIA, BRASIL ............................................................................. 52
RADIOCARBON DATING AND ISOTOPIC PALEOECOLOGY (δ13C, δ18O) OF Glossotherium sp.
FROM MIDWEST BRAZIL (SANTA ELINA, MT) ........................................................................ 53
THE SOUTH AMERICAN FOSSIL RECORDS OF LISSAMPHIBIANS: A TAXONOMIC
PERSPECTIVE ...................................................................................................................... 54
POTENTIAL GEOGRAPHIC DISTRIBUTION OF Notiomastodon platensis (Ameghino, 1888) IN
THE LATE QUATERNARY OF SOUTH AMERICA ..................................................................... 55
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FEEDING ECOLOGY OF LATE MIOCENE GROUND SLOTHS FROM SOUTHWESTERN AMAZON

(SOLIMÕES FORMATION, ACRE BASIN) THROUGH TOOTH MICROWEAR ANALYSIS ............. 56
UMA VISITA VIRTUAL GUIADA À EXPOSIÇÃO PALEONTOLÓGICA DE ANIMAL CROSSING
NEW HORIZONS ................................................................................................................... 57
MÉTODOS VOLUMÉTRICOS NA ESTIMATIVA DE MASSA DE Massetognathus sp.
(MESOTRIÁSSICO, SUPERSEQUENCIA SANTA MARIA) ......................................................... 58
SESSÃO CENOZÓICO .............................................................................................................. 59
PRESENCIA DE Mammuthus columbi EN SANTA LUCIA, ESTADO DE MÉXICO, MÉXICO: UN
REPORTE PRELIMINAR ......................................................................................................... 59
NUEVOS REGISTROS DE MAMÍFEROS DEL PLEISTOCENO-HOLOCENO EN PAMPA DEL
TAMARUGAL, REGIÓN DE TARAPACÁ, CHILE ....................................................................... 60
NOVOS REGISTROS DE SQUAMATA (LEPIDOSAURIA) PARA O QUATERNÁRIO NORDESTINO,
LAJEDO DE SOLEDADE, BACIA POTIGUAR .......................................................................... 61
PALEODIET OF A NEW EXTINCT ALOUATTINAE (PRIMATES: PLATYRRHINI: ATELIDAE):
INFERENCES FROM TOOTH MORPHOLOGY ......................................................................... 62
GLIPTODONTÍDEOS JUVENIS DO PLEISTOCENO TARDIO DA BAHIA ................................... 63
PHYLOGENETIC RELATIONSHIP OF MACRAUCHENIIDAE (MAMMALIA,
PANPERISSODACTYLA, LITOPTERNA) AT GENERA LEVEL .................................................... 64
PALEOECOLOGY OF LATE PLEISTOCENE CAMELIDS FROM THE CHUY CREEK FOSSIL
ASSEMBLAGE OF THE BRAZILIAN PAMPA ............................................................................ 65
ANALYSES OF ENCRUSTED BONES COLLECTED IN LIMESTONES CAVES IN MINAS GERAIS
AND BAHIA ........................................................................................................................... 66
MORFOMETRIA GEOMÉTRICA 2D DO MOLDE ENCEFÁLICO DE FOLIVORA E OUTROS
XENARTHRA ......................................................................................................................... 67
PRIMEIRO REGISTRO DE RESTOS DE MEGAFAUNA QUATERNÁRIA PARA O QUADRILÁTERO
FERRÍFERO .......................................................................................................................... 68
OCORRÊNCIA DE Catonyx cuvieri (XENARTHRA; SCELIDOTHERIIDAE) EM NOVA CAVIDADE
FOSSILÍFERA, NORTE DE MINAS GERAIS, BRASIL ............................................................... 69
SURVEY OF SMALL AND MEDIUM-SIZED MAMMALS FROM THE PLEISTOCENE OF THE
ATLANTIC FOREST ............................................................................................................... 70
TAXONOMIC REVIEW OF THE GENUS Asmithwoodwardia (MAMMALIA:
PANPERISSODACTYLA LITOPTERNA) OF THE PALEOGENE OF SOUTH AMERICA ................. 71
ESTUDO SISTEMÁTICO DO MATERIAL DE TOXODONTIDAE (MAMMALIA, NOTOUNGULATA)
DO QUATERNÁRIO DO ESTADO DA BAHIA ........................................................................... 72
NOVOS ACHADOS PALEONTOLOGÓGICOS PARA O CARSTE DE ARCOS-PAINS-
DORESOPOLIS, MINAS GERAIS ............................................................................................ 73
NEW OCCURRENCE OF Notiomastodon platensis (MAMMALIA, PROBOSCIDEA) IN
AGUALINDA, NORTE DE SANTANDER, QUATERNARY OF COLOMBIA ................................... 74
vi
- Programação -
Terça-feira (19 de julho)
Martes (19 de julio)
Tuesday (july 19)
MINICURSOS
workshops
https://us06web.zoom.us/j/8328
Biologia Sistemática
0745883?pwd=b1hoV3l3TjVRQXR
Dra. Lucy Souza
DK0N3NEZxYU0zQT09
Introdução à osteologia de Theropoda: um guia
teórico-prático para identificar e descrever a
3097523?pwd=MitERGxobDVNcH
morfologia
J1d0JDTFhDWnhrdz09
MSc. Tainá Constância & MSc. Natan Brilhante
Sistemática e Paleobiologia dos dinossauros https://us06web.zoom.us/j/8560
sauropodomorfos 5868999?pwd=aXM0S2llcDhUUV
MSc. Kamila Nogueira JlU2pSbldWUlVVQT09
Origem e evolução de tetrápodes não-amniotas
2555351?pwd=SlpFRlBFOHh3V3I
Dr. Lucas Barcelos & Dr. Rodolfo Santos
5NHdKUCtjTjlTZz09
PALESTRA DE ABERTURA (19h00-20h00)

conferencia de apertura - openning lecture
A microcosmos in a fossil bone
8274202?pwd=c3RqNVlpMWVvUE
Dra. Mirian Pacheco (UFSCAR, Brasil)
93alliRFByS0NTdz09
Quarta-feira (20 de julho)

Miércoles (20 de Julio)
Wednesday (July 20)
PALESTRA (09h00-10h00)
conferencia - lecture
The Value of Enamel Microwear for https://us06web.zoom.us/j/8506
Reconstructing Paleodiet and Paleohabitats 2660324?pwd=L1gzVkNhU2NXbD
Dra. Gina Semprebon (Bay Path University, EUA) A1RTNXK1d3cWpodz09
APRESENTAÇÃO DE TRABALHOS
Presentación de los trabajos – sessions/symposia
Sessão Mesozóico 2891513?pwd=aUNWcWxFeUczU
kMyQmE5L3VkcHBBZz09
UM ENIGMÁTICO NOVO ARCOSSAURO DE ARMADURA DO
10h00-10h20 TRIÁSSICO SUL-BRASILEIRO (CARNIANO, ZONA DE ASSOCIAÇÃO
DE HYPERODAPEDON) - Letícia Rezende de OLIVEIRA et al.
REVIEWING THE DIVERSITY OF “SICKLE CLAWS” IN
10h20-10h40 DROMAEOSAURIDAE (DINOSAURIA, THEROPODA) - Natan Santos
BRILHANTE et al.
PATTERNS OF EARLY PAN-AVIAN DIVERSIFICATION - Pedro L.
10h40-11h00
GODOY & Max Cardoso LANGER
7
A HISTORICAL REVIEW OF COMPSOGNATHIDS IN THE 1800s - Joâo

11h00-11h20
Marcus Vale CAETANO et al.
NOVOS REGISTROS DE DINOSAURIA (SAUROPODOMORPHA) DA
11h20-11h40 SEQUÊNCIA CANDELÁRIA (TRIÁSSICO SUPERIOR,
SUPERSEQUÊNCIA SANTA MARIA) - Debora MORO et al.
THE FOSSIL RECORD QUALITY OF BRAZILIAN SAUROPODOMORPHS
11h40-12h00 (SAUROPODOMORPHA: DINOSAURIA): A PRELIMINARY STUDY -
Carolina PINHO et al.
Sessão Cenozóico 4209044?pwd=UkFleG83dzh6RnV
Tb1Y0VHk4KzEzQT09
PRESENCIA DE Mammuthus columbi EN SANTA LUCIA, ESTADO DE
10h00-10h20 MÉXICO, MÉXICO: UN REPORTE PRELIMINAR - Víctor Adrián
PÉREZ-CRESPO et al.
NUEVOS REGISTROS DE MAMÍFEROS DEL PLEISTOCENO-
10h20-10h40 HOLOCENO EN PAMPA DEL TAMARUGAL, REGIÓN DE TARAPACÁ,
CHILE - Natalia VILLAVICENCIO et al.
NOVOS REGISTROS DE SQUAMATA (LEPIDOSAURIA) PARA O
10h40-11h00 QUATERNÁRIO NORDESTINO, LAJEDO DE SOLEDADE, BACIA
POTIGUAR - Silvia Oliveira LOMBA et al.
PALEODIET OF A NEW EXTINCT ALOUATTINAE (PRIMATES:
11h00-11h20 PLATYRRHINI: ATELIDAE): INFERENCES FROM TOOTH
MORPHOLOGY - Andre Vieira ARAUJO et al.
GLIPTODONTÍDEOS JUVENIS DO PLEISTOCENO TARDIO DA BAHIA -
11h20-11h40
Nancy de F. C. REGO et al.
PHYLOGENETIC RELATIONSHIP OF MACRAUCHENIIDAE
11h40-12h00 (MAMMALIA, PANPERISSODACTYLA, LITOPTERNA) AT GENERA
LEVEL - Leonardo LOBO et al.
Simpósio Evolução dos Roedores https://us06web.zoom.us/j/8803
 Dra. Pati Pérez (Museo Egidio Feruglio, Argentina) 9372934?pwd=RS9jWXFuYUg5Q3
Dr. Leonardo Kerber (CAPPA/UFSM, Brasil) dObHBCb2lpclVCdz09
EL REGISTRO FÓSIL DE LOS PAN-CHINCHILLIDAE (RODENTIA,
10h00-10h20 CAVIOMORPHA): SU IMPORTANCIA BIOESTRATIGRÁFICA Y
PALEOAMBIENTAL – Luciano Luis RASIA
LATE MIOCENE DINOMYIDAE, NEOEPIBLEMAE, AND
HYDROCHOERINAE CAVIOMORPHS (RODENTIA: HYSTRICOGNATHI)
10h20-10h40
FROM ENVIRA RIVER, NORTHERN BRAZIL – Emmanuelle Fontoura
MACHADO et al.
ADVANCES IN THE STUDY OF NEOEPIBLEMIDAE RODENTS
10h40-11h00 (HYSTRICOGNATHI: CAVIOMORPHA) FROM THE LATE MIOCENE OF
BRAZIL – Leonardo KERBER & Francisco Ricardo NEGRI
GIGANTISM EVOLUTION IN CAVIOMORPHS: ASSESSING THE
CRANIAL ENDOCAST ANATOMY AND BRAIN SIZE OF
11h00-11h20
Josephoartigasia monesi (RODENTIA: DINOMYIDAE) – Jose Darival
FERREIRA et al.
REDESCRIPTION OF Neoreomys huilensis FIELD (RODENTIA,
HYSTRICOGNATHI) FROM THE MIDDLE MIOCENE OF LA VENTA
11h20-11h40
(VILLAVIEJA, HUILA, COLOMBIA) – Francisco Javier Urrea
BARRETO et al.
ANÁLISIS PRELIMINAR DE LA DIVERSIDAD DE CAVIDOS DEL
11h40-12h00 MIOCENO-PLEISTOCENO EN ARGENTINA – Maria Carolina Madozzo
JAEN & Pati PÉREZ
8
MESA REDONDA (14h00-16h00)

roundtable
Colonialismo científico na Paleontologia
Dra. Aline Ghilardi (UFRN, Brasil) https://us06web.zoom.us/j/8936
Dr. Juan Carlos Cisneros (UFPI, Brasil) 9819289?pwd=SzFVYWlHQlVXSG
Dra. Bolortsetseg Minjin (ISMD, Mongolia) NGOUQwdUw0VmlTdz09
Dra. Nussaïbah Raja (FAU, Alemanha)
Sessão Mesozóico 1267617?pwd=ejFmMVlqMFAvL0
xUZE9qZUtWU3V4UT09
A PRELIMINARY OVERVIEW ON APPENDICULAR CHARACTERS
16h00-16h20 WITHIN TITANOSAURIA (SAUROPODA; SAURISCHIA) – Maria Luiza
Peres BERTOLOSSI et al.
AMAZONIAN TYRANT: REAPPRAISAL OF A THEROPOD TOOTH FROM
16h20-16h40 THE CRETACEOUS ALTER DO CHÃO FORMATION, BRAZIL – Joaquin
Pedro Bogado DINIZ et al.
PALEOECOLOGY AS A CORRELATION BETWEEN HABITAT USE AND
FEEDING BEHAVIOR OF BAURUSUCHUS FROM THE LATE
16h40-17h00
CRETACEOUS, BRAZIL – Paulo Roberto de Figueiredo SOUTO &
Felipe Mesquita de VASCONCELLOS
DESCOBERTA INÉDITA DE COPRÓLITOS DE VERTEBRADOS NA
17h00-17h20 FORMAÇÃO AÇU, BACIA POTIGUAR, (CRETÁCEO SUPERIOR) –
Thiago Bragança Paduam GONÇALVES et al.
ISOTOPIC NICHES (δ13C, δ18O) OF PREDATORS FROM THE LAJE DO
17h20-17h40 CORINGA FLAGSTONE (ALCANTARA FORMATION, CENOMANIAN,
NORTHEASTERN BRAZIL) – Raquel Soares de MORAES et al.
PALEOHISTOLOGY OF AN AVIAN (ORNITHUROMORPHA) BONE
17h40-18h00 FRAGMENT FROM THE LÓPEZ DE BERTODANO FORMATION
(MAASTRICHTIAN), ANTARCTICA – Bruno Alvez BULAK et al.
Simpósio Tafonomia e Paleoecologia
Dr. Hermínio Araújo Jr. (UERJ, Brasil)
2575244?pwd=SVZablZZb3JWWH
Dra. Carolina Scherer (UFRB. Brasi)
FwbHZWb2dPelB3UT09
Dr. Leonardo Lobo (UFMT, Brasil)
NEW OCCURRENCE AND TAPHONOMY OF Myrmecophaga tridactyla
16h00-16h20 LINNAEUS, 1758 IN A CAVE IN NORTHEASTERN BRAZIL – Lais Alves
SILVA et al.
BIOLOGICAL AND TAPHONOMIC BIAS OF LISSAMPHIBIAN RECORDS
16h20-16h40
IN SOUTH AMERICA – Lucas BARCELOS & Rodolfo SANTOS
ESTUDO TAFONÔMICO DE VERTEBRADOS TERRESTRES FÓSSEIS
16h40-17h00 DA FORMAÇÃO AÇU (?APTIANO-CENOMANIANO DA BACIA
POTIGUAR), NORDESTE DO BRASIL – Leticia Paiva BELFORT et al.
REGISTRO E ASPECTOS TAFONÔMICOS DE UM PHASIANIDAE
17h00-17h20 INDETERMINADO (AVES) EM DEPÓSITO CÁRSTICO DA BAHIA,
BRASIL – Ronaldo Araujo LEONI et al.
RADIOCARBON DATING AND ISOTOPIC PALEOECOLOGY (δ13C, δ18O)
17h20-17h40 OF Glossotherium sp. FROM MIDWEST BRAZIL (SANTA ELINA, MT) –
Thais Rabito PANSANI et al.
9
THE SOUTH AMERICAN FOSSIL RECORDS OF LISSAMPHIBIANS: A

17h40-18h00
TAXONOMIC PERSPECTIVE – Rodolfo SANTOS & Lucas BARCELOS
Simpósio Evolução dos Roedores https://us06web.zoom.us/j/8899
 Dra. Pati Pérez (Museo Egidio Feruglio, Argentina) 2449918?pwd=RE9wN2RYQkNGel
Dr. Leonardo Kerber (CAPPA/UFSM, Brasil) JlMG1mRXBmeFcxQT09
THE AUDITORY REGION OF Neoreomys australis AMEGHINO, 1887
16h00-16h20 (HYSTRICOGNATHI, CAVIOIDEA) FROM THE EARLY MIOCENE OF
PATAGONIA – Maria Eugenia ARANUDO & Pati PÉREZ
DESCRIPTION OF THE SKULL OF Acarechimys (CAVIOMORPHA, PAN-
16h20-16h40 OCTODONTOIDEA) AND EARLY EVOLUTIONARY CONSIDERATIONS –
Michelle ARNAL et al.
Novidades sobre o sítio do Arroyo del Vizcaíno,
Uruguai: em busca de mais evidências de https://us06web.zoom.us/j/8286
presença humana muito antiga num depósito de 1903693?pwd=QmJUVkhyVmt6R
mamíferos gigantes do Pleistoceno UphMVllOEduMm0vdz09
Dr. Richard Fariña (Un. de La Republica, Uruguai)
Quinta-feira (21 de julho)

Jueves (21 de Julio)
Thursday (July 21)
A new pliosaur from the Paja Formation of
Colombia supports high top predator diversity
5618651?pwd=azNJb0pVYUJ2SH
during the early Cretaceous
gxb0RCQU5jQlRFdz09
Dra. Dirley Cortés (McGill University, Canadá)
Sessão Mesozóico 3544856?pwd=L2VNQ0hrcEpOUT
FhY0l4SWVGTms3UT09
NEW PARAVIAN THEROPOD REMAINS FROM THE TURONIAN OF
10h00-10h20 THE ARAÇATUBA FORMATION (PARANÁ BASIN, BAURU GROUP) –
Arthur S. BRUM et al.
THERE AND BACK AGAIN: CHARACTERS, AND ONLY NOT TAXA
10h20-10h40 MUST BE EXPLORED FOR A CRITICAL REAPPRAISAL OF
TITANOSAURIA SYSTEMATICS – Kamila L. N. BANDEIRA
ATHLETIC CAPABILITY IN PTEROSAURS – Daniel Ferreira CEROULA
10h40-11h00
et al.
SYNONYMY BETWEEN Brasileodactylus araripensis AND
Cearadactylus atrox (PTEROSAURIA, ANHANGUERIDAE; APTIAN,
11h00-11h20
ROMUALDO FORMATION) – Rodrigo Vargas PEGAS & Fabiana
Rodrigues Costa NUNES
RESISTÊNCIA DA COROA DENTÁRIA EM TERÓPODES NÃO-AVIANOS
11h20-11h40
(DINOSAURIA) – Tainá Constância de FRANÇA et al.
10
SYSTEMATIC REVIEW ON NON-AVIAN DINOSAUR TEETH FROM

11h40-12h00
CRETACEOUS BRAZIL – Theo Baptista RIBEIRO et al.

roundtable
Aplicações das Tecnologias 3D nas coleções e
exibições
Dr. Sergio Alex Asevedo (Museu Nacional, Brasil)
5848988?pwd=N2ExV0xyVnJWY
Dr. Augustin Ruella (Museo de La Plata, Argentina)
mc1UVJLLzlzdWNxQT09
Dr. Martín Batallés (Un. de La Republica, Uruguay)
Dr. Francisco Garrido (MHNH, Chile)
Dr. David Rubilar (MHNH, Chile)
Sessão Mesozóico 5539426?pwd=dVJtM2ZuYUNId3
BSU2VVN2xiL2RLdz09
A CAVIDADE CEREBRAL DE Clevosaurus brasiliensis
16h00-16h20 (LEPIDOSAURIA, RHYNCHOCEPHALIA) DO TRIÁSSICO SUPERIOR DO
SUL DO BRASIL – Livia Roese MIRON & Annie S. HSIOU
JUVENILE SPECIMEN OF Notosuchus terrestris FROM BAJO DE LA
16h20-16h40 CARPA FORMATION (UPPER CRETACEOUS), NEUQUÉN, ARGENTINA
– Emmanuel Facundo ROUSILLON & Jorge O. CALVO
WHAT DID THEY EAT? USING STABLE ISOTOPES TO STUDY THE
16h40-17h00 DIET OF A TRIASSIC VERTEBRATE FAUNA FROM BRAZIL –
Leonardo CORECCO et al.
TEETH OF Priohybodus arambourgi (HYBODONTIFORM) FROM THE
17h00-17h20 TUCANO BASIN (ALIANÇA FORMATION - JURASSIC): MORPHOLOGY,
MORPHOMETRY AND HISTOLOGY – Estevan ELTINK et al.
PALEOHISTOLOGIA DE ESCAMAS DE PEIXES TELEÓSTEOS DO
17h20-17h40 CRETÁCEO DO GRUPO SANTANA, BACIA DO ARARIPE – Ísis da
Rocha SOUSA & Yuri Modesto ALVES
A NEW PLIOSAUR FROM THE PAJA FORMATION OF COLOMBIA
17h40-18h00 SUPPORTS HIGH TOP PREDATOR DIVERSITY DURING THE EARLY
CRETACEOUS – Dirley CORTÉS et al.
Simpósio Tafonomia e Paleoecologia
Dr. Hermínio Araújo Jr. (UERJ, Brasil)
5819609?pwd=R2RqU3BiZkpHWF
Dra. Carolina Scherer (UFRB. Brasi)
dXR21EbEFERjdmZz09
POTENTIAL GEOGRAPHIC DISTRIBUTION OF Notiomastodon platensis
16h00-16h20 (AMEGHINO, 1888) IN THE LATE QUATERNARY OF SOUTH AMERICA
– Tawane Carvalho Fonte Boa MACHADO et al.
FEEDING ECOLOGY OF LATE MIOCENE GROUND SLOTHS FROM
SOUTHWESTERN AMAZON (SOLIMÕES FORMATION, ACRE BASIN)
16h20-16h40
THROUGH TOOTH MICROWEAR ANALYSIS – Lidiane ASEVEDO et
al.
UMA VISITA VIRTUAL GUIADA À EXPOSIÇÃO PALEONTOLÓGICA DE
16h40-17h00
ANIMAL CROSSING NEW HORIZONS - Beatriz HÖRMANSEDER et al.
11
MÉTODOS VOLUMÉTRICOS NA ESTIMATIVA DE MASSA DE

17h00-17h20 Massetognathus sp. (MESOTRIÁSSICO, SUPERSEQUENCIA SANTA
MARIA) – Pedro Daniel SAMPAIO et al.
NEW OCCURRENCE OF Notiomastodon platensis (MAMMALIA,
17h20-17h40 PROBOSCIDEA) IN AGUALINDA, NORTE DE SANTANDER,
QUATERNARY OF COLOMBIA – Jaime Y. SUÁREZ-IBARRA et al.
A tale of two Gondwanas: late Cenozoic
crocodyliforms of East Africa and response to
7995204?pwd=U2xKNGllOU0zc3o
global and regional environmental change
4a09ob2lmL3pFUT09
Dr. Cristopher Brocchu (University of Iowa, EUA)
Sexta-feira (22 de julho)

Jueves (22 de Julio)
Friday (July 22)
Explorando os lagos permianos do Nordeste do https://us06web.zoom.us/j/8438
Brasil 1172027?pwd=UWxKTXV5T08yN
Dr. Juan Carlos Cisneros (UFPI, Brasil) mdKYWI4YTByTzVnQT09
Sessão Mesozóico 9242002?pwd=enNudlJyUnB1a0F
kczl4L0J1RDRBZz09
NEW RECORD OF Brasilosuchus mendesi (CROCODYLIA:
10h00-10h20 TOMISTOMINAE?) IN CAJUEIRO LOCALITY, BOCA DO ACRE-AM –
Yasmin Catarina Alves DANTAS et al.
FIRST FOSSIL RECORD OF Melanosuchus cf Melanosuchus niger
10h20-10h40 (CAIMANIDAE: CROCODYLIA) ON CAJUEIRO SITE, BOCA DO ACRE,
AMAZONAS – Lavinia Tavarez PAZ et al.
ESTUDO COMPARATIVO DE ESPINHOS FÓSSEIS DE
10h40-11h00 CTENACANTIFORMES (FORMAÇÃO PEDRA DE FOGO, ESTADO DO
TOCANTINS) - Jhonathan Guimarães Sousa COSTA et al.
Sessão Cenozóico 0595945?pwd=cFdibkVBNXpxelB
vc04yUkhCa21UZz09
PALEOECOLOGY OF LATE PLEISTOCENE CAMELIDS FROM THE
10h00-10h20 CHUY CREEK FOSSIL ASSEMBLAGE OF THE BRAZILIAN PAMPA –
Renato Pereira LOPES et al.
ANALYSES OF ENCRUSTED BONES COLLECTED IN LIMESTONES
10h20-10h40
CAVES IN MINAS GERAIS AND BAHIA – Ingrid FERNANDES et al.
MORFOMETRIA GEOMÉTRICA 2D DO MOLDE ENCEFÁLICO DE
10h40-11h00 FOLIVORA E OUTROS XENARTHRA – Roberta Veronese do AMARAL
et al.
12
PRIMEIRO REGISTRO DE RESTOS DE MEGAFAUNA QUATERNÁRIA

11h00-11h20 PARA O QUADRILÁTERO FERRÍFERO – André Gomide
VASCONCELOS et al.
OCORRÊNCIA DE Catonyx cuvieri (XENARTHRA; SCELIDOTHERIIDAE)
11h20-11h40 EM NOVA CAVIDADE FOSSILÍFERA, NORTE DE MINAS GERAIS,
BRASIL – Alexandre LIPARINI et al.
SURVEY OF SMALL AND MEDIUM-SIZED MAMMALS FROM THE
11h40-12h00 PLEISTOCENE OF THE ATLANTIC FOREST – Rodrigo Ventura
GERMANO & Taissa Rodrigues Marques da SILVA

roundtable
Mulheres na Paleontologia Sul-americana
Dra. Mirian Pacheco (UFSCAR, Brasil)
1826843?pwd=TDdpazFQcEZuc2
Dra. Carolina Scherer (UFRB, Brasil)
RQM2lYNTArZEVxdz09
Dra. Dirley Cortés (McGill University, Canadá)
Sessão Cenozóico 1481840?pwd=dFg2MGt5YWc3NX
FEc2dJNU1TcFZXUT09
TAXONOMIC REVIEW OF THE GENUS Asmithwoodwardia
(MAMMALIA: PANPERISSODACTYLA LITOPTERNA) OF THE
16h00-16h20
PALEOGENE OF SOUTH AMERICA – Tábata Zanesco FERREIRA et
al.
ESTUDO SISTEMÁTICO DO MATERIAL DE TOXODONTIDAE
(MAMMALIA, NOTOUNGULATA) DO QUATERNÁRIO DO ESTADO DA
16h20-16h40
BAHIA – Carolina Saldanha SCHERER & Denise Fonseca dos
SANTOS
NOVOS ACHADOS PALEONTOLOGÓGICOS PARA O CARSTE DE
16h40-17h00 ARCOS-PAINS-DORESOPOLIS, MINAS GERAIS – Luciano VILABOIM
et al.
REUNIÃO DE ENCERRAMENTO (19h00-20h00)

reunión de cierre - closing meeting
Escolha do local do futuro evento; premiação da
9325964?pwd=QS9CeTdGa2JPY2l
apresentação dos melhores aluno(a)s; o que ocorrer
JUGUyYlh3VEQzdz09
13
- Sessão Mesozóico -
UM ENIGMÁTICO NOVO ARCOSSAURO DE ARMADURA DO TRIÁSSICO SUL-
BRASILEIRO (CARNIANO, ZONA DE ASSOCIAÇÃO DE HYPERODAPEDON)
Letícia Rezende de OLIVEIRA

l.rezende2707@gmail.com
Voltaire Dutra PAES
voltairearts@gmail.com
Rodrigo Temp MÜLLER
rodrigotmuller@hotmail.com
Leonardo KERBER
leonardokerber@gmail.com
Flávio PRETTO
flavio.pretto@ufsm.br
O Sítio Pivetta (Sequência Candelária, Zona de Associação de Hyperodapedon,

Triássico Superior), localizado em São João do Polêsine, RS, possui diversos
materiais isolados que nunca foram devidamente registrados, incluindo osteodermas,
vértebras, dentes, entre outros elementos, todos de pequeno tamanho.
Recentemente, novos materiais articulados foram encontrados, e ainda que em
preparação, indicam que alguns dos elementos isolados, previamente coletados,
pertencem a um mesmo táxon. A morfologia peculiar dos osteodermos e vértebras
indica que os elementos pertencem a um táxon inédito. O novo espécime apresenta
crânio, vértebras, falanges e possíveis metapodiais, além de distintos morfótipos de
osteodermos, incluindo elementos articulados formando uma carapaça dorsal. As
vértebras possuem o processo transverso expandido lateralmente, representando
cerca de 70% da largura total. Além disso, o espinho neural é reduzido, com sua
altura equivalente a menos da metade da altura do arco neural. Entre os
osteodermos já identificados, os seguintes morfótipos se destacam: 1) osteodermos
paramedianos dorsais (com ampla variação de tamanho), subretangulares em vista
dorsal e com uma eminência dorsal cônica se projetando posterodorsalmente; 2)
osteodermos da coluna lateral dorsal, ainda de posição indefinida, com um formato
retangular em vista dorsal. Esses osteodermos possuem duas eminências dorsais
cônicas, uma no centro e outra próxima a margem lateral; 3) osteodermos laterais
dorsais com uma articulação estilo “tongue-and-groove” e apenas uma grande
eminência dorsal achatada dorsoventralmente e curvada posterolateralmente. Esse
conjunto de características é bastante particular, ocorrendo apenas em alguns
membros de Aetosauromorpha, grupo que inclui os aetossauros e outros
pseudossúquios de armadura. As características dos osteodermos lembram
Euscolosuchus olseni. Além disso, nos materiais articulados, é possível observar três
tipos de osteodermos em linha, como em Acaenasuchus geoffreyi onde há um
osteodermo paramediano e dois osteodermos laterais. As vértebras, com seu espinho
neural extremamente curto, são uma característica marcante do clado formado por
Acaenasuchus e Euscolosuchus. O material é também intrigante por possuir uma
combinação de características de diversos Aetosauromorpha, dificultando num
primeiro momento seu posicionamento filogenético. Entretanto, confirmando-se a
hipótese de os novos espécimes representarem um aetossauromorfo não-aetossauro,
este será o primeiro registro do grupo para o Gondwana, além de representar o
registro mais antigo mundialmente.
14
REVIEWING THE DIVERSITY OF “SICKLE CLAWS” IN DROMAEOSAURIDAE

(DINOSAURIA, THEROPODA)
Natan Santos BRILHANTE

natan.biologia@gmail.com
Tainá Constância de FRANÇA
taina.constancia@gmail.com
Leandro Sanches da COSTA
leandro_sanches84@hotmail.com
Rodrigo Vargas PÊGAS
rodrigo.pegas@hotmail.com
Philip John CURRIE
pjcurrie@ualberta.ca
Rafael DELCOURT
rafael.delcourt@gmail.com
Sergio Alex AZEVEDO
sazevedo@mn.ufrj.br
Dromaeosauridae is a diverse family of theropods dinosaurs generally characterized

by specialized second pedal digits with hypertrophied, strongly recurved, and
trenchant ungual phalanges. Despite the ubiquity of these raptor dinosaurs in the
literature, many aspects of their anatomy remain poorly understood. Because they
are strongly curved, dorsoventrally tall, and lateromedially compressed, many
disarticulated or isolated unguals have been tentatively assigned to this family.
However, ungual phalanges are usually not described in the kind of detail necessary
to determine their relationships. It has led to anatomical misidentifications in some
of these unguals. The characteristics of the second pedal unguals in dromaeosaurid
theropods and possibly in some other basal avialans are re-examined using
qualitative and quantitative analyses. Examining the diversity of “sickle claws” after
an extensive review of the literature, ten features were established for 23 paravian
taxa and 39 specimens to create a practical guide to recognize new records. Some of
these morphological trends are common in most taxa examined, including a broad
lateral cotyle on the articular end (proximal view); the higher position of the blood
groove on the lateral surface; a relatively well-developed flexor tubercle that is lateral
in position; a ventral surface that is offset laterally below the blood groove; and, in
proximal view, the outline is slightly more convex on the lateral side than the medial
side. In addition, our study also reveals relatively subtle anatomical differences
between the relative angles of the outer curvatures of dromaeosaurid unguals in the
same species, and they decrease in relative curvature through ontogeny. Based on
bivariate analyses between the angles of curvature and femoral lengths, no noticeable
correlation was found. At the end of this study, new information on the anatomy of
these “sickle claws” will be dealt with in detail to correctly identify differences and
similarities among dromaeosaurids. These features are conducive to the taxonomic
identification of disarticulated or isolated second pedal unguals, as well as for
determining the side of the body that they come from. [CAPES 88882.183275/2018-
01]
15
PATTERNS OF EARLY PAN-AVIAN DIVERSIFICATION
Pedro L. GODOY
pedrolorenagodoy@gmail.com
Max Cardoso LANGER
mclanger@ffclrp.usp.br
Previous studies provided hypotheses to explain the successful early Mesozoic

diversification of Pan-Aves (=Avemetatarsalia), the lineage of archosaur reptiles closer
to birds than to crocodiles. Some of these suggested that environmental events, such
as climate change and volcanism, were possible drivers, especially during the Late
Triassic. Here, we explore patterns of pan-avian diversification to test the influence of
such events on the group’s radiation. We characterized Triassic to Early Jurassic
terrestrial Pan-Aves diversification (excluding pterosaurs) using phylogenetic-based
and occurrence-based approaches. As diversity metrics, we quantified raw species
richness (using occurrence/count data) and phylogenetic diversity (using time-
calibrated phylogenetic trees). We also estimated net diversification rates using two
methods: PyRate and the episodic fossilized-birth-death (EFBD) model (implemented
in RevBayes), which respectively use occurrence data and time-calibrated trees as
input data. Our results indicate that, when phylogenetic information is taken into
account, diversification events tend to occur earlier in time. For example, occurrence-
based results show a Carnian increase in diversity and higher diversification rates in
this Stage than in the Ladinian. This would be consistent with hypotheses suggesting
that the Carnian Pluvial Episode (CPE) was crucial for the early pan-avian
diversification. However, phylogenetic-based results indicate otherwise, with higher
diversity/diversification rates in the Ladinian, suggesting that the CPE was not a
significant driver of pan-avian diversification. Our results also show no significant
increase in diversity/diversification across the Triassic-Jurassic boundary. Even
though the end-Triassic mass extinction possibly emptied regions of the ecospace for
dinosaurs to expand into during the Jurassic, our results suggest that this
expansion did not occur quickly and homogeneously for the entire group. For the
Early Jurassic, all methods show a sustained reduction in diversity and sub-zero net
diversification rates for the Hettangian and the Sinemurian, with rates increasing
again near the Toarcian. Overall, we did not consistently identify major diversity
shifts across all the results, suggesting that the early diversification of pan-avians
was more nuanced and complex than anticipated. [CAPES 88887.583087/2020-00;
National Science Foundation NSF DEB 1754596; FAPESP 2020/07997-4]
16
A HISTORICAL REVIEW OF COMPSOGNATHIDS IN THE 1800s
João Marcus Vale CAETANO

joaomarcus1997@edu.unirio.br
Luiza PONCIANO
luiza.ponciano@unirio.br
Rafael DELCOURT
The first compsognathid described was Compsognathus longipes, with Wagner

initially classifying it as a lizard instead of a dinosaur. Therefore, he mostly compared
the holotype to squamates, especially regarding cranial bones, but also acknowledged
its oddity regarding the postcranium. Gegenbaur (1863) provided the first extensive
comparison between Compsognathus and birds, suggesting that Compsognathus
presented a tibiotarsus and it corresponded to an intermediate stage between reptiles
and birds. The tarsal region similarity of Compsognathus towards birds was
consistently cited in the literature afterwards. Cope (1867) erected the dinosaur class
Symphypoda, which comprised in a first moment solely Compsognathus, but that
later included Ornithotarsus, Anchisaurus, and (perhaps) Euskelosaurus. The name
Compsognathus gracilis was used by Cope to designate the holotype of C. longipes, C.
gracilis is then invalid. Huxley erected Compsognatha to accomodate Compsognathus
and decided to remove it from Dinosauria, but uniting both into Ornithoscelida. Later
Cope (1871) erected Compsognathidae (Compsognathus) and Ornithotarsidae
(Ornithotarsus) as subtaxa of Symphypoda. Gegenbaur (1874) proposed C. longipes
and Archaeopteryx could be grouped together into a taxon named Saururi,
comprising intermediate forms between ‘reptiles’ and birds. Saururi, however, quickly
became confused with Haeckel's Sauriurae, a class of birds. Owen (1876) described
Aristosuchus pusillus, a dinosaur that later would be classified among the
compsognathids. Initially, Owen referred it to the genus Poekilopleuron and the
crocodilian Family Coelospondylia. But most naturalists believed it was a dinosaur.
Williston argued both Nanosaurus and Compsognathus were the most bird-like of the
fossil reptiles. Owen (1878) erected the species Poekilopleuron minor, which is
claimed by some authors as being a new name for Poekilopleuron (= Aristosuchus)
pusillus. It is understandable because their epithets are words with similar meanings
in Latin (‘pusillus’ = ‘very small’, ‘minor’ = ‘smaller’). Marsh (1879) described Coelurus
fragilis, a possible compsognathid. During the 1800s seven species were classified
within this genus, including possibly Aristosuchus pusillus. Marsh classified all three
compsognathids species within Theropoda, but in diferent families (Compsognathus
in Compsognathidae, Aristosuchus in Megalosauridae, and Coelurus in Coeluridae).
This abstract is part of a larger ‘state of art’ review of Compsognathidae. [CNPQ]
17
NOVOS REGISTROS DE DINOSAURIA (SAUROPODOMORPHA) DA SEQUÊNCIA

CANDELÁRIA (TRIÁSSICO SUPERIOR, SUPERSEQUÊNCIA SANTA MARIA)
Débora MORO
deboramorod@gmail.com
jossano de rosso MORAIS
jossanomorais42@gmail.com
Leonardo KERBER
Rodrigo Temp MÜLLER
rodrigotmuller@hotmail.com
Flávio PRETTO
Descobertas recentes têm contribuído para a compreensão da evolução dos primeiros

dinossauros, sobretudo dos sauropodomorfos. Este grupo é o melhor representado
entre os dinossauros triássicos. Aqui reportamos dois novos materiais de
sauropodomorfos da Sequência Candelária (Zona de Associação de Hyperodapedon),
sul do Brasil, coletados no Sítio Piche, município de São João do Polêsine, Rio
Grande do Sul. Os materiais, em preparação, se preservaram em nível
conglomerático com materiais desarticulados de Dinosauria e de Rhynchosauria, que
recobre um segundo esqueleto articulado de Dinosauria, repousando em nível
pelítico. O espécime desarticulado (CAPPA/UFSM 0244) é preliminarmente atribuído
a um único indivíduo por possuir elementos compatíveis em tamanho, sem
elementos duplicados. A amostra inclui até o momento vértebras de toda a série
axial, costelas fragmentadas, úmero direito, ílio esquerdo, púbis esquerdo, fêmur
direito, tíbia direita e astrágalo esquerdo. O espécime articulado (CAPPA/UFSM
0268) já revelou elementos pós-cranianos da base do tronco até o começo da cauda,
incluindo apêndices articulados e em bom estado de preservação. Em primeira
análise, a morfologia dos espécimes é similar. Ambos possuem úmero com mais de
60% do comprimento femoral; fêmures com cabeça direcionada craniomedialmente,
plataforma trocantérica bem desenvolvida; ílio com a margem ventral do acetábulo
reta, ala pré-acetabular mais curta que o pedúnculo púbico e ala pós-acetabular
alongada. A morfologia geral dos espécimes se assemelha a dos sauropodomorfos da
ZA de Hyperodapedon. O material (assim como Buriolestes) difere de Pampadromaeus
e Saturnalia por não ter uma fossa deltoide (ou qualquer escavação similar) no
úmero. O espécime é ligeiramente mais robusto que os demais espécimes de
Buriolestes, o que pode ser fruto de variação intraespecífica, como observado em
alguns dinossauros triássicos. Entre os elementos ainda em preparação, há também
elementos da mão, ainda pouco conhecidos entre os representantes Carnianos de
Sauropodomorpha. Desse modo, a conclusão da preparação e descrição dos
espécimes, além de futuramente brindar uma atribuição taxonômica menos
inclusiva, ajudará a esclarecer aspectos anatômicos ainda obscuros dentre os
primeiros Sauropodomorpha.
18
THE FOSSIL RECORD QUALITY OF BRAZILIAN SAUROPODOMORPHS

(SAUROPODOMORPHA: DINOSAURIA): A PRELIMINARY STUDY
Carolina PINHO
carolinapinho.g@gmail.com
Maria Luiza Peres BERTOLOSSI
malu.peres@gmail.com
Kamila L.N. BANDEIRA
kamilabandeira@yahoo.com.br
André Eduardo Piacentini PINHEIRO
andre.eduardo.pinheiro@uerj.br
Sauropodomorpha is a clade of predominantly herbivorous dinosaurs that lived

through most of the Mesozoic, especially in South America. However, few studies
have focused on its completeness and how it affects the taxa scoring. In this study,
we evaluate the skeletal completeness of six Brazilian sauropodomorphs (Saturnalia
tupiniquim, Pampadromaeous barberenai, Buriolestes schultzi, Bagualosaurus
agudoensis, Unaysaurus tolentinoi and Macrocollum itaquii) in order to verify the
quality data current used in the phylogenetic analyses. For a better investigation, the
total scoring has been split in cranial, axial and appendicular characters. We use a
more recent cladistic dataset focused on basal sauropodomorphs, which sample has
the largest number of Brazilian sauropodomorphs, with 277 characters. The
Brazilian taxa analyzed here come from Santa Maria and Caturrita formations,
respectively containing four taxa, where two of them pass the 50.0% of total scoring
and two taxa, where only one has a total encoding greater than 50.0%. In general, for
these six taxa, the common elements are the jaw, the dorsal vertebrae and tibia.
Macrocollum and Unaysaurus are the taxa that have more complete skeletal
sampling, with 18 preserved elements, being Bagualosaurus the less sampled among
it, with only 14 preserved elements. However, despite Macrocollum and Unaysaurus
presents a high skeletal sampling, the scoring did not reflect it as we could verify -
Macrocollum has 84.1% of scoring and Unaysaurus has 29.9%. Buriolestes and
Macrocollum are the taxa with the highest percentages of cranial scorings (>80.0%).
Buriolestes is also the more scorable sauropodomorph among analyzed taxa (85.2%),
within a high percentage of axial remains (97.4%), while Macrocollum is the taxon
with highest appendicular scoring (86.8%). Macrocollum has the main representative
scoring average, with cranial, axial and appendicular characters greater than 80.0%.
We observe that the relationship between scoring and completeness is not directly
proportional, as observed here to Macrocollum and Unaysaurus. Given our
preliminary results, two scenarios are speculated: 1 - character hypotheses
formulations need to be better explore the anatomy of sauropodomorphs or, 2 - some
skeletal portions of sauropodomorphs are naturally conservative and in fact have
poor extractable anatomical information for hypotheses of homologies. [FAPERJ]
19
A PRELIMINARY OVERVIEW ON APPENDICULAR CHARACTERS WITHIN

TITANOSAURIA (SAUROPODA; SAURISCHIA)
Maria Luiza Peres BERTOLOSSI

malu.peres@gmail.com
Titanosauria (Sauropoda, Saurischia) is a well-established clade of sauropods,

diverse mainly during the Cretaceous of Gondwana landmasses. However, its
phylogenetic relationships are still far to be resolved. In this scenario, the
appendicular characters are the most consistent in the cladistic analysis for the
group, characterizing some less inclusive clades (e.g., Saltasauridae). In this work,
we provide a preliminary overview concerning the use of appendicular characters
throughout decades. We evaluate the dataset from nine different cladistic analyses
dated from 1997 to 2021 and focused on titanosaur relationships. We observe the
total number of characters, the number of appendicular characters, and the number
of characters about one specific element, being them: sternal plate, humerus, and
femur. We also chose six taxa that bring the elements (Andesaurus, Malawisaurus,
Epachtosaurus, Aeolosaurus, Opisthocoelicaudia, and Saltasaurus) to observe
potential changes in the amount of appendicular character scored over the years. As
result, a total of 2,047 characters were evaluated, where 780 are appendicular
characters. However, their distribution is not equal to the analyzed matrices. The
oldest matrix (from 1997) and two from 2016 are the ones with the highest number
of appendicular characters when compared to their original datasets (47.3%; 37.3%
and 48.6% respectively). The most recent, with 552 osteological characters, has only
35.6% of appendicular ones. The femur and the sternal plate are the most consistent
appendicular elements, being present in all the analyzes observed here with at least
one character since 1997. The femur is also the morphological element that presents
more scoreable features, with an average of six characters in datasets. Andesaurus is
the taxon less scoreable for appendicular characters, scored by nine characters while
Opisthocoelicaudia is the most, within 46 characters scored. Although preliminary,
our study shows that the appendicular characters have lost prominence in cladistic
analyses in the last years, which can be a directional to phylogenetic datasets in the
future.
20
AMAZONIAN TYRANT: REAPPRAISAL OF A THEROPOD TOOTH FROM THE

CRETACEOUS ALTER DO CHÃO FORMATION, BRAZIL
Joaquin Pedro Bogado DINIZ

jpbogadodiniz@gmail.com
Theo Baptista RIBEIRO
theobribeiro1@gmail.com
Paulo Victor Luiz Gomes da Costa PEREIRA
paulovictor29@yahoo.com.br
The record of Mesozoic reptiles in the Northern region of Brazil is extremely limited,
being composed mostly of fossils that have since been lost to science or reinterpreted
as misidentified Cenozoic material. The only definite occurrence of body fossils of
Mesozoic reptiles in the region consists of two ziphodont teeth recovered from an oil
well core in the municipality of Nova Olinda do Norte, AM, from strata of the
Cretaceous Alter do Chão Formation of the Amazonas Basin. One of these teeth is
presently unaccounted for, but the other is now part of the Collection of Fossil
Reptiles of the Museu de Ciências da Terra (MCT), as specimen MCT.R.514. This
tooth was studied by Llewellyn Ivor Price (1905-1980), who regarded it as an
indeterminate carnosaur. Since Price’s original description of the fossil the material
has attracted little attention, being only briefly discussed in stratigraphic studies of
the Amazonas Basin and reviews about the Brazilian dinosaur record. In this study
we aim to reevaluate MCT.R.514 using the most recent methodologies available for
the identification of isolated theropod teeth, such as morphometric (Linear
Discriminant Analyses and Clustering Analyses) and cladistic analyses, in order to
obtain a new and more robust assignment for the specimen. MCT.R.514 showcases a
transitory morphology between a non-tyrannosaurid tyrannosauroid and a pachidont
tyrannosaurid, with a moderately narrow blade-shaped crown and an irregular
enamel texture. This morphology is usually seen in early tyrannosaurids with
ziphodont teeth, such as Alioramus, or in juvenile pachidont tyrannosaurids. Three of
the six morphometric analyses recovered MCT.R.514 as a tyrannosauroid, similar to
tyrannosaurids such as Alioramus or Gorgosaurus. Two of the analyses recovered it
as either an abelisaurid or a non-megalosaurid megalosaurian, and one analysis
brought inconclusive results. The two cladistic analyses assigned MCT.R.514 to a
basal position within Tyrannosauroidea, more derived than Eotyrannus yet more
basal than many tyrannosaurid taxa. MCT.R.514 represents the second known
record of Tyrannosauroidea in South America and the first tooth fossil of this group
in the continent, further expanding our still limited knowledge about the distribution
and dental evolution of Gondwanan tyrannosauroids. [CAPES; FAPERJ]
21
PALEOECOLOGY AS A CORRELATION BETWEEN HABITAT USE AND FEEDING

BEHAVIOR OF Baurusuchus FROM THE LATE CRETACEOUS, BRAZIL
Paulo Roberto de Figueiredo SOUTO

prfsouto15@gmail.com
Felipe Mesquita de VASCONCELLOS
fmv@geologia.ufrj.br.com
We described a paleoecological study based on new remains of fossil crocodyliforms

found in outcrops located in General Salgado, Ibirá, Jales, Adamantina, Monte Alto,
and Votuporanga counties, São Paulo, southeastern Brazil. In the last years the
discovery of fossil eggs and coprolites associated with Baurusuchus sp.
(Sebecosuchia), a Late Cretaceous genus of crocodyliforms, has contributed to the
understanding of their habitat and behavior. The fossil materials from the
Adamantina Formation (Bauru Group) fossil materials are usually preserved in fine-
grained carbonate-cemented sandstones and siltstones (and a few red mudstones) of
Turonian to Santonian age. Recent investigations support that animal behavior plays
a key role in the adaptations of biological functions. The behavioral interpretation
was carried out from the analysis of organic remains preserved within coprolites and
the stomach cavity in association with nesting areas, and sedimentary features. In
most occurrences, the fossil remains of young and adult individuals are found
articulated in proximity and directly associated with vertebrates (coprolites,
eggshells, and nests) and invertebrate ichnofossils. The coprolites are characterized
by cylinder forms, with organic inclusions consisting of bone fragments, teeth, and
scales, with evidence of cannibalism of newborn individuals. The abdominal area
displays bones with chemically etched surfaces by digestive juices with mechanical
abrasion. In at least three almost complete individuals; the abdominal area was
abundant with Taenidium sp. ichnofossils and shallow root-like systems. Some
clutches with hatched ellipsoidal eggs were found in a nest-like semicircular
organization, suggesting they all hatched. This collected evidence corroborates that
Baurusuchus sp. may be a large population. They adapted to a seasonally-controlled
environment displaying extreme precipitation during a brief rainy season, responsible
for the biostratinomic stage of transport and burial (shot-to-no transport and fast
burial, high presence of coprolites, and partially fragmented eggshells and nest-like
clutches), followed by a long-lasting drought season. This strategy is a pattern found
in environments faced by some of the extant species of crocodyliforms and points out
that these fossils evidence is indicative of territorial behavior. [UNIRIO P-0063/2011]
22
DESCOBERTA INÉDITA DE COPRÓLITOS DE VERTEBRADOS NA FORMAÇÃO

AÇU, BACIA POTIGUAR, (CRETÁCEO SUPERIOR)
Thiago Bragança Paduam GONÇALVES

tp-goncalves@hotmail.com
Lilian Paglarelli BERGQVIST
bergqvist@geologia.ufrj.br
Paulo Roberto de Figueiredo SOUTO
prfsouto15@gmail.com
A Bacia Potiguar está localizada na região litorânea do nordeste do Brasil, se

estendendo pelos estados de Rio Grande do Norte e Ceará. Sua formação é
relacionada ao processo de estiramento crustal ocorrido na abertura do oceano
Atlântico durante o Cretáceo. O registro de fósseis de vertebrados na Formação Açu
ainda é pouco expressivo, tendo sido verificada, nos últimos anos, a ocorrência restos
fósseis de peixes e de répteis. Entretanto, apenas recentemente foram documentadas
pela primeira vez, estruturas biogênicas de vertebrados, mais especificamente,
coprólitos. Neste estudo foram analisados mais de cem exemplares de coprólitos
coletados em atividades de campo realizadas pelo Laboratório de Macrofósseis -
UFRJ, no período de 2015 a 2020. A maioria dos exemplares se caracteriza pela
forma cilíndrica, com tamanhos variando de milímetros até seis centímetros de
comprimento e com até cinco centímetros de espessura. Vinte exemplares
apresentam marcas de espiral. O estado de preservação dos coprólitos sugere uma
condição alóctone, com transporte posterior à fossilização, o que é evidenciado pelo
expressivo número de espécimes apresentando polimento da superfície e quebras
parciais. A respeito dos possíveis produtores, segundo as características e a
morfologia estrutural dos exemplares mais bem preservados, os coprólitos cilíndricos
menores e com espirais podem ser associados a peixes cartilaginosos ou ósseos,
enquanto as formas cilíndricas maiores possuem relação com vertebrados de terra
firme, possivelmente crocodilídeos ou dinossauros. A análise da estrutura interna e
composição química estão em fase de elaboração, o que irá contribuir para o
entendimento da dieta e das possíveis interações paleoecológicas da comunidade que
habitou essa região da Bacia Potiguar durante o Cretáceo.
23
ISOTOPIC NICHES (δ13C, δ18O) OF PREDATORS FROM THE LAJE DO CORINGA

FLAGSTONE (ALCANTARA FORMATION, CENOMANIAN, NORTHEASTERN
BRAZIL)
Raquel Soares de MORAES

moraesraquel.bio@gmail.com
Mario Andre Trindade DANTAS
matdantas@yahoo.com.br
Manuel Alfredo MEDEIROS
medeirosalf@gmail.com
Taissa Rodrigues Marques da SILVA
taissa.rodrigues@gmail.com
The Laje do Coringa Flagstone is a bonebed pertaining to the Alcântara Formation,

cropping out along the shore in Cajual Island, in the vicinity of São Luís (Maranhão
state, Brazil). Its fossils are abundant but found incomplete and isolated.
Nevertheless, it has yielded hundreds of specimens, providing a unique glimpse of an
early Late Cretaceous Gondwanan biota. The majority of fossils pertains to
vertebrates, such as theropod and sauropod dinosaurs, crocodylomorphs,
pterosaurs, turtles, and bony and cartilaginous fishes. As is the case for other
deposits of the same age, they represent a large number of predators but few
herbivorous taxa, thus bringing the question of how their ecological niches were
partitioned. We thus analyzed the isotopic niche of selected predatorial taxa as a
proxy for their ecological niche. We obtained the δ13C and δ18O signatures of the
tooth enamel of carcharodontosaurids, spinosaurids and anhanguerians (five
specimens each), of four undetermined elasmobranch dental plates, and of two
hybodontiform dorsal spine fragments. Additional δ13C and δ18O values of three
carcharodontosaurids and eight spinosaurids were taken from the literature. To
estimate their niche breadth and overlap, we ran a multivariate niche analysis using
taxon-specific enrichment values and used the SIBER package in R to extrapolate
Bayesian ellipses. The results of the former indicate that spinosaurids and
anhanguerians were more specialists, while carcharodontosaurids, hybodontiforms,
and the undetermined elasmobranchs were more generalist taxa. Pairwise taxon
comparisons showed moderate or high niche overlaps between all taxa, being lower
between the spinosaurids and the elasmobranchs and hybodontiforms. The analyses
in SIBER showed that anhanguerians had the smallest isotopic niche, closely
followed by the elasmobranchs, and that spinosaurids and carcharodontosaurids
had large niches. Our results agree with widespread evidence that anhanguerians
were specialized piscivorous animals that, despite having some overlap with
elasmobranchs, had a different niche, especially in regard to their δ18O signature: as
terrestrial reptiles, pterosaurs would get water from both their food and by actively
drinking freshwater. Our results also agree with several lines of evidence that point
out that spinosaurids were specialized in certain prey such as fishes, thus differing
from carcharodontosaurids.
24
PALEOHISTOLOGY OF AN AVIAN (ORNITHUROMORPHA) BONE FRAGMENT

FROM THE LÓPEZ DE BERTODANO FORMATION (MAASTRICHTIAN),
ANTARCTICA
Bruno Alves BULAK

brunoabulak@gmail.com
Geovane Alves de SOUZA
geovanesouza@ufrj.br
Esaú Victor de ARAÚJO
esauvictor13@gmail.com
Juliana Manso SAYAO
jmsayao@mn.ufrj.br
Alexander Wilhelm Armin KELLNER
kellner@mn.ufrj.br
Marina Bento SOARES
marina.soares@mn.ufrj.br
The PALEOANTAR project (Museu Nacional-UFRJ) has been conducting its fieldwork
in the James Ross archipelago of the Antarctic Peninsula, specifically in the Late
Cretaceous (Maastrichtian) strata of the López de Bertodano Formation, from which
several tetrapods were recorded, including plesiosaurs, mosasaurs, non-avian and
avian theropods. Concerning the latter, two stem-anseriformes (Ornithuromorpha,
Neornithes) species were previously reported: Vegavis iaai and Polarornis gregorii. In
the expedition 2018-2019, the project’s crew collected in Vega Island a small bone
fragment with an external smooth texture (3 cm in length and 1 cm in diameter),
identified solely as part of an indeterminate long bone. As the fragment did not show
any diagnostic anatomical characteristics, paleohistology was used to refine the
identification. The bone was embedded in epoxy resin for sectioning. One of its sides
was ground and polished with a series of sandpapers, glued onto a petrographic slide
and the free side was further ground and polished. The histological slides were
analyzed under polarized and incident light using a petrographic microscope. The
thick cortex is composed of well-vascularized fibrolamellar bone, with a reticular
vascularization pattern, opening up to the periosteal surface. Cyclical growth marks
are absent. The perimedullary region is occupied by many resorption cavities,
indicating an osteosclerotic bone. A well-defined endosteal lamella can be observed.
These characteristics are consistent with those found in Late Cretaceous
ornithuromorph birds and differ from those of the same sized non-avian theropods
because the latter exhibited less vascularized cortices consisting of slow depositing
parallel-fibered bone. When comparing the histology of the known birds from the
López de Bertodano Formation, the studied specimen is more similar to P. gregorii,
due to its reticular vascularization, than to V. iaai, which shows semi-reticular
grading towards the subperiosteum to longitudinal vascularization. The sample's
thicker cortex shows a relative bone wall thickness (RBT) of approximately 35%,
similar to that of P. gregorii (RBT=37%), supporting an aquatic/diving lifestyle. These
results are relevant both because Cretaceous records of Aves are still rare and
because they guide the work through the already collected material by the
PALEOANTAR project in search of bones with avian structures. [CNPq]
25
NEW PARAVIAN THEROPOD REMAINS FROM THE TURONIAN OF THE

ARAÇATUBA FORMATION (PARANÁ BASIN, BAURU GROUP)
Arthur S. BRUM
arthur7sbc@gmail.com
The paravian theropods reached a wide distribution and diversity during the
Cretaceous, comprising troodontids, dromaeosaurids, and aves. In the Bauru Group,
the paravian record is scarce and fragmentary, with few records from the
Adamantina Formation (Campanian-Maastrichtian sequences) and Marília Formation
(Maastrichtian), with only one species described to the last—the unenlagiine
dromaeosaurid Ypupiara lopai. The fossil vertebrates of the Araçatuba Formation
(Turonian) are still scarce, with only one archosaur species described so far: the
sphagesaurian crocodyliform Coronelsuchus civali. Here, we present new paravian
remains from Araçatuba Formation from the fieldwork of 2019 in the Western São
Paulo state by the LAPA. The specimens comprise three caudal vertebrae (two of
them are associated). The caudals are morphologically similar and from the same
horizon. They exhibit laterally projected transverse processes and tall neural spines,
which are typical to the caudal transition point—mid anterior portion of the tail—in
paravian theropods. The ventral surface of the centra exhibits articular facets to the
articulation of the haemal arches. On the lateral surface, in the infradiapophyseal
region, the specimens are devoid of laminae, which is similar to the observed among
dromaeosaurids and troodontids. The morphology of the zygapophyses is similar to
the observed in the paravians Buitreraptor, Rahonavis, Anchiornis, and
Archaeopteryx, with retracted prezygapophyses and dorsally directed; elongated
postzygapophyses, surpassing the posterior margin of the centrum; and the pre-
postzygapophyseal articulation between the level of the intervertebral articulation
and the mid-length of the centrum. These paravian remains extend the lower
temporal range of the taxon to the Bauru Group, indicating that it was early present
in the Turonian times. Such a report also indicates a hidden diversity in these small-
sized theropods in the Bauru Group. [CAPES 88887.336584/2019-00; FAPERJ
PDR10 E-26/201.995/2020; Proatec/DEPESQ/UERJ]
26
THERE AND BACK AGAIN: CHARACTERS, AND ONLY NOT TAXA MUST BE
EXPLORED FOR A CRITICAL REAPPRAISAL OF TITANOSAURIA SYSTEMATICS

The last few decades have been crucial to understanding the evolutionary traits and
systematics aspects of the Titanosauria clade, with the growing description of almost
complete specimens. However, most of the cladistic analyzes do not contemplate the
diversity of the clade, and the internal relationships are far from being well-
understood. More recent works focused especially on the resolution of instability and
polytomies based primarily on scripts searching clades with higher support and
exclusion of unstable taxa. Here I provide an overview after evaluating 55 datasets
(+3,000 characters) concerning the phylogenetic relationships of Titanosauria (or at
least the Titanosauriformes clade). Most analyses included twelve (or fewer)
titanosaur taxa as terminal. Some incomplete species act as a stable specifier, such
as (1) Andesaurus as basalmost titanosaur and presenting >70% of missing data
(m.d.); (2) Mendozasaurus (>65% of m.d.) and Futalognkosaurus (>68% of m.d.) as a
key to Lognkosauria; and (3) Rinconsaurus (>63% of m.d.) to define Rinconsauria. The
simple inclusion of more complete taxa nor the creation of statistical methods did not
bring great resolutions in analyzes with a greater number of inserted titanosaurs,
causing the pruning of some taxa with a higher score of missing data (>50%). These
results agree with previous work where even fragmentary specimens have enough
combination of features that enable a precise recognition of their evolutionary
affinities. Among morphological characters, the cranial has grown in recent decades,
starting from 2% to 4% in older datasets, but occupying around 29% of characters in
more recent analyses. Appendicular characters, albeit numerous (32%) remain poorly
explored when more specific elements are observed (such as the radii and fibula with
less than 10%). Some morphological characters (>18%) describe variation in
quantitative traits (i.e., ratios), which hinders the coding of more incomplete taxa.
Moreover, some regions of the appendicular skeleton (such as the ischia, hand, and
pes) exhibit remarkable variability and are still underrepresented in phylogenetic
analysis. Osteoderms type and paleohistological features are also practically absent.
Lastly, the detection of these patterns in the cladistic datasets used provides insight
into an ongoing work of titanosaurian lower-level relationships.
[Proatec/DEPESQ/UERJ]
27
ATHLETIC CAPABILITY IN PTEROSAURS
Daniel Ferreira CEROULA

ciroula@gmail.com
Fabiana Rodrigues Costa NUNES
fabianarodriguescosta@gmail.com
Pterosaurs were the first vertebrates to actively fly. They lived during the Mesozoic
Era and exhibited a wide spectrum of shapes and sizes. Because of their unique
morphology, a direct comparison with living vertebrates is not possible. Regarding
terrestrial locomotion, inferences on the terrestrial proficiency of pterosaurs can be
made by calculating Athletic Capability Indices, which measure the resistance of
bones to axial compression and lateral flexion forces. With the aim of determining
these indices in pterosaurs, stylopod bones (humerus and femur) of 25 individuals
(from different pterosaur families with range of mass from 0,10 to 330 kg) were used.
Calculations were made by A/amg for resistance to axial compressions, and Z/amgx
for resistance to lateral bending, in which A stands for the cross-sectional area of the
stylopod shaft, Z is the section modulus, a the fraction that each limb supports
during walking, mg the animal weight and x corresponds to the distance between the
distal portion of the bone to the diaphysis where the cross section was measured.
Values were given in GPa. Comparisons were made between the obtained values and
those of extant (quadruped mammals) and extinct vertebrates (dinosaurs and
terrestrial xenarthrans) from the literature. Values generally ranged from 16 to 276
GPa for the humerus, and 5.1 to 224 GPa for the femur. The highest values of both
indices were found for humerus in 20 individuals. As high values indicate high
relative strength of bones, the here achieved results point to forelimbs with higher
resistance than that of hindlimbs. Values were higher for femur in five individuals,
all of these small (<0,5kg) excepting for Pteranodon longiceps (63kg). Compared to
those of other quadruped vertebrates, these values are consistent with a good
proficiency for terrestrial locomotion for all pterosaurs here analyzed, even for basal
forms previously thought to have little proficiency in terrestrial locomotion.
28
SYNONYMY BETWEEN Brasileodactylus araripensis AND Cearadactylus atrox

(PTEROSAURIA, ANHANGUERIDAE; APTIAN, ROMUALDO FORMATION)

fabianarodriguescosta@gmail.com
The Romualdo Formation (Aptian) is famous for its exquisite three-dimensional fossil
preservation and outstanding pterosaur record. The first pterosaur species
represented by cranial material (a partial mandible) from there was Brasileodactylus
araripensis, described in 1984, followed by Cearadactylus atrox in 1985 (based on an
almost complete skull). Brasileodactylus has been consistently regarded as valid due
to a prominent autapomorphy: a series of subtle lateral grooves connected to the
median occlusal dentary groove (termed here "accessory grooves"). Cearadactylus has
been regarded as distinct from Brasileodactylus mainly due to (1) the absence of
accessory grooves, (2) the presence of a bifurcation of the anterior tip of the median
occlusal groove, located between the first pair of alveoli (autapomorphy), and (3) a
distinct pattern of rosette alveoli spacing. However, we notice that the bone surface of
the dentary of the holotype of C. atrox is quite weathered, meaning that the absence
of subtle accessory grooves cannot be reliably confirmed. Furthermore, we observe
that, in the holotype of B. araripensis, the absence of a bifurcated end of the median
groove cannot be confirmed either – the anterior tip of the dentary (at the level of the
first pair of alveoli) has been almost entirely scrapped off, so that the alveoli
previously identified as the first pair actually represent the second one. A pair of
subtle depressions on the dentary anterior surface would be eroded vestiges of the
first pair of alveoli. Furthermore, we observe that in specimen SMNS 55414, referred
to Brasileodactylus araripensis in the literature (due to the presence of accessory
grooves), a bifurcated anterior tip of the median groove is present, though unnoticed
before. Also, in SMNK PAL 1281, referred to Cearadactylus atrox in the literature
(due to the presence of a bifurcated median groove), accessory grooves are present,
though also unnoticed before. Finally, upon reinspection of these specimens, we
noticed that differences in tooth spacing are actually nonexistent (all exhibit, in the
rosette, interalveolar spacing under alveolus diameter). We thus conclude that C.
atrox is indistinguishable from B. araripensis, which has priority. [FAPESP
2019/102316; CNPq 421772/2018-2]
29
RESISTÊNCIA DA COROA DENTÁRIA EM TERÓPODES NÃO-AVIANOS

(DINOSAURIA)
Tainá Constância de FRANÇA

taina.constancia@gmail.com
Rodrigo PÊGAS
rodrigopegas42@gmail.com
Natan Santos BRILHANTE
natan.biologia@gmail.com
Rafael DELCOURT
fabiana.costa@ufabc.edu.br
Terópodes não-avianos consistem em um grupo que apresenta uma grande

diversidade de formas e tamanhos corporais, com distribuição global, e ocupando
todos os continentes durante o Mesozóico. Essa diversidade reflete uma grande
variedade morfológica de coroas dentárias que estaria associada a diferentes nichos
alimentares, tal como acontece com mamíferos viventes. Neste estudo a resistência
da coroa dentária em dinossauros terópodes foi comparada com a de mamíferos
carnívoros viventes e fósseis. A resistência dentária foi calculada a partir da
modelagem mecânica da coroa dentária como uma viga de extremidade fixa e seção
transversal elíptica, conforme protocolo extraído da literatura para o cálculo da
resistência de caninos em mamíferos da ordem Carnivora (Canidae, Felidae e
Hyaenidae). O mesmo modelo foi aplicado em representantes de seis grupos de
dinossauros terópodes (Abelisauridae, Carcharodontosauridae, Ceratosauridae,
Dromaeosauridae, Tyrannosauroidea não-tiranossaurídeos e Neovenatoridae), nos
quais se considerou o maior dente da maxila como equivalente funcional dos caninos
dos mamíferos carnívoros (i.e., ambos aplicariam a maior parte da força durante a
perfuração da presa). Foram utilizadas no modelo medidas de comprimento da coroa
dentária, diâmetro labiolingual (y) e diâmetro mesiodistal (x). Os valores de
resistência obtidos para os caninos de mamíferos foram plotados em dois gráficos (de
regressão linear e de boxplot) na linguagem GNU-R 4.1.2 para a comparação dos
resultados, que indicaram uma resistência no eixo labiolingual (Sy) maior ou igual à
resistência no eixo mesiodistal (Sx) para todos os grupos. A maior resistência
dentária foi representada pelo gênero Acrocanthosaurus (Carcharodontosauridae), e
os menores valores foram observados em Bambiraptor (Dromaeosauridae) e Dilong
(Tyrannosauroidea não-tiranossaurídeo). Os intervalos de valores foram similares
para Dromaeosauridae e Neovenatoridae, que ocuparam intervalo similar ao clado
Canidae. Abelisauridae ocupou intervalo de resistência similar a Canidae, Felidae e
Hyaenidae. A resistência dentária demonstrou grande potencial para identificar
guildas alimentares de terópodes não-avianos, em conjunto com outras variáveis
como tamanho corporal, conforme tem sido observado para mamíferos viventes. Por
exemplo, mamíferos com resistência dentária mais alta apresentariam maior
potencial para se alimentar de presas maiores e o mesmo poderia ser aplicado para
os dinossauros terópodes, corroborando ou inferindo novas estratégias alimentares já
propostas previamente. [CAPES 88887.645497/2021-00]
30
SYSTEMATIC REVIEW ON NON-AVIAN DINOSAUR TEETH FROM CRETACEOUS

BRAZIL
Theo Baptista RIBEIRO

theobribeiro1@gmail.com
Paulo Marques Machado BRITO
pbritopaleo@yahoo.com.br
The fossil record of Brazilian dinosaurs is very rich, with findings from all the periods
of the Mesozoic and more than 20 taxa being described from Cretaceous period.
Many studies have been recently made with the aim of cataloging and reviewing the
diversity of Brazilian dinosaurs, with the majority of them focusing only on
paleobiogeographical issues, while few try to typify which kind of body fossils are
found and their representativity on the fossil record. This study aims to account for
and discuss the representativity of non-avian dinosaur teeth (isolated or in situ) and
also review the methods used for the identification of those dental elements.
Databases such as SciElo and Google Scholar, together with available abstract and
reference lists, were searched in order to obtain the literature used in this review, in
which 123 papers were selected. Our results show that 49.8% (n = 934) of all the
dinosaur fossil record (n = 1872) found in Brazilian Cretaceous rocks is composed by
dental elements, isolated or in situ, with the former representing 42.3% of all fossil
elements found thus far in the country. Within this sampling of teeth, the most
abundant groups were Spinosauridae, Deinonychosauria, Abelisauridae, and
indeterminate theropods. Deinonychosauria, Carcharodontosauridae, Spinosauridae
and indeterminate theropods had a larger ratio of teeth findings per fossil element.
Theropod teeth were more abundant in comparison to Sauropod teeth. Among the
twenty lithostratigraphic units analyzed in this study, seven units did not possess
preserved non-avian dinosaur teeth, while the dinosaur record of two of units were
represent solely by isolated teeth. Alcântara Formation showcased the highest
percentage of dinosaur teeth per fossil elements (90.2%), followed by Itapecuru
(51.6%) and Presidente Prudente (50.0%) Formations. Despite representing a large
share of the Brazilian fossil record, the majority of the isolated theropod teeth were
described with outdated and/or dubious methods that may lead to
misidentifications, making the reassessment of these specimens through the latest
methods essential. Other aspects besides taxonomic identification can also be further
explored using the latest technologies on dinosaur teeth, allowing a deeper
understanding of the paleobiology of these animals. [CAPES; FAPERJ]
31
A CAVIDADE CEREBRAL DE Clevosaurus brasiliensis (LEPIDOSAURIA,

RHYNCHOCEPHALIA) DO TRIÁSSICO SUPERIOR DO SUL DO BRASIL
Lívia Roese MIRON

livia.roem@hotmail.com
Annie S. HSIOU
anniehsiou@ffclrp.usp.br
Apesar de atualmente estarem representados por apenas uma espécie da Nova

Zelândia, durante a primeira metade do Mesozoico o clado de répteis lepidossauros
Rhynchocephalia era consideravelmente diverso, com cerca de cinquenta espécies
conhecidas de morfologia e ecologia variada. O gênero de rincocefálios Clevosaurus
era particularmente diversificado no início do Mesozoico, apresentando distribuição
cosmopolita. Duas espécies são conhecidas nos sedimentos do Triássico Superior do
Rio Grande do Sul (Formação Caturrita, Noriano), sendo a mais abundante
Clevosaurus brasiliensis. Apesar da grande diversidade fóssil do gênero no globo,
pouco se sabe sobre a anatomia de suas cavidades endocranianas. Moldes
endocranianos obtidos a partir de Tomografia Computadorizada (CT) podem prover
informações neuroanatômicas e biológicas sobre as espécies estudadas, sendo
particularmente informativos em espécies extintas. Assim, este estudo busca
descrever pela primeira vez a morfologia da cavidade cerebral de C. brasiliensis. Com
o uso de CT e segmentação manual com auxílio de uma mesa digitalizadora, um
molde digital foi gerado para um espécime adulto da espécie (MCN PV 2852), e uma
série de medidas foi feita para complementar a descrição qualitativa. O volume,
comprimento e largura do molde foram 111.89 mm³, 18.5 mm e 5.98 mm,
respectivamente. O modelo é alongado e praticamente linear. Os hemisférios
cerebrais, lobos ópticos e cerebelo não são distinguíveis, assim como a glândula
pituitária. O trato olfatório é estreito, e os bulbos expandem-se anteriormente. A
superfície dorsal do prosencéfalo possui formato losangular, e as faces laterais da
medula oblongata são aproximadamente paralelas. Apresenta uma organização geral
similar ao conhecido para Sphenodon punctatus, a única espécie vivente de
Rhynchocephalia, mas a forma mais retilínea e organização mais tubular sugerem
um cérebro composto por características mais plesiomórficas. Esta é a primeira
descrição da cavidade cerebral de um rincocefálio fóssil e uma das primeiras para
lepidossauros extintos, expandindo o conhecimento da história evolutiva da
neuroanatomia do clado. [CNPq 130381/2020-9; FAPESP 2020/06819-5]
32
JUVENILE SPECIMEN OF Notosuchus terrestris FROM BAJO DE LA CARPA

FORMATION (UPPER CRETACEOUS), NEUQUÉN, ARGENTINA
Emmanuel Facundo ROUSSILLON

rousillon1095@gmail.com
Jorge O. CALVO
jocalvomac@gmail.com
We report a Mesoeucrocodylia juvenile skull from Bajo de la Carpa Formation,

Neuquén Group (Upper Cretaceous). It was found in the northeast of Neuquén City
(38°56’28” S; 68°03’07” W), Argentina. The material, MUCPv-118, has a length of 4.6
cm; 2.6 cm width and 2.5 cm high. Just the posterior fragment of the premaxila is
preserved showing at the suture premaxila/maxilla the insicivum foramen. On the
left side a caniniform tooth is partially preserved, while on the right side a replace
tooth is seen. The maxilla, arranged vertically, show neurovascular foramens. the
subtriangular anteorbital fenestra shows rounded edges without roughness. The
orbit has 1.7 cm in diameter. The right squamosal is preserved partially and present
a smooth surface without striations; it is laterally projected and contact with the
quadrate covering the otic region. Parietal present two longitudinal redounded crest
separates by a slightly width depression. The supratemporal fenestra is elongated
anteroposteriorly occupying a big portion of the cranial roof. The jugal preserves the
right dorsal ramus contacting the postorbital. Its elongated anterior process contacts
with the maxilla, and the posterior process is incomplete and contact with the
quadratojugal. The quadratojugal, dorsoanteriorly directed, contacts with the jugal
and the postorbital. quadratojugal and Quadrate contact parallelly. The narrow
frontal presents, in the middle, a low crest directed anteroposteriorly. The nasals are
elongated and have a medial longitudinal inflection, also present in other Notosuchus
specimens. Dentary has a slightly sigmoidal shape. The bottom is worn showing the
roots of preserved teeth. The symphysis is projected anterodorsally at an acute angle,
very similar to others adult exemplars like MUCPv-35 and 147. The Meckelian groove
is visible between the dentaries and splenial. Posterior processes of both pterygoids
are incipiently observed. Surangular is concave and posteroventrally projected. The
angular is poorly observed in right lateral. Based on the general morphology, the
specimen MUCPv-118 correspond to Notosuchus terrestris. [Proyecto UNCo 04/I231;
Empresa Chevron SA and Fundación Luciérnaga]
33
WHAT DID THEY EAT? USING STABLE ISOTOPES TO STUDY THE DIET OF A
TRIASSIC VERTEBRATE FAUNA FROM BRAZIL
Leonardo CORECCO
leocorecco@gmail.com
Justin CREVIER
justincrevier@u.boisestate.edu
Linda M. REYNARD
lindareynard@boisestate.edu
Matthew J. KOHN
mattkohn@boisestate.edu
Cesar SCHULTZ
cesar.schultz@ufrgs.br
Bones and especially tooth enamel incorporate chemical elements from their food
and water sources that can provide paleoecological and paleoenvironmental
information. Some of this information can be recovered by using stable isotopes
(δ13C). Here we present isotopic analyses of three taxa from the Brazilian Middle
Triassic Dinodontosaurus AZ fauna (Pinheiros-Chiniquá Sequence, base of Santa
Maria Supersequence) in the context of previous food sources that have been
attributed to them according to their morphological characteristics. Seven samples of
enamel were analyzed for δ13C (13C/12C). Taxa include Prestosuchus chiniquensis
(three samples; hypercarnivorous); Dinodontosaurus sp. (three samples; herbivorous),
and Decuriasuchus quartacolonia (one sample; carnivorous, but a juvenile individual).
The analyzed fossils belong to the Paleovertebrate Section of the Geosciences
Institute of the Universidade Federal do Rio Grande do Sul collection (UFRGS-PV-T).
Our preliminary data show a Prestosuchus chiniquensis δ13C = -8.8±0.6‰
indistinguishable from the Dinodontosaurus sp. δ13C = -8.5±0.4‰. This similarity
could indicate that Prestosuchus could have preyed on Dinodontosaurus or other
herbivore. Decuriasuchus quartacolonia shows a δ13C = -7.6‰. This lower value
compared to P. chiniquensis suggests a distinct diet pattern of D. quartacolonia,
related to ontogeny (as all known specimens of D. quartacolonia are considered
juveniles) or to feeding on different prey. Preliminary MEV analyses on P. chiniquensis
(D1) and Dinodontosaurus sp. (D2) samples shows that D1 have secondary REE
phosphate, mainly Ce (probably florencite), a common secondary mineral that occurs
in supergenic geochemical processes (e.g. soils, lateritic profiles, sedimentary rocks).
Barite veins succeeded by calcite veins were also observed across this sample. D2 is
predominantly composed by fluorapatite, but also presents secondary phosphates
with Fe (ferric florencite?). In general, D2 is less changed than D1, but it is important
to keep in mind that many fossils, including some of the same taxa, that were
recovered at the same locality that our samples were collected, have also some
signals of diagenetic affects (e.g. carbonatic concretions). [CNPQ 141216/2020-4]
34
TEETH OF Priohybodus arambourgi (HYBODONTIFORM) FROM THE TUCANO

BASIN (ALIANÇA FORMATION - JURASSIC): MORPHOLOGY, MORPHOMETRY
AND HISTOLOGY
Estevan ELTINK
estevan.eltink@univasf.edu.br
Kelly Roberta da SILVA
kellyroberta755@gmail.com
Paula Eduarda Gama de Carvalho ROCHA
pauladudarocha@gmail.com
Daniel Vieira De SOUSA
daniel.vsousa@univasf.edu.br
Marco Aurélio Gallo de FRANÇA
marco.franca@univasf.edu.br
Recent discovery of the hybodontiform shark, Priohybodus arambourgi, in the

southern Tucano Basin (Aliança Formation: Upper Jurassic) represents the first
record of the species in Brazil. P. arambourgi had a widespread geographic
distribution in Jurassic and Cretaceous of the Gondwana and the new record
increases the occurrence of the taxon. The morphology of P. arambourgi teeth is
characterized by labiolingual flattening, a very symmetrical teeth, broad and
triangular central cusp (smooth on both sides) with the presence of serrated edges,
and a minimum of three pairs of accessory cusps with smaller sizes. Measurements,
such as the height of the central cusp and the total length of the teeth, demonstrate
a linear correlation. The comparison among P. arambourgi specimens from different
localities reveals a significant variation in the size proportion of the teeth. Teeth from
the Chicla Formation (Libya) are the largest among the analyzed specimens, followed
by the intermediate size teeth from the Aliança Formation (Brazil), and the smallest
ones from Tacuarembó Formation (Uruguay). Geographic isolation or changes in the
diet of this species may have influenced this discrepancy. The histology provides a
microscopic view of layers as enameloid, orthodentin and osteodentin that could be
recognized along with inner teeth structures, such as denteons. P. arambourgi has an
osteodont tooth type, as seen in some other Hybodontiform sharks, such as
Hybodus, and extant Neoselachii. P. arambourgi bears derived characters as
comparatively thinner deposition of orthodentin, and extensive vascular tubes in the
innermost layers of osteodentin. However, P. arambourgi has a monocrystalline layer
of enameloid, which is commonly shared with primitive orthodont teeth. It
demonstrates the complexity in the histology and morphology of fossil sharks teeth.
[FAPESB 5171/2019; CNPq-UNIVASF 05/2020 - PIBIC 3624; FACEPE APQ-1119-
1.07/21]
35
PALEOHISTOLOGIA DE ESCAMAS DE PEIXES TELEÓSTEOS DO GRUPO

SANTANA (CRETÁCEO INFERIOR), BACIA DO ARARIPE
Ísis da Rocha SOUSA

isisdarochasousa@gmail.com
Yuri Modesto ALVES
alves_modesto@yahoo.com.br
O Grupo Santana da Bacia do Araripe, Nordeste do Brasil, é amplamente estudado

devido aos bem preservados afloramentos fossilíferos que contêm uma notável
diversidade de organismos. Fósseis de peixes teleósteos também são abundantes e
diversificados nessa unidade, apresentando, muitas vezes, escamas muito bem
conservadas. Entretanto, apesar dos inúmeros registros paleontológicos, poucos são
os estudos paleohistológicos acerca de suas escamas. Sob essa perspectiva, o
presente trabalho realizou a descrição histológica das escamas de quatro espécies de
teleósteos, †Notelops brama, †Rhacolepis buccalis, †Cladocyclus gardneri e †Vinctifer
comptoni. Os fósseis de estudo foram coletados no Grupo Santana, no distrito de
Santana do Cariri, Ceará, preservados em bloco de concreção calcária de cor
acinzentada. Atualmente eles estão armazenados na Coleção de Paleontologia do
Laboratório de Paleobiologia da Universidade Federal do Tocantins, tombados sob os
acrônimos UFT0481, UFT0484, UFT0485, UFT0492 e UFT0500. Para isso, foram
confeccionadas lâminas petrográficas das escamas triadas dos espécimes para
visualização em microscópio óptico. As análises indicam que as escamas estudadas
possuem dois padrões básicos: ganoide lepidosteoides, o caso de †V. comptoni, e
elasmoide cicloide, caso das outras espécies. Em †V. comptoni, a dentina é ausente e
a placa basal totalmente vascularizada com canalículos de Williamson, semelhante
ao padrão encontrado em Lepisosteidae, todos os †Pachycormidae exceto
†Pachycormus macropterus e Teleostei basais como †Pholidophoridae. Em †N. brama
e †R. buccalis, a ganoína é perdida e são visualizadas uma camada óssea externa que
cobre outra camada óssea. †N. brama e †C. gardneri apresentam uma camada
hipermineralizada de cristas que indicam a ornamentação das escamas, sobreposta à
camada óssea externa, como em Notopterus notopterus, Hiodon alosoides, Megalops
atlanticus, Albula vulpes, Clupea harengus, Chanos chanos e Cyprinus carpio.
Camadas de osso lamelar que atravessam a placa basal ou se localizam na parte
inferior da escama estão presentes em †C. gardneri. Para melhor descrição possível
das escamas e complementação dos resultados, estudos comparativos posteriores
serão realizados. Os resultados poderão fornecer informações importantes acerca das
quatro espécies e uma melhor compreensão dos aspectos sistemáticos dos grupos
basais de Teleostei.
36
A NEW PLIOSAUR FROM THE PAJA FORMATION OF COLOMBIA SUPPORTS

HIGH TOP PREDATOR DIVERSITY DURING THE EARLY CRETACEOUS
Dirley CORTES
dirley.cortes@mail.mcgill.ca
Hans C.E. LARSSON
hans.ce.larsson@mcgill.ca
Mary Luz Parra RUGE
mLparra@centropaleo.com
The Paja Formation in central Colombia was deposited in a shallow marine sea
during the Early Cretaceous. The formation has already yielded abundant
ichthyosaur, thalattosuchian, plesiosaur, fish, and turtle taxa along with several
ammonites, molluscs, crabs, and a dinosaur. Despite the relatively well studied
taxonomy of the Paja Formation’s top predators, little is known regarding their actual
diversity and ecological roles. A new, exquisitely preserved pliosaur from the Paja
Formation is presented. It consists of a large skull about 1.2 m long with complete
dentition and features many autapomorphies including: a laterally constricted
dentary and upper jaw to near orbits, almost like an odontocete whale; a nearly
tubular snout for almost half the skull length (in contrast to the constricted snout of
Stenorhynchosaurus that is only about ? of total skull length - Monquirasaurus and
Sachicasaurus are not comparable as those skulls are significantly larger and
allometrically robust with no obvious snout constriction; and a hypertrophied
frontal-parietal contact with rugose, striated ornamentation over the interorbital
region, constructing a wide interorbital dorsal surface. A phylogenetic analysis places
the new pliosaur with Brachaucheninae and contributes to the taxonomic and
anatomical diversity of this clade. This new taxon is coeval, at nearly the same
horizon in the Paja Formation, with Stenorhychosaurus, Monquirasaurus, and
Sachicasaurus. The high diversity of the top predators in the Paja Formation is
striking and differs from most other Mesozoic marine ecosystems. The different skull
shapes suggest niche partitioning among these taxa and an equally complex
ecosystem, in spite of the paucity of smaller vertebrate fauna. We present
morphometric comparisons across the teeth of these Paja brachauchenines and
hypothesize their feeding guilds within the known ecosystem. Our work on this new
pliosaur taxon provides useful data on the Paja’s ecosystem complexity and food web
interactions and sheds light on the ecological roles of the last surviving top predators
during the late Mesozoic before their imminent extinction.
37
NEW RECORD OF Brasilosuchus mendesi (CROCODYLIA: TOMISTOMINAE?) IN

CAJUEIRO LOCALITY, BOCA DO ACRE-AM
Yasmin Catarina Alves DANTAS

ydantas001@gmail.com
Lucy SOUZA
souzalucyg@gmail.com
Peter Mann De TOLEDO
peter.toledo@hotmail.com
Thiago Silva LOBODA
loboda_bio@yahoo.com.br
Edson GUILHERME
guilherme.edson@gmail.com
Brasilosuchus is considered to be part of the Tomistominae clade, popular known as

“false gharials”, which the only living species is the Tomistoma schlegelii, from
Malasian. The Brasilosuchus genera lived where today is the north of Brazil, in the
Acre state, and had piscivorous habits, as indicated by its logirostrine and its long,
conical and homodont dentition. This animal lived alongside with others longirostrine
Crocodylia, like the Gavialoidea genera Gryposuchus. During the Profª. Rosalie
Benchimol expedition, a partnership between MUSA and UFAC, in 2019, an
incomplete mandible of a longirostrine Crocodylia has been found in the Cajueiro
locality, in Boca do Acre municipality, Amazonas, Brazil, on the left margin of Purus
River. The fossil MUSA 3.1.129 is composed by the premaxilla and part of the right
maxilla, preserving the premaxilla alveoli, three maxilla alveoli and three teeth. It can
be assigned to the Brasilosuchus genera by the presence of five premaxilla alveoli,
with the second and fifth been the smallest, the anteriorly projection of the
premaxilla, causing the naris to be distant from the tip of the snout, and the notches
between each alveolus of the premaxilla and maxilla. It differs from Gryposuchus by
its dentition (Gryposuchus has only four premaxilla alveoli) and by its incisive
foramen, almost absent, even smaller than the ones in Gryposuchus. This is a new
specimen of B. mendesi, been a little more complete, and its geographic distribution
has been expanded to the Amazonas state.
38
FIRST FOSSIL RECORD OF Melanosuchus cf. Melanosuchus niger (CAIMANIDAE:

CROCODYLIA) ON CAJUEIRO SITE, BOCA DO ACRE, AMAZONAS
Lavínia Tavares PAZ

laviniatpaz@gmail.com
Caio Cunha FERREIRA
caiocunhaferreira@gmail.com
Peter Mann De TOLEDO
peter.toledo@hotmail.com
Thiago Silva LOBODA
loboda_bio@yahoo.com.br
Edson GUILHERME
guilherme.edson@gmail.com
Lucy SOUZA
souzalucyg@gmail.com
The Miocene-Pliocene Epochs of the Amazon are notorious for the high diversity of
crocodylian fauna, in which the Cajueiro site represents one of the notable and
richest fossil deposits. The present work identifies a new Caimaninae fossil record for
this region (specimen MUSA 3.1.130, a preserved partial snout from a small adult
consisting of the right half of the rostrum, including the maxillary bone and a
fragment of the premaxillary bone), assigned as Melanosuchus cf. niger according to
the following diagnostic characters: (1) the presence of 13 maxillary teeth and (2) a
preserved and well marked rostral ridge in dorsal view; In addition, the relative width
of the maxilla in relation to its length suggests a broader and more dorsoventrally
flattened snout compared to Caiman crocodilus, Caiman yacare, Paleosuchus
palpebrosus and Paleosuchus trigonatus, but also an anteroposteriorly longer snout
when compared to Caiman latirostris. In comparison to the representatives of the
genus Melanosuchus, M. niger has thirteen maxillary teeth, while M. latrubessei
presents 12. The maxilla-maxilla contact in ventral view of specimen MUSA 3.1.130
extends until not after the 4th alveoli, as M. niger, which has a maxilla-maxilla
contact´s length between the 2nd and 4th maxillary teeth, while M. latrubessei
extends from third to the sixth maxillary alveoli. The fossil also presents four
preserved heterodont teeth. The premaxilla-maxilla suture is not preserved in its
entirety, with the fossil lacking Melanosuchus’ characteristic exposed vomer in said
suture. This record contributes to a better understanding of the palaeobiodiversity in
South America, serving not only as a piece for the reconstruction of the crocodilian
fauna of the period but also as valuable data for evolutionary and palaeoecological
studies.
39
ESTUDO COMPARATIVO DE ESPINHOS FÓSSEIS DE CTENACANTIFORMES

(FORMAÇÃO PEDRA DE FOGO, ESTADO DO TOCANTINS)
Jhonathan Guimarães Sousa COSTA

biojhonathansousa@gmail.com
Daniel FORTIER
fortier@ufpi.edu.br
Juan C. CISNEROS
juan.cisneros@ufpi.edu.br
A Formação Pedra de Fogo localizada na Bacia do Parnaíba possui uma rica

diversidade de peixes fósseis referentes ao Período Permiano, e alguns destes
correspondem ao táxon Ctenacanthus, um gênero de tubarão fóssil representado em
seu registro na maioria das vezes por espinhos de nadadeiras dorsais. O gênero foi
descrito em 1837 utilizando um holótipo através de características como
ornamentações e vista transversal e designado como espécie tipo Ctenacanthus
major. O presente trabalho tem como objetivo realizar a descrição de espinhos fósseis
de tubarão que se encontram armazenados na coleção científica do Museu de
Arqueologia e Paleontologia da Universidade Federal do Piauí. Levando em
consideração caracteres utilizados na descrição de outros tubarões fósseis do
Permiano em bibliografia especializada, foram analisados 11 espinhos fósseis
medindo de 2cm à 7cm de comprimento, possuindo costelas lisas e/ou com
ornamentação pectinada, arredondada, achatada e justaposta, presentes tanto na
região anterior e nas faces laterais, como em apenas uma dessas regiões, sendo
também mais abundante em alguns fragmentos do que em outros, as costelas finas e
pouco espaçadas se repetem sequencialmente em sentido longitudinal, com
pequenos tubérculos que se alternam em suas inserções nas costelas, originando um
espaçamento entre cada tubérculo, dispostos intercaladamente, como “dentes de um
zíper”. Os espécimes analisados se assemelham a espécie Ctenacanthus
amblyxiphias, semelhança essa já sugerida por autores brasileiros para outros
registros de espinhos fósseis encontrados no país. No entanto, desde a primeira
descrição do gênero, foram adicionadas a estas várias espécies referentes a espinhos
isolados, tornando a sistemática do grupo de difícil compreensão. Somando isso ao
fato de ainda serem debatidas as características que de fato definem o gênero
Ctenacanthus. No presente trabalho o material está sendo considerado como
Ctenacanthiformes. Espera-se que tanto este trabalho como futuros agreguem
informações para uma classificação mais específica não só do material aqui
analisado mas também para o gênero. [FAPEPI]
40
- Simpósio Evolução dos Roedores -
EL REGISTRO FÓSIL DE LOS PAN-CHINCHILLIDAE (RODENTIA,

CAVIOMORPHA): SU IMPORTANCIA BIOESTRATIGRÁFICA Y PALEOAMBIENTAL
Luciano Luis RASIA
lucianorasia@hotmail.com
Pan-Chinchillidae es un grupo de roedores caviomorfos de América de Sur con un

registro fósil muy rico y cronológicamente extenso pero con pocos representantes en
la actualidad, incluyendo a la vizcacha (Lagostomus maximus), las chinchillas
(Chinchilla spp.) y los chinchillones o vizcachas de montaña (Lagidium spp.). Estudios
recientes incluyen como stem Chinchillidae a †Eoviscaccia (Oligoceno temprano-
Mioceno temprano de Argentina, Chile y Bolivia), †Incamys (Oligoceno tardío de
Argentina y Bolivia), †Loncolicu (Oligoceno tardío de Argentina), †Garridomys y
†Saremmys (ambos del Mioceno temprano de Argentina); mientras que las
subfamilias con representantes vivientes (Lagostominae y Chinchillinae) forman el
crown group de Chinchillidae. Los lagostominos son conocidos desde el Mioceno
temprano y medio con los géneros †Prolagostomus en Argentina, Chile y Bolivia, y
†Pliolagostomus en Argentina. Desde el Mioceno tardío se registran varias especies
del género viviente Lagostomus en Argentina, Uruguay, Bolivia y Brasil. La historia
evolutiva de los chinchillinos fue prácticamente desconocida hasta la descripción de
†Miochinchilla del Mioceno temprano y medio de Chile y Bolivia, cercanamente
relacionado con Chinchilla y Lagidium. Si bien en tiempos recientes se le ha dedicado
mayor atención al estudio de este grupo, la descripción de nuevos géneros y especies
sugiere que el conocimiento que tenemos de los pan-chinchíllidos es incipiente en
comparación con otros caviomorfos. El objetivo de esta contribución es revisar el
registro fósil de los pan-chinchíllidos, resaltando su importancia bioestratigráfica y
paleoambiental. La distribución estratigráfica de los géneros de pan-chinchíllidos
muestra que pueden ser útiles como herramientas bioestratigráficas a nivel regional,
y los estudios de la distribución estratigráfica de las especies de Lagostomus han
probado su utilidad bioestratigráfica también a escala regional y local, pero se
necesitan más estudios del resto de los pan-chinchíllidos a nivel específico. Estudios
evolutivos del grupo han mostrado que ciertos factores como el grado de hipsodoncia,
la simplificación dentaria o el tamaño corporal, podrían estar cercanamente
relacionados con cambios climático-ambientales, tales como: procesos geológicos,
oscilaciones en la temperatura global y cambios en la flora.
41
LATE MIOCENE DINOMYIDAE, NEOEPIBLEMAE, AND HYDROCHOERINAE

CAVIOMORPHS (RODENTIA: HYSTRICOGNATHI) FROM ENVIRA RIVER,
NORTHERN BRAZIL
Emmanuelle Fontoura MACHADO

emmanuelle.fontoura@acad.ufsm.br
Ana Maria RIBEIRO
anamariaribeiro69@gmail.com
Francisco Ricardo NEGRI
frnegriufac@gmail.com
Leonardo KERBER
Caviomorpha is a taxonomic and phenotypic diverse clade of South American

hystricognath rodents. Its stratigraphic distribution ranges from the middle Eocene
to recent times. In the Solimões Formation, located in northwest Brazil, a rich fossil
record of the Neogene has been documented in recent years, including marsupials,
rodents, xenarthrans, primates, birds, snakes, invertebrates, plants, among others.
Here, we report preliminary data on a new fossiliferous assemblage of rodents from
the Solimões Formation recovered in fieldwork on the Envira River (Acre). The
specimens were collected in 2010 and 2019 in sedimentary strata of the Neogene
exposed on the banks of this river and are deposited in the paleontological collection
of the Federal University of Acre, Campus Floresta, Cruzeiro do Sul (UFAC-CS). The
fossils were found using the screen-washing method with an aqueous solution of
hydrogen peroxide and subsequent manual screening under a stereomicroscope. In
this work, we report isolated teeth and a portion of the maxilla assigned to
Dinomyidae (Potamarchus sp.), Neoepiblemidae (Neoepiblema sp.), and
Hydrochoerinae (Cardiatherium sp.) rodents. The studied locality has not yet been
dated using radiometric methods. However, the biochronology of the taxa studied
here constrains the age of the fossiliferous levels to the Upper Miocene. Other
fossiliferous outcrops of Solimões Formation dated by radiometric methods have a
Tortonian age. The study of rodents found in outcrops of the Envira River fills a gap
in the knowledge of the distribution of fossils from this stratigraphic unit. [CNPq
140639/2021-7]
42
ADVANCES IN THE STUDY OF NEOEPIBLEMIDAE RODENTS (HYSTRICOGNATHI:

CAVIOMORPHA) FROM THE LATE MIOCENE OF BRAZIL
Leonardo KERBER
Neoepiblemidae is a clade of Chinchilloidea caviomorphs (pacaranas, chinchillas,

vizcachas, and extinct forms) characterized by the large dimensions they reached
during the late Miocene. The clade comprises Perimys, Doryperimys (late
Oligocene/early Miocene), Neoepiblema, and Phoberomys (late Miocene). These
rodents are an expressive component of the paleofauna of the Solimões Formation
(upper Miocene, northern Brazil). The quality of the fossil record of this sedimentary
formation, which includes dozens of isolated teeth, postcranial bones, and the best-
preserved known cranial materials of this group, permitted new interpretations of the
lifestyle of these animals. In recent years, there have been advances in the study of
the Brazilian fossil record, which include: 1) taxonomic and anatomical revision of
specimens; 2) phylogenetic reassessment of neoepiblemids in the context of extant
and extinct chinchilloids; 3) paleoecological interpretations of the fossil record; 4)
analysis of cheek teeth replacement in Neoepiblema; 5) endocranial studies and
paleoneurological approaches; 6) body mass estimates based on allometric equations
and volumetric models; 7) functional morphology analyses of the postcranial bones.
These studies have promoted a greater understanding of the diversity, phylogenetic
relationships, and paleobiology of the giant rodents (Neoepiblema and Phoberomys)
that inhabited the Brazilian Amazon 10 million years ago.
43
GIGANTISM EVOLUTION IN CAVIOMORPHS: ASSESSING THE CRANIAL

ENDOCAST ANATOMY AND BRAIN SIZE OF Josephoartigasia monesi (RODENTIA:
DINOMYIDAE)
Jose Darival FERREIRA

darival.ferreira@gmail.com
A. RINDERKNECHT
rinderk@adinet.com.uy
M.T. DOZO
dozo@cenpat-conicet.gob.ar
M.R. SÁNCHEZ-VILLAGRA
info@freunde-zsm.de
Leonardo KERBER
The high ecomorphological diversity of caviomorph rodents in South America

includes extinct forms that have reached great dimensions. The largest known rodent
was the Dinomyidae Josephoartigasia monesi from the Pliocene of Uruguay. This
work aims to study the neuroanatomy and encephalization of J. monesi (MNHN 921,
San José Formation, Uruguay) using virtual endocranial models. For this study 69
specimens of most caviomorph families were also scanned (CT-Scan). To estimate the
encephalization quotient (EQ) of each specimen, we employed the endocranial volume
converted to brain mass and estimated the body mass based on the average of linear
regressions from eight cranial linear measurements. The EQ was estimated based on
an equation designed specifically for rodents. Our preliminary results demonstrate
that the cranial endocast of J. monesi shows: a long olfactory tract, gyrencephaly,
superior sagittal sinus forming a longitudinal ridge on the endocranial surface,
marked rhinal fissure, and absence of evident paraflocculi. The estimated body mass
is 845 kg, and the volume of the endocast is 192,144 mm³. The estimated EQ of J.
monesi is 0.31. This is a low encephalization coefficient when compared to extant
caviomorphs, which we analyzed here (Erethizontidae: 1.04-1.77, Caviidae: 0.75-
1.18, Cuniculidae: 1.04-1.44, Dasyproctidae: 1.13-1.33, Chinchillidae: 0.85-1.49,
Dinomyidae: 0.97, Echimyidae: 0.76-1.13, Abrocomyidae: 0.94, Octodontidae: 0.88-
0.99), a pattern also found recently in neoepiblemids (0.36), another lineage of giant
rodents. It has been hypothesized that the adaptive value of a low energetic cost and
other ecological factors could explain the presence of relatively small brains in giant
Neogene rodents.
44
REDESCRIPTION OF Neoreomys huilensis FIELD (RODENTIA, HYSTRICOGNATHI)

FROM THE MIDDLE MIOCENE OF LA VENTA (VILLAVIEJA, HUILA, COLOMBIA)
Francisco Javier Urrea BARRETO

fjurreab@gmail.com
Andrés Link OSPINA
a.link@uniandes.edu.co
Juan CARRILLO
juanchorrillo@gmail.com
Andrés VANEGAS
andresvanegasv@gmail.com
Pati PÉREZ
mperez@mef.org.ar
Neoreomys is among the most representative genus of South American Miocene

rodents. The systematic of this taxon have been uncertain and some studies have
suggested their affinity to Myocastorinae, Capromyidae, or Dasyproctidae. Currently,
the most accepted hypothesis based on taxonomic and phylogenetic analysis
proposed Neoreomys is within Cavioidea. Neoreomys is extremely abundant at high
latitudes, and well represented by skulls, teeth, and entire skeletons. Three Miocene
species are currently recognized: N. australis and N. pinturensis both restricted to the
early to middle Miocene of Patagonia, and N. huilensis from the middle Miocene of La
Venta, Colombia in tropical latitudes. N. huilensis has been known from a few
fragmentary specimens, and Fields (1957) detailed description of its holotype. Recent
expeditions have led to the discovery of new material (mandibular and lower teeth) of
N. huilensis, adding new information about this taxon. The aim of this work is to the
reanalyze the systematics, phylogenetic relations and biogeography of Neoreomys
huilensis based on old and new collected materials. N. huilensis is the smaller species
of the genus, lower-crowned than the type species. The molariforms are
tetralophodont with transverse, long, and wide crests. The metalophulid II is well
developed, similar to metalophulid I. The fossetids are more persistent than in other
species of Neoreomys. Some individuals have a flexid on the metalophulid I that is
eventually closed with wear. The dp4 is mesiodistally long, and lower-crowned than
the other teeth, and its general morphology is similar to the dp4 of some specimens
of Dasyprocta. Our phylogenetic analysis supports the monophyly of the genus and
reveals that Neoreomys is not a dasyproctid, but instead corresponds to a basal form
within the clade known as Cavioidea s.s. which includes the crown group of Caviidae.
Although a broader analysis that includes other caviomorphs representatives is still
required, our results suggests that the origin of the genus is Patagonian with a later
distribution towards lower latitudes. The disappearance of this species of Patagonia
and the persistence of the lineage in the tropical latitudes could be related to the
climatic deterioration that occurred after the middle Miocene warming event.
45
ANÁLISIS PRELIMINAR DE LA DIVERSIDAD DE CAVIDOS DEL MIOCENO-

PLEISTOCENO EN ARGENTINA
Maria Carolina Madozzo JAEN

cmadozzo@csnat.unt.edu.ar
Pati PÉREZ
mperez@mef.org.ar
En la actualidad los roedores caviomorfos están representados por cuatro

superfamilias: Cavioidea, Chinchilloidea, Erethizontoidea y Octodontoidea. Los
cavioideos concentran su riqueza en los Andes centrales y la Caatinga en Brasil. Los
registros más antiguos de la Caviidae provienen del Mioceno medio y a partir del
Mioceno tardío se diversifican en las tres subfamilias que llegan hasta la actualidad
(maras, cuises y carpinchos). Esta contribución tiene por objetivo realizar un análisis
preliminar de la diversidad y abundancia de cávidos en el registro fósil del Mioceno
tardío-Pleistoceno de Argentina. Se confeccionó una base de datos (51 entidades
taxonómicas y 500 restos fósiles) con registros publicados, como de estudios en
curso de sedimentos del norte (Formaciones Andalhuala, Chiquimil), noreste
(Conglomerado osífero), Cuyo (Fmaciones Huayquerías y Tunuyán), en la región
pampeana (Fms. Caleufú, Arroyo Chasicó, Monte Hermoso, Chapadmalal) y en la
Patagonia (Fm. Puerto Madryn) siendo la Fm. Chapadmalal la más productiva en
términos de abundancia. Los resultados es este estudio muestran que el registro de
Caviidae está dominado en un 20% por Dolicavia minuscula, Plioceno de la formación
Chapalmalal y “Orthomyctera” andina con más del 10%, del Mioceno tardío-Plioceno
del NOA y Cuyo y Palaeocavia de la Formación Cerro Azul, de La Pampa (Mioceno
tardío-Plioceno) con el 7%. A partir del Plio-Pleistoceno se produce un marcado
recambio de especies quedando representados solo géneros que llegan hasta la
actualidad, así mismo el número de especies y la abundancia de las mismas decrece
notoriamente. A nivel biogeográfico, la región del NOA comparte con Argentina
Central (Plioceno temprano-tardío de la localidad de Chapadmalal, Provincia de
Buenos Aires) dos taxones (Caviodon cf. C. pozzi, y D. minuscula) del Plioceno
temprano-Mioceno tardío del Formación Andalhuala, y con las Formaciones
Huayquerías y Tunuyán de Mendoza tres (“Orthomyctera” andina, Palaeocavia
mawka y Prodolichotis prisca), esta última región comparte solo un taxón con la
Formación Cerro Azul de La Pampa (Neocavia pampeana). Para realizar estudios más
precisos de abundancia y diversidad son necesarios nuevos muestreos en sedimentos
del Cenozoico tardío, tanto en Argentina, como en otras localidades de América del
Sur que incrementen el número de ocurrencia de cada especie.
46
THE AUDITORY REGION OF Neoreomys australis AMEGHINO, 1887

(HYSTRICOGNATHI, CAVIOIDEA) FROM THE EARLY MIOCENE OF PATAGONIA
Maria Eugenia ARANUDO

mearnaudo@gmail.com
Pati PÉREZ
mperez@mef.org.ar
Neoreomys australis Ameghino, is one of the most representative caviomorph rodents

of the early Miocene (Santacrucian age) in regards to the abundance and quality of
the preservation. Neoreomys has been traditionally considered as an ancient lineage
related to the living Dasyprocta and Myoprocta. However, recent morphological and
phylogenetic analysis corroborated its inclusion as part of the basal forms of
Cavioidea sensu stricto. Although N. australis is remarkably abundant in the fossil
record, its anatomical and paleobiological characteristics have been scarcely
analyzed, except very rarely. The purpose of this contribution is the exploration of the
internal anatomy of the auditory region of Neoreomys australis (FMNH P 13155) in
the Cavoidea context. The most notorious features are the number of turns of the
cochlea, which reached almost five and makes Neoreomys one of the caviomorphs
with the highest amount of cochlea turns, and much higher than the average for
mammals in general. The anterior semicircular canal is the biggest, followed by the
lateral (LSC) and finally by the posterior, which is the smallest. A largest LSC than
the posterior implies more agility movements. The middle ear cavity is globose in
Neoreomys but septae are absent. The epitympanic recess is also well developed in
comparison with other cavioids (e.g., Dasyprocta, Dolichotis, Cuniculus). Middle ear
ossicles are similar in shape to those described for other caviomorphs taxa (e.g.,
freely movile and with a malleoincudal complex). A longer cochlea could indicate
wider frequency hearing range, with a major capacity to detect low frecuency sounds
than those mammals with less coiled cochleae. Also, a well developed middle ear
cavity is chracteristic of low frecuency hearing adaptations. From this preliminary
analysis of the auditory region of N. australis, it is expected to infer its paleobiological
and evolutionary significance within Cavioidea. [ANPCyT PICT 2010-2613 (MEP);
ANPCyT PICT 2016-0566 (MEP)].
47
DESCRIPTION OF THE SKULL OF Acarechimys (CAVIOMORPHA, PAN-

OCTODONTOIDEA) AND EARLY EVOLUTIONARY CONSIDERATIONS
Michelle ARNAL
michoarnal@gmail.com.ar
Pati PÉREZ
mperez@mef.org.ar
Maria Eugenia ARANUDO
mearnaudo@gmail.com
Maria Susana BARGO
msbargo@fcnym.unlp.edu.ar
Sergio F. VIZCAINO
vizcaino@fcnym.unlp.edu.ar
Pan-octodontoidea is the earliest caviomorph lineage to differentiate and the most

taxonomically diverse group with the greatest morphological disparity. Acarechimys
is the pan-octodontoid rodent with the longest biochron (late Oligocene-middle
Miocene) and the widest geographic distribution (Argentinean Patagonia to La Venta,
Colombia). The genus comprises five species: A. leucotheae from the late Oligocene of
Chubut Province (Patagonia); A. gracilis and A. constans from the early Miocene of
Chubut and Santa Cruz provinces (Patagonia); and the most common species, A.
minutus and A. minutissimus, from the early middle Miocene of Patagonia, Bolivia,
and Colombia. Despite its abundant fossil record (composed mainly of mandibles and
maxillary fragments with teeth), no complete skull has been reported to date. Based
on its generalized low-crowned and tetralophodont teeth, Acarechimys has been
considered to be a generalized caviomorph rodent. In this work, we described an
almost complete skull, only lacking both zygomatic arches, recovered from outcrops
of the Santa Cruz Formation (early-middle Miocene) in the locality of Río Chalía,
Santa Cruz Province (Patagonia). Based on the cheek teeth morphology and size, this
specimen is assigned to Acarechimys minutus. Its most striking aspect is the delicate
anterior half of the skull, with a narrow snout, incisors, and orbital region, that
contrasts with a much wider and robust basicranium with large auditory bullae. This
general morphology differs from that of other fossil pan-octodontoids. However, the
presence of low-crowned cheek teeth and large auditory bullae, with a well-developed
epitympanic sinus and mastoid region, is a striking character also present in the
early Miocene Prospaniomys priscus from Patagonia. Acarechimys minutus supports
the hypothesis that a basicranium with large auditory bullae could represent a
generalized morphology and that adaptations to enhance low-frequency hearing, a
primitive condition for, at least, pan-octodontoid caviomorphs. [ANPCyT PICT 2016-
0566 (MEP, MA); ANPCyT PICT 2017-1081 (MSB); UNLP N867 (SFV)].
48
- Simpósio Tafonomia & Paleoecologia -
NEW OCCURRENCE AND TAPHONOMY OF Myrmecophaga tridactyla LINNAEUS,

1758 IN A CAVE IN NORTHEASTERN BRAZIL
Lais Alves SILVA

allveslais@gmail.com
Ronaldo Araujo LEONI
ronaldoaleoni@gmail.com
Fernando Henrique de Souza BARBOSA
fhsbarbosa@gmail.com
Herminio Ismael ARAUJO JR
herminio.ismael@yahoo.com.br
Karst deposits constitute an important basis for Quaternary Paleontology as they are
extremely favorable environments for processes that lead to the fossilization of
organic remains, presenting well-preserved fossils. Here, we present a new fossil
record of Myrmecophaga tridactyla collected at Gruta do Engrunado, Nova Redenção,
Bahia, and its taphonomic aspects. The material is housed at the paleontological
collection of Laboratório de Ecologia e Geociências, Universidade Federal da Bahia,
Vitória da Conquista/BA. The fossils were analyzed macroscopically for anatomical
identification and observation of the ontogenetic stage. In addition, the
biostratinomic signatures were analyzed, including: physical integrity,
disarticulation, weathering, transportability, abrasion, incrustation, corrosion,
coloration and biogenic marks. The specimens were found associated, with some
elements still articulated. The bones collected were: skull, vertebrae (atlas, five
cervicals, 13 thoracics, three lumbars, two sacrals and 23 caudals), 14 vertebral
discs, right and left scapula, right and left humerus and their proximal epiphysis,
right and left radius and ulna, 24 fragments of ribs, two fragments of pelvis, right
and left femurs and their distal and proximal epiphysis, right and left tibias, two
fragments of fibula, right and left calcaneus and astragalus, two metacarpals and 10
phalanges. The morphological characteristics and the presence of articulated
materials allow to assigne the material to a single subadult individual. Of this
sample, 70.63% of the bones are complete, while 23.81% are partial. As it was an
almost complete individual, bones from the three Voorhies Groups were recorded.
About 79.37% showed incrustation and 80.16% of corrosion. Abrasion, weathering
and biogenic marks were not observed. All bones are blackish. The absence of signs
of abrasion, the presence of bones in all Voorhies Groups and the presence of
complete and articulated skeletal elements suggest an in situ deposition.
Incrustation and corrosion indicate wet conditions during fossilization, as the former
occurs by precipitation of CaCO3 in the surrounding waters, while the latter occurs
under acidic water conditions. The dark color may be related to the incorporation of
MnO2 in the bones, indicating that these remains were probably immersed in water
or wet sediment, which corroborates the incrustation and corrosion data. [CAPES;
CNPq; FAPERJ]
49
BIOLOGICAL AND TAPHONOMIC BIAS OF LISSAMPHIBIAN RECORDS IN SOUTH

AMERICA
Lucas BARCELOS
lucasabarcelos@gmail.com
Rodolfo SANTOS
rodolfosantos013@gmail.com
Lissamphibians are typically small-sized specimens with fragile bones that breathe
through permeable skin, and present physiological and reproductive restrictions
related to moisture, suggesting a marked sensitivity to environmental conditions.
Assuming that their fossil relatives had similar restrictions, these biological factors
could affect the occurrence of lissamphibian remains in fossiliferous deposits.
Beyond that, taphonomic biases (e.g. decay, erosion, weathering, action of scavengers
and microbial mats) influence the formation of thanatocoenoses and taphocoenoses
of lissamphibians fossils. Our review on fossil record of South America
lissamphibians suggests records are biased in the type of remains preserved and in
relation to the depositional environments represented in their bearing rocks. Fossils
of Gymnophiona and Caudata are mainly found in lacustrine depositional
environments, preserved as vertebral remains. We recognize that their fossil record is
biased, favoring taxa that live next to or in aquatic environments. In contrast, most
fossorial caecilians rarely live where deposition of sediments occurs and, thus,
fossilization of their remains is rare. Record of Caudata in South America comprises
only a few isolated vertebrae, so it is difficult to propose general patterns from it.
Anura records are mostly concentrated in rocks representing fluvial settings, followed
by bearing rocks characterizing lacustrine environments. That is congruent with the
physiological and ecological features of anurans, which use those kinds of
environments as mating and oviposition sites. Skeleton portion predominantly
preserved and main depositional environments of the records are: Ceratophryidae
(skull; fluvial), Pipimorpha (skeleton; lacustrine), Calyptocephalellidae (postcranial;
fluvial), and Bufonidae (postcranial; fluvial). We infer that this pattern found in
anuran fossils are justified by the microhabitat occupied and anatomical features of
those families: the records are mainly from aquatic (Pipimorpha, Ceratophryidae
Lepidobatrachus, and Calyptocephalellidae) or hyperossified taxa (Ceratophryidae,
Calyptocephalellidae, Bufonidae, and Pipimorpha). These findings corroborate the
idea that skeletal remains of aquatic and hyperossified taxa could further resist the
destruction process and fulfil the biostratinomy and fossildiagenesis process. We
recognize that lacustrine environments present most features that enable the
complete preservation of a skeleton, as exemplified by pipid’s record. [CAPES;
FAPESP n° 2019/24466-5]
50
ESTUDO TAFONÔMICO DE VERTEBRADOS TERRESTRES FÓSSEIS DA

FORMAÇÃO AÇU (?APTIANO-CENOMANIANO DA BACIA POTIGUAR), NORDESTE
DO BRASIL
Letícia Paiva BELFORT

leticiapaivabelfort@hotmail.com
A Bacia Potiguar é conhecida por seus fósseis marinhos da Formação Jandaíra e, até
meados da década de 1960, a Formação Açu foi considerada afossilífera. Por meio de
pesquisas recentes na unidade operacional 4 da Formação Açu, pesquisadores
coletaram fósseis de vertebrados de grande porte, os quais foram atribuídos a
dinossauros, com associação a peixes e outros crocodiliformes. Este trabalho tem
como objetivo descrever e interpretar assinaturas tafonômicas macroscópicas de
fósseis coletados em alguns pontos específicos da área aflorante. O material
analisado consiste em 147 espécimes, e sua maior parte é composta por vértebras e
ossos longos. Os fósseis estão tombados na Coleção de Paleontologia do
Departamento de Geologia (DEGEO), Instituto de Geociências (IGEO), da
Universidade Federal do Rio de Janeiro (UFRJ). Foram utilizados três pontos de
coleta denominados de Ponto 1.1, Ponto 1.2 e Ponto 2.3. Para isso, foram avaliados:
(A) articulação e fragmentação; (B) representatividade óssea; (C) equivalência
hidráulica; (D) padrões de quebra; (E) marcas de intemperismo; (F) abrasão; (G)
marcas de pisoteio; (H) bioerosão causada por organismos vivos; (I) marcas de raízes;
(J) padrões de coloração. Todos os espécimes apresentam-se desarticulados. Apesar
de grande parte dos fósseis terem sido classificados como indeterminados, a análise
de transportabilidade por FTI indica uma assembleia periférica pelo predomínio do
Grupo I em 53,45%, sendo 50% de vértebras. Em relação à abrasão, 34%
apresentam leve abrasão e 32% de abrasão moderada, sugerindo um moderado
transporte. Há uma prevalência do grau 2 de intemperismo com 40%, sugerindo
moderada exposição subaérea. Marcas de dentes reafirmam essa exposição subaérea
e a presença de bioerosão por insetos propõe um ambiente úmido. Há presença de
fóssil com marcas de raízes, sugerindo que houve presença de vegetação no local do
soterramento. Poucos ossos longos aparecem na assembleia mas, quando presentes,
um padrão de descamação é dominante. Por fim, a análise preliminar indica um
ambiente úmido devido as marcas de inseto, com períodos anormalmente mais secos
derivado das taxas de intemperismo, e os fósseis submetidos a moderado transporte.
[FAPERJ]
51
REGISTRO E ASPECTOS TAFONÔMICOS DE UM PHASIANIDAE

INDETERMINADO (AVES) EM DEPÓSITO CÁRSTICO DA BAHIA, BRASIL
Ronaldo Araujo LEONI

ronaldoaleoni@gmail.com
Lais Alves SILVA
Carlos Micael Bonfim LESSA
biologolessa@gmail.com
André Vieira de ARAUJO
anddrevieira@gmail.com
Registros de aves fósseis em cavernas no Brasil são raros. Dentre as razões estão a
sua morfofisiologia e consequentemente a inadaptabilidade do grupo ao ambiente
cavernícola. Este estudo propõe a identificação taxonômica e análise tafonômica dos
remanescentes de uma ave resgatada na caverna Toca da Boa Vista, Campo
Formoso, Bahia. O espécime (LEG 2288-2312) está depositado na coleção
paleontológica do Laboratório de Ecologia e Geociências da Universidade Federal da
Bahia (IMS/CAT). Os elementos coletados constituem um fragmento de crânio, três
vértebras cervicais, fúrcula, fragmento do esterno, dois fragmentos de úmero,
escápula, fragmento de coracoide, ulna, carpometacarpo, fragmento de sinsacro,
fragmento do fêmur, fragmento de tibiotarso e fragmento de tarsometatarso. O
tamanho e forma da abertura do osso nasal, o processo articular esternal do
coracoide e a morfologia geral de elementos como escápula, tarsometatarso, sinsacro
apontam para um indivíduo da ordem Galliformes, mais especificamente da família
Phasianidae. Os elementos foram encontrados a aproximadamente 2-2.5 km da
entrada da caverna, em um conduto de quatro metros de largura, completamente
desarticulado e incompleto, seus elementos separados por poucos centímetros de
distância entre si, com uma notável ausência de elementos menores, exceto por três
vértebras cervicais. Além disso, a maioria dos elementos foram encontrados
fragmentados, com quebras irregulares, em sua maioria paralelas às fibras ósseas.
Não foram encontrados vestígios de tecidos moles, caracterizando os restos no
estágio 2 de Behrensmeyer, conforme a mesma classificação, os restos ficaram
expostos aos agentes intempéricos de 2 a 5 anos, após a necrólise. Todo o esqueleto
apresenta um aspecto marrom avermelhado, indicando a composição química do
deposito fossilífero. Perfurações ovaladas e irregulares podem ser notadas ao longo
do fêmur, com autoria provável de um pequeno carnívoro. Durante a necrólise, pode
ocorrer um fenômeno de aderência temporária da carcaça ao solo, esse efeito pode
ter contribuído para a permanência do esqueleto no ambiente externo, impedindo
quaisquer tipos de transporte temporariamente. A desarticulação, presença de todos
os grupos de Voorhies e a ausência de abrasão classificam a acumulação como
periférica, e um agente hidráulico de alta energia como causa de transporte, agentes
hidráulicos de baixa energia posteriores não são descartados.
52
RADIOCARBON DATING AND ISOTOPIC PALEOECOLOGY (δ13C, δ18O) OF

Glossotherium sp. FROM MIDWEST BRAZIL (SANTA ELINA, MT)
Thais Rabito PANSANI

thais-pansani@hotmail.com
Mário André Trindade DANTAS
matdantas@yahoo.com
Lidiane de Asevedo SILVA
lidi.asevedo@gmail.com
Alexander CHERKINSKY
acherkin@uga.edu
Agueda Vilhena VIALOU
avialou@gmail.com
Mírian Liza Alves Forancelli PACHECO
forancelli@gmail.com
The use of stable isotopes is a successful tool applied to the paleodiet and
paleoenvironment reconstitution of extinct animals, such as the Pleistocene
megafauna. In the context of few isotopic data available for the giant ground sloth
Glossotherium in South America, here we provide radiocarbon dating, isotopic data
(δ13C, δ18O), and paleoecological interpretation for two individuals from the Santa
Elina archaeological site, Mato Grosso state, Central Brazil. The analyses were
performed at the Center for Applied Isotope Studies, University of Georgia, USA. The
radiocarbon dating reveals that the specimen from unit III is dated to ~22,500 Cal yr
BP, and its carbon isotopic results showed a generalist diet consisting of 70% of C4
plants and 30% of C3 plants (δ13C = -3.3‰), with a broader niche breadth (BA =
0.79). The radiocarbon dating of the specimen from unit II reveals the age of ~15,000
Cal yr BP, and the carbon isotopic results also showed a generalist diet, consisting of
80% of C4 plants and 20% of C3 plants, with a slightly narrow niche breadth (BA =
0.50). The carbon results suggest a generalist diet with preference for C 4 plants for
both individuals, but their oxygen data differ slightly, with the specimen from unit III
exhibiting a lower δ18O value (-4.5‰) than the one from unit II (δ18O = -2.8‰). The
feeding behavior of Glossotherium phoenesis remained similar during these two
distinct phases, and the predominance of C4 resources in their diet allows us to
suggest that they lived in an open habitat during the late Pleistocene (~15,000–
22,500 Cal yr BP) in Central Brazil. The difference in their oxygen isotopic data may
suggest variations in the composition of water sources, or a variation in humidity
through time, with climatic conditions becoming drier through the late Pleistocene.
Our data contribute to the study of paleoecological and paleoenvironmental aspects
of Pleistocene ground sloths of South America. [CNPq: 141078/2019-7; CAPES:
88887.569989/2020-00; CAPES: 88882.443670/2019-01]
53
THE SOUTH AMERICAN FOSSIL RECORDS OF LISSAMPHIBIANS: A TAXONOMIC

PERSPECTIVE
Rodolfo SANTOS
rodolfosantos013@gmail.com
Lucas BARCELOS
lucasabarcelos@gmail.com
Lissamphibians form a highly diversified clade of non-amniote tetrapods, which

includes three extant subgroups: Anura, Urodela, and Gymnophiona. Nowadays,
these animals exhibit an almost cosmopolitan distribution, being present in all
continents except Antarctica. In this context, South America stands out because
approximately 3000 species (about 35% of all modern lissamphibians) inhabit the
continent. Some key geological events in the history of South America (e.g., the
Gondwanan breakup, its isolation during parts of Mesozoic and Cenozoic, the
Andean uplift, and the formation of the Panamá isthmus) have been pointed out
among the reasons to explain this huge biodiversity. In the present contribution, we
provide the most comprehensive bibliographic review on the South American fossil
lissamphibians to date, in which published (e.g., scientific papers, book chapters)
and unpublished (e.g., theses and dissertations) studies were included. Compared to
previous similar reviews, around 85% of the fossil records here compiled correspond
to materials revisited still not addressed by previous reviews. The South American
fossil record of Lissamphibia ranges from the Early Jurassic to the Quaternary and
includes materials from 164 fossil-bearing localities spread over eight of the twelve
South American countries. We compiled a total of 273 records, most of them frogs
(97.5%), followed by caecilians (~1.5%) and salamanders (~1%). The record of Anura
is heavily biased towards Neobatrachia (~62.8%), specially Ceratophryidae (~18.5%),
Calyptocephalellidae (13.3%), Bufonidae (~12.6%), and Leptodactylidae (~6.3%), but
pipoids and stem-anurans are also well represented (~18.2%). The occurrences of
caecilians are limited to isolated and damaged vertebrae of uncertain affinities,
whereas the salamander record is limited to the poorly known Noterpeton
bolivianum. According to our results, some fossil occurrences fall outside the
distribution area of the modern taxa, suggesting differences in their geographic range
over time. Additionally, we found that the fossil occurrences are unequally
distributed within the geological time, and consequently, some groups exhibit a
scattered temporal range (e.g., pipoids), whereas other taxa have occurrences
concentrated within a more restricted but continuous interval (e.g., ceratophryids).
Our review shows that most of the records comprise specimens still undescribed or
assigned only to more inclusive taxa, which suggests an underestimated diversity.
54
POTENTIAL GEOGRAPHIC DISTRIBUTION OF Notiomastodon platensis

(Ameghino, 1888) IN THE LATE QUATERNARY OF SOUTH AMERICA
Tawane Carvalho Fonte Boa MACHADO

tawanecfbmachado@gmail.com
Douglas RIFF
driff2@gmail.com
The proboscid Notiomastodon platensis (Ameghino, 1888) expressively integrated the

extinct South American megafauna. The numerous and well-distributed records
indicate that the species was present in almost all countries of the subcontinent
between at least 530 ka (Middle Pleistocene) and 6 ka (Early Holocene). Here, we
estimated the potential geographic distribution of N. platensis for the periods of 120
ka (Last Interglacial - LI), 21 ka (Last Glacial Maximum - LGM) and 6 ka (mid-
Holocene). Paleospecies Distribution Models (PSDMs) were created using 29 locations
in South America where fossils of the species were found and dated to approximately
21 ka. Each model was replicated 10 times (Bootstrap) and validated with 25% of the
data. Climate reconstructions for 120 ka, 21 ka e 6 ka were acquired from the images
database of the Worldclim project (v. 1.4). The climatic variables used for analysis
were: annual mean temperature, mean diurnal range, maximum temperature in the
warmest month, minimum temperature in the coldest month, annual precipitation,
precipitation of wettest month, and precipitation of driest month. The variables were
selected as suggested by the literature and, therefore, a collinearity test was not
performed. The predictive maps were produced using the MaxEnt (Machine-Learning
Maximum Entropy Model) algorithm and interpreted based on the minimum training
presence threshold. The model for the 120 ka period (area under the curve -AUC- of
0.91), characterized by higher temperatures and humidity, indicates that N. platensis
was restricted to small territories in the northeast, southeast, southwest and
northwest of the continent (Andean Region of Venezuela to Northern Peru, Brazilian
Intertropical Region, Southern Pampas and Chile). The model for 21 ka (AUC 0.95),
in turn, indicates a significant expansion of the species, occupying more widely such
regions (including Southern Peru, Bolivia, Paraguay and Northern Amazon Basin) in
a pattern similar to that proposed by other studies. In the 6 ka period (AUC 0.95),
the area with potential use by the species would be only a little smaller and more
restricted than that of the LGM, corroborating the hypothesis that climate change
was not the only responsible for the extinction of the group [FAPEMIG12630;
PQ/CNPq311003/2019-2].
55
FEEDING ECOLOGY OF LATE MIOCENE GROUND SLOTHS FROM

SOUTHWESTERN AMAZON (SOLIMÕES FORMATION, ACRE BASIN) THROUGH
TOOTH MICROWEAR ANALYSIS
Lidiane de Asevedo SILVA

Gina M. SEMPREBON
gsempreb@baypath.edu
Alceu RANZI
alceuranzi@hotmail.com
Cenozoic sediments are present in the western Amazon and contain a great diversity
of fossil vertebrates. One of the most diverse faunas from late Miocene is found in the
Solimões Formation, Acre basin, southwestern Brazilian Amazon. Regarding
mammals, fossils of Xenarthra are common, although little is known about their
paleoecology. Here, we aim to reconstruct the paleodiet of two ground sloth taxa,
Octodontobradys puruensis (Orophodontidae) and Urumacotherium campbelli
(Mylodontidae) from Talismã and Niterói sites, Amazonas and Acre states, using
tooth microwear analysis. We analyzed 22 molariforms (M1-5/m1-4), isolated and
associated with the mandible/maxilla from 11 individual specimens. One specimen
of the Quaternary Eremotherium laurillardi (Megatheriidae) from the Chandless River
(Acre state) with previous isotopic data was also examined for comparisons. Two
different regions on the mesial surface of the orthodentine were examined with a
stereomicroscope (35x) in a standard counting square of 0.16 mm². The microwear
features analysed include mean number of scratches and pits, presence or absent of
hypercoarse scratches, gouges, > four cross-scratches, > four large pits and
predominant scratch texture (fine, coarse and mixture). The compiled data was
compared with previous stereoscopic microwear of living and extinct species of
Xenarthra, using variance tests, principal component analysis (PCA) and hierarchical
cluster analyses. The fossil sloths show mean numbers of scratches (>10) compatible
to the extant sloths, Bradypus and Choloepus, and differ from armadillos with
carnivore-omnivore and insectivore diets. O. puruensis has a non-polished tooth
surface with a great proportion of coarse features and mixed scratch textures, similar
to E. laurillardi, whose isotopic data pointed to a browsing diet in a forested habitat.
Ingestion of seeds, branches, grasses and also exogenous grit could explain the high
proportion of coarse features in their tooth surfaces. In turn, with relatively polished
tooth surfaces, U. campbelli, records a higher number of scratches (~15) that are
exclusively fine or mixed, and the fewer coarse features support a predominant
folivorous diet. The Late Miocene heterogenous wetland habitats of southwestern
Amazonia could have facilitated the niche partitioning between the ground sloth
species. [CAPES 88882.443670/2019-01]
56
UMA VISITA VIRTUAL GUIADA À EXPOSIÇÃO PALEONTOLÓGICA DE ANIMAL

CROSSING NEW HORIZONS
Beatriz HÖRMANSEDER
b.marinho.h@gmail.com
Virgínia Maria do Rêgo Codá dos SANTOS
virginiacoda@gmail.com
Juliana Sampaio da COSTA
sampaiojcosta@gmail.com
Matheus Lewi Cruz Bonaccorsi de CAMPOS
matheusdrlewi@outlook.com
Rodrigo Lima VELOSO
rodrigobiolima@gmail.com
Ana Carolina de CARVALHO
ana.carvalho@cnpem.br
Devido à crise sanitária mundial, as visitas guiadas tiveram que ser adaptadas e
transferidas para o virtual. Muitos museus e instituições de pesquisa investiram
mais nessa prática e na digitalização de seu acervo para ampliar seu alcance. O
aumento da informação digital durante a pandemia da COVID-19 não impactou
apenas cientistas: a procura e o consumo de mídias digitais afetou todos, fenômeno
evidente com o lançamento de Animal Crossing New Horizons em março de 2020 pela
Nintendo. Um jogo eletrônico de simulação social que se passa em tempo real. Foi
um grande sucesso, muito devido a seu lançamento concomitante ao confinamento e
crescente busca por interações sociais. No jogo, é possível coletar espécies de peixes,
artrópodes, obras de arte e fósseis que podem ser doados ao museu. Através desse
ambiente didático, elaboramos um material audiovisual de divulgação científica para
o público gamer, composto majoritariamente de amantes de videogames lusófonos de
idade variada e de todos os gêneros. Embora não disponhamos de dados sobre o
perfil socioeconômico desta população, o material foi disponibilizado abertamente no
YouTube e divulgado através de redes sociais, de modo a tentar alcançar a maior
parcela da população. A partir da gravação e narração de uma visita ao museu
virtual, conseguimos acrescentar ao universo digital um conteúdo científico de
qualidade. Observamos uma preocupação do jogo em apresentar um conteúdo
cientificamente acurado, além de exibir uma disposição instrutiva de sua exposição.
Os fósseis estão organizados em três salas, sendo interpretadas pelas três Eras
geológicas do Fanerozóico. Interligando as salas visualizamos no chão uma árvore
filogenética que se origina na entrada da exposição e se ramifica até os espécimes
dispostos. É possível perceber de forma muito didática a proximidade taxonômica
dos fósseis através desta simples ferramenta visual. Os fósseis, possuem uma
incrível riqueza em sua anatomia, assim como sua descrição, visto que cada fóssil
possui uma explicação que pode ser acessada através da interação com o
personagem responsável pela curadoria do museu. O material pode ser acessado em
nosso canal “DLC Ciência”, disponibilizado com foco em divulgação científica e como
material de apoio para o ensino de ciências.
57
MÉTODOS VOLUMÉTRICOS NA ESTIMATIVA DE MASSA DE Massetognathus sp.

(MESOTRIÁSSICO, SUPERSEQUENCIA SANTA MARIA)
Pedro Daniel SAMPAIO

pedrods@hotmail.com
Daniel de Simão OLIVEIRA
doliveira.simao@gmail.com
Débora MORO
deboramorod@gmail.com
Flávio PRETTO
Massetognathus é um cinodonte sul-americano do Triássico Médio - início do

Triássico Superior - amplamente amostrado. Sua massa corporal foi avaliada por
estudos recentes a partir de uma série de equações lineares que relacionam a massa
com o comprimento craniano e/ou comprimento ou circunferência estilopodial
(circunferência de fêmur + circunferência do úmero). Aqui reavaliamos a massa
corporal de espécimes de Massetognathus usando valores volumétricos calculados
pelo método de Casca Convexa Mínima (Minimum Convex Hull). Um modelo-base foi
digitalizado com um scanner de superfície ZScan 700 a partir do espécime
UNIPAMPA 0625. Este espécime foi escolhido por sua completude. Após a
digitalização, o esqueleto foi montado virtualmente em posição anatomicamente
neutra, assumindo a premissa de isometria corporal. Foi possível reescalonar o
modelo-base a partir de medidas tomadas da literatura, notavelmente o comprimento
femoral (CF) e comprimento craniano (CC). Para cada modelo escalonado, foi
calculado o volume do Convex Hull, usando o software livre Meshlab. A estes
volumes, foram aplicados valores de densidade corporal entre 863 kg/m³ e 1049
kg/m³, observadas em mamíferos atuais, de modo a estimar a massa dos modelos
volumétricos. Com base nesses cálculos obtivemos faixas aproximadas de massas
corporais para cada espécime (e espécie) avaliados. Para M. pascuali (2 espécimes)
obtivemos uma faixa de 0,642 kg (CF= 53mm) a 0,926 kg (CF= 56mm). Para M.
ochagaviae (3 espécimes) tem-se uma faixa de 2,984 kg (CC=127mm) a 8,701 kg
(CC=170mm). Por fim, para Massetognathus sp. (10 espécimes) obtém-se uma faixa
de 2,469 kg (CF=83mm) a 52,758 kg (CC=310mm). Uma das principais vantagens
dessa metodologia é que, uma vez tendo-se um modelo-base, é possível calcular
volumes e massas corporais independentemente de qual elemento ósseo seja
recuperado do espécime em questão, bastando apenas escalonar o modelo-base
adequadamente. Isto contorna o “single bone problem” enfrentado por estimativas
com base em medidas lineares, e aumenta a quantidade de espécimes passíveis de
avaliação de massa corporal.
58
- Sessão Cenozóico -
PRESENCIA DE Mammuthus columbi EN SANTA LUCIA, ESTADO DE MÉXICO,

MÉXICO: UN REPORTE PRELIMINAR
Víctor Adrián PÉREZ-CRESPO

vapc79@gmail.com
Joaquin ARROYO-CABRALES
arromatu@hotmail.com
Rubén Manzanilla LÓPEZ
rubenmlo@prodigy.net.mx
Gerardo Arturo RODRÍGUEZ-FRANCO
gerardoRF9@hotmail.com
El mamut colombino (Elephantidae, Mammuthus columbi) es una de las cuatro

especies de proboscidios que habitaron en México durante el Pleistoceno y sus restos
se han encontrado en todo el país, con excepción de la Península de Yucatán. Las
recientes excavaciones realizadas en la Base Área Militar de Santa Lucía
perteneciente a la Fuerza Área Mexicana con motivo de la construcción del
Aeropuerto Internacional General Felipe Ángeles, han puesto al descubierto, en tres
niveles estratigráficos, restos de diversas especies de mamíferos del Pleistoceno,
tanto extintas como existentes, incluyendo équidos, perezosos, camélidos,
carnívoros, vertebrados pequeños y, en especial mamuts, cuyos restos van de un solo
elemento óseo a esqueletos casi completos y que podrían representar a más de 500
individuos, lo cual pudiera indicar que Santa Lucia es la localidad con el mayor
número de individuos de mamuts hallados en México y quizás de Norteamérica. Las
descripciones morfológicas, así como las medidas osteológicas y las inferencias de
edad realizadas a 13 molares de esta especie, revelan que estas piezas eran molares
M3/m3, representando a individuos adultos con una edad estimada en promedio de
40 años elefante; muestras de estos individuos fueron tomadas con objeto de
analizar los isótopos estables y reconocer el tipo de alimentación y el ambiente en
que vivieron. Sin embargo, dado el gran número de individuos hallados en Santa
Lucía es posible que se encuentren representados individuos infantiles, juveniles,
adultos y ancianos, por lo que nuevos trabajos contribuirán a comprender la
estructura de edad de la población de mamuts hallado en Santa Lucía.
Agradecimientos: PAPIIT IN101321, Consejo de Paleontología-INAH, SEDENA,
Proyecto Prehistoria y Paleoambiente del Noreste de la Cuenca de México.
59
NUEVOS REGISTROS DE MAMÍFEROS DEL PLEISTOCENO-HOLOCENO EN

PAMPA DEL TAMARUGAL, REGIÓN DE TARAPACÁ, CHILE
Natalia VILLAVICENCIO
natalia.villavicencio@uoh.cl
Rafael LABARCA
r.labarca.e@gmail.com
Francisco CARO
fco.carolivares@gmail.com
Francisco PREVOSTI
protocyon@hotmail.com
Jaqueline CORREA-LAU
jcorreal@academicos.uta.cl
José CAPRILES
jmcapriles@gmail.com
Claudio LATORRE
clatorre@bio.puc.cl
Calogero SANTORO
calogero_santoro@yahoo.com
Numerosas evidencias paleoclimáticas muestran que las cuencas del núcleo

hiperárido del Desierto de Atacama tuvieron una mayor disponibilidad hídrica
durante la última transición glacial-interglacial. Entre estas, la cuenca de Pampa del
Tamarugal es actualmente un paisaje extremo con reducida diversidad faunística y
florística, pero que en el Pleistoceno tardío fue más diversa, con presencia de
sistemas fluviales permanentes y extensos humedales. Si bien, se han reportado
hallazgos de mamíferos pleistocenos desde finales del siglo XIX en esta cuenca, los
escasos datos de proveniencia y de excavación han limitado las inferencias
cronológicas y paleoecológicas hechas en torno a ellos. En este trabajo, presentamos
numerosos hallazgos óseos de mamíferos pleistocenos, todos descubiertos en el
marco de un estudio interdisciplinario destinado a reconstruir ecosistemas del
Pleistoceno-Holoceno en la Pampa del Tamarugal. El aspecto de los elementos óseos
evidencia periodos prolongados de exposición en superficie y un grado alto de
meteorización. Análisis taxonómicos identificaron micro y mega mamíferos
pertenecientes a cf. Vicugna sp. (Artiodactyla, Camelidae, 2 localidades), Hippidion
sp. (Perissodactyla, Equidae, 2 localidades, extinto), Equus sp. (Perissodactyla,
Equidae, 1 localidad, extinto), Megatherium sp. (Pilosa, Megatheriidae, 5 localidades,
extinto), Ctenomys sp. (Rodentia, Ctenomyidae, 5 localidades), Abrocoma sp.
(Rodentia, Abrocomidae, 1 localidad) y Canidae cf. Aenocyon dirus (Carnivora, 1
localidad, extinto). Análisis de 14C AMS en bioapatita ubican el ensamble de mega
mamíferos en el Pleistoceno tardío. Tres fechados ubican a los especímenes de
Megatherium sp. entre 16670-13650 cal AP, mientras que dos fechados sitúan a
Hippidion sp. entre 15950-13970 cal AP. Un fechado para Canidae cf. A. dirus, lo
ubica en torno a 14660 cal AP. Ctenomys sp. fue datado en torno a 9620 cal AP. La
cronología obtenida sitúa a la fauna extinta en épocas previas a las primeras
ocupaciones humanas reportadas para la región, datadas en torno a 12800 cal AP.
Estos hallazgos complementan los análisis florísticos realizados en esta cuenca
evidenciando condiciones de mayor productividad en el ecosistema. Finalmente, los
nuevos descubrimientos reportados permiten una reconstrucción más detallada de
estos paisajes hoy desaparecidos. [FONDECYT-1201786, FONDECYT-1160744,
FONDECYT POSTDOCTORADO-3170706]
60
NOVOS REGISTROS DE SQUAMATA (LEPIDOSAURIA) PARA O QUATERNÁRIO

NORDESTINO, LAJEDO DE SOLEDADE, BACIA POTIGUAR
Sílvia Oliveira LOMBA

silvialomba06@gmail.com
Hermínio Ismael ARAUJO JR
Annie S. HSIOU
anniehsiou@ffclrp.usp.br
A paleofauna cenozóica do nordeste brasileiro é relativamente bem conhecida

taxonomicamente. No entanto, o registro paleoherpetológico da região ainda é pouco
explorado, sendo descritos fósseis de espécies viventes na Bahia e no Ceará. Os
depósitos dessa região são principalmente dos tipos tanque ou cavernas e ravinas,
onde se acumulam e preservam elementos esqueléticos de diversos táxons. Um
desses sítios é o Lajedo de Soledade, no Rio Grande do Norte. O Lajedo de Soledade é
um sitio paleontológico e arqueológico localizado na Chapada do Apodi, sendo a
maior exposição de rocha carbonática da Formação Jandaíra, Bacia Potiguar, Rio
Grande do Norte. Nesta localidade, o processo de carstificação alargou falhas na
rocha formando cavernas e ravinas onde são acumulados inúmeros fragmentos
fósseis associados a depósitos do Cenozóico. Os fósseis de mamíferos pleistocênicos
são bem conhecidos e documentados na literatura, no entanto, demais grupos
taxonômicos provenientes desta localidade ainda não apresentam descrição formal.
Neste trabalho, são apresentados os primeiros fósseis de lagartos e serpentes
(Squamata) para o Quaternário potiguar, que estão profisoriamente depositados no
Laboratório de Paleontologia da Faculdade de Geologia da Universidade do Estado do
Rio de Janeiro (UERJ, Rio de Janeiro), ainda sem números de tombo. Até o momento,
os espécimes fósseis, representados por restos cranianos e pós-cranianos isolados,
foram comparados com espécies viventes da herpetofauna brasileira, onde foram
identificados representantes de Tropiduridae (Tropidurus sp.), Boidae (Epicrates sp.),
Viperidae (Crotalus durissus) e serpentes Colubroides indeterminadas. Além dos
escamados, restos pós-cranianos de anuros também foram triados, porém ainda em
fase de identificação. Essas descrições ampliam o conhecimento paleoherpetológico
do local e do Quaternário do nordeste brasileiro. [CAPES]
61
PALEODIET OF A NEW EXTINCT ALOUATTINAE (PRIMATES: PLATYRRHINI:

ATELIDAE): INFERENCES FROM TOOTH MORPHOLOGY
Andre Vieira ARAUJO

andre.vieira@ifsertao-pe.edu.br
Lais Alves SILVA
Alexandre LIPARINI
alexandreliparini@yahoo.com.br
Mario Alberto COZZUOL
mario.cozzuol@gmail.com
Mário André Trindade DANTAS
Hermínio Ismael ARAUJO JR
Despite phylogenetic relationship and similarity in body size, the Atelinae and
Alouattinae primates show disparate dietary behaviors. It is difficult to classify their
diet and identify morphological correlates, particularly when it corresponds to dietary
profiles of extinct species. Recently, several researches on dental morphology have
been proposed paleodiet reconstructions. In this context, the molar shearing quotient
(SQ) is a successful approach that has been employed in the paleodiet reconstruction
of different primate taxa. Here, we present the paleodiet insights of a new specimen
of a giant Alouattinae primate fossil from a limestone cave of Bahia State, Brazil,
using molar shearing quotient (SQ). The fossil material was recovered from
Engrunado cave, located at the municipality of Nova Redenção (12°86’30”S,
41°06’81”W), and is housed at the Laboratório de Ecologia e Geociências,
Universidade Federal da Bahia (IMS/CAT). The SQ was calculated by regressing the
total length of shearing crests on a first lower molar (m1) against a proxy for body
size. We measured the lengths of six principal shearing crests on the m1 using a
stereomicroscope. The crests measured were medial protocristid, paracristid, cristid
obliqua, postcristid, postmetacristid, and pre-entocristid. All the measurements were
summed in total first molar lower shear. The measurements must be standardized
for the body size through a linear equation. The shear crests measurements and the
Shearing Quocient (SQ) found for m1 of this Alouattine fossil were 11.25 mm and
14.30, respectively. These results demonstrate that the primate fossil from
Engrunado fall in the range of the most folivores among the extinct and extant new
world monkeys. This new data allows us to contribute to the evolutionary discussion
on species related to the extraordinary Howler monkey lineage. This new Alouattine
fossil could represent an intermediate evolutionary step between big size Atelidae
fossil lineages, such as Cartelles coimbrafilhoi with a frugivore dentition, and the
extant Alouatta, with most folivorous dentition.
62
GLIPTODONTÍDEOS JUVENIS DO PLEISTOCENO TARDIO DA BAHIA
Nancy de F. C. REGO
nancyfcr@gmail.com
Jorge F. M. de JESUS
moradejesus@gmail.com
Marcelo A. FERNANDES
mafernandes@ufscar.br
Os Cingulata são caracterizados por sua armadura corporal composta por

osteodermos articulados; sendo classificados nas famílias Protobradyidae,
Peltephilidae, Paleopeltidae, Pampatheriidae, Pachyarmatheriidae, Dasypodidae e
Chlamyphoridae. Dentro desta última, encontramos a sub-família Glyptodontinae.
Os gliptodontídeos mostram indícios de sua presença desde o final do Eoceno na
América do Sul, expandindo-se por toda América durante o Grande Intercâmbio
Americano e extinguindo-se no inicio do Holoceno. Seu registro fóssil consiste
principalmente de osteodermos desarticulados e fragmentos de crânios. Existe muita
informação sobre a filogenia, taxonomia, dieta e locomoção desses animais, mas o
registro fóssil e o conhecimento de gliptodontídeos em suas fases juvenis ainda é
incipiente. A escassez de conhecimento das fases juvenis pode induzir ao erro de
classificar um estágio ontogenético como uma espécie diferente ou mesmo uma nova
espécie. Aqui descrevemos dois espécimes juvenis de gliptodontídeos, coletados na
Gruta da Lapinha, Iramaia, BA, em expedições nos anos de 2012 e 2014. O material
(registrado como LPP-PV-005 e LPP-PV-006) está depositado no Laboratório de
Paleoecologia e Paleoicnologia da Universidade Federal de São Carlos. O espécime
juvenil LPP-PV-005 compreende fragmentos das mandíbulas direita e esquerda,
fragmento do esfenoide, hemimandíbula esquerda quase completa, 50 osteodermos
dorsais da carapaça e diáfise do fêmur direito. O espécime juvenil LPP-PV-006
compreende diáfise do fêmur direito, vértebras caudais e alguns osteodermos dorsais
da carapaça. A datação por carbono da bioapatita resultou em uma idade de
13.130±30 BP para o LPP-PV-005. Esta é a primeira descrição de ocorrência de
osteodermos associados a elementos esqueléticos de gliptodontídeos juvenis.
Características dimensionais e anatômicas do espécime LPP-PV-005 comparadas às
de um espécime adulto indicam que é muito jovem e a ausência de desgaste
observada nos dentes sugere que estivesse em fase de amamentação. O fêmur do
espécime LPP-PV-006 é maior do que o do espécime LPP-PV-005, mas ainda é muito
curto comparado a fêmures de adultos. Comparações com outros espécimes juvenis e
adultos sul-americanos reforçam a ideia de que o LPP-PV-005 e o LPP-PV-006 são do
gênero Glyptotherium. Apoio: Coordenação de Aperfeiçoamento de Pessoal de Nível
Superior - Brasil (CAPES); Programa de Pós-Graduação em Ecologia e Recursos
Naturais - PPGERN/UFSCar.
63
PHYLOGENETIC RELATIONSHIP OF MACRAUCHENIIDAE (MAMMALIA,

PANPERISSODACTYLA, LITOPTERNA) AT GENERA LEVEL
Leonardo LOBO
leoloboo@gmail.com
Javier N. GELFO
jgelfo@fcnym.unlp.edu.ar
Sergio Alex AZEVEDO
sazevedo@mn.ufrj.br
In the last decade, Macraucheniidae systematic has been the focus of several studies
to enhance the understanding of the diversity and genera relationships. This
contribution proposes a new phylogenetic hypothesis of the Macraucheniidae genera
after adding new characters and new outgroups. We included eight non-
macraucheniid taxa as outgroups, three Didolodontidae, one Protolipternidae (also
considered as a junior synonym of didolodontids), two Proterotheriidae, and two
Adianthidae; and a priori we included as macraucheniids the 16 genera that are
currently accepted to belong to this family by the other cladistics studies. The matrix
used in the analysis comprehends 73 characters, subdivided into cranial (17), upper
dentition (24), lower dentition (25), and postcranial (7). The matrix was constructed
on Mesquite and the cladistic analysis was conducted on TNT. We performed an
implicit enumeration with the assumptions of non-ordered and equal weight
characters. The cladistic analysis produced 28 most parsimonious trees of 205 steps
long (CI=0.507, and RI=0.660) and the consensus with 213 steps long (CI=0.488, and
RI=0.633). The strict consensus tree shows the following relationship between the
taxa: (Didolodus (Ernestokokenia, Ricardocifellia, Miguelsoria, (Polymorphis,
((Diadiaphorus, Tetramerorhinus) (Tricoelodus (Proadiantus, ((Pternoconius,
(Coniopternium, Cramauchenia, Theosodon)) (Paranauchenia, (Llullataruca, Cullinia,
Scalabrinitherium, Huayqueriana, (Oxyodontherium, (Promacrauchenia,
(Windhausenia, (Xenorhinotherium, (Macraucheniopsis, Macrauchenia))))))))))))). The
genus Polymorphis was recovered as the sister taxon of the clade composed by
Proterotheriidae + Adianthidae + Macraucheniidae, not as macraucheniid.
Disregarding Polymorphis, all other genera attributed to macraucheniid were
recovered as a monophyletic, as well as Proterotheriidae. Adianthidae in contrast was
paraphyletic, being the stem group of macraucheniids. There were two
macraucheniid clades, one formed by Cramaucheniinae + Theosodon, and the other
by all the Macraucheniinae with the addition of Llullataruca, being placed in this
subfamily for the first time. The late Pliocene Windhausenia, was recovered as the
sister group of “Quaternary macraucheniids”, and Xenorhinotherium as sister group
of the Macraucheniopsis + Macrauchenia. Based on the topology found, Polymorphis
must be considered as incertae sedis, the Macraucheniidae temporal range is late
Oligocene – late Pleistocene / Holocene; Llullataruca became the oldest recorded for
the Macraucheniinae; and the cladogenetic event of “Quaternary macraucheniids”
should occur before the middle Pleistocene.
64
PALEOECOLOGY OF LATE PLEISTOCENE CAMELIDS FROM THE CHUY CREEK

FOSSIL ASSEMBLAGE OF THE BRAZILIAN PAMPA
Renato Pereira LOPES

paleonto_furg@yahoo.com.br
Carolina Saldanha SCHERER
carolina.ss@ufrb.edu.br
Jamil Corrêa PEREIRA
jamilpereira2168@gmail.com
The Pleistocene fossil assemblage in continental beds exposed along the banks of
Chuy Creek, in the coastal Pampa of southern Brazil, includes several mammal
species of the extinct megafauna, with scarce camelid remains. Here is presented an
analysis of the stable isotope ratios (δ13C, δ18O) in tooth enamel of camelid molars
found along the creek, consisting of two Hemiauchenia paradoxa and three Lama
guanicoe. Four specimens were collected in colluvium, but one molar of H. paradoxa
was found in situ at a stratigraphic level within a loess unit (Cordão Formation)
apparently deposited around the last glacial maximum (LGM, 26.5-19 ka) based on
luminescence ages. The carbon isotope ratios show that the analysed individuals fed
mostly on C3 plants, although the less negative δ13C of L. guanicoe (average of -7.5‰)
indicate ingestion of a substantial (~30%) proportion of C4 plants or succulent CAM
plants. The more negative δ13C (average of -11.9‰) of H. paradoxa indicate pure C3
diet with few (<10%) C4/CAM in one specimen. The higher δ18O and calculated
evaporation sensitivity than the other mammals from the assemblage show that the
camelids obtained evaporatively 18O-enriched water from the ingested plants, thus
point to drier conditions at the time they were living. A comparison with δ13C values
of Holocene L. guanicoe from archaeological sites of Argentina indicates the
landscape inhabited by the individuals, especially the LGM H. paradoxa, was similar
to the Monte and Patagonic xerophytic steppes found in arid/semiarid areas of
Argentina. The isotopic ecology of the fossil camelids indicate that southern Brazil
experienced drier conditions and change of floras from about 30 ka onwards,
possibly driven by a shift of the arid belt of South America, which may have
contributed for the local extinction of other elements of the megafauna absent within
the Cordão Formation but present in the Santa Vitória Alloformation below. The
camelids could have survived in such dry conditions but would have disappeared as
a result of expansion of wetland environments and changes in vegetation
communities of the coastal plain driven by wetter and warmer climate around the
Pleistocene-Holocene transition. [Sepkoski Grant 2017 to R.P. Lopes]
65
ANALYSES OF ENCRUSTED BONES COLLECTED IN LIMESTONES CAVES IN

MINAS GERAIS AND BAHIA
Ingrid FERNANDES
ingrid-fernandes@hotmail.com
André Gomide VASCONCELOS
andregomide86@gmail.com
Alexandre LIPARINI
Mariangela Garcia Praça LEITE
mgpleite@gmail.com
This paper presents the results obtained from slide analyses, X-ray diffractometry
(XRD), X-ray spectroscopy (EDS) and scanning electron microscopy (SEM) of 40
bones collected from deposits in 10 limestone caves, distributed in the states of
Bahia and Minas Gerais. The material belongs to the Quaternary mammal
megafauna: Tayassuidae, Equidae, Dasypoidae, Scelidotheridae, Megatheridae,
Nothrotheridae, Felidae and Proboscidae. Firstly, the material was analyzed
macroscopically, and it was observed that carbonates and other compounds were
deposited both on the surface and inside the bones. Through slide analysis, it was
possible to differentiate and classify these minerals/materials, and infer that one of
the main fouling materials was calcite mosaic. Additionally, the fouling also occurred
in the form of prismatic calcite mosaic, microcrystalline calcite, aragonite (with
acicular fabric), authigenic argillominerals, Fe and Mn oxides/hydroxides,
autochthonous (e.g. calcite fragments) and allochthonous (e.g. quartz crystals)
sediments. Precipitation can be said to depend on the amount of water present,
temperature, reducing and oxidizing conditions, and the minerals that compose the
depositional environment. These factors control the amount of materials that
participate in the process, the durability of the bone in the fossil record, and the
intensity of the diagenetic events, which determine the variations in the thickness of
the encrusted layers. From the analyses, it was noted that the minerals/materials
involved in the fossilization process in the different caves were the same, however the
forms of precipitation in the bone, were not similar. In addition, it can be inferred
that encrusted bones are in a better state of preservation, regardless of the
encrusting materials. Therefore, the microscopic analyses of the collected samples
and the understanding of the physical and chemical processes that occurred, allowed
the real dimensioning of the quantity of participating elements and the variability of
events that collaborate with the encrustation process and, consequently, with the
preservation of the fossil.
66
MORFOMETRIA GEOMÉTRICA 2D DO MOLDE ENCEFÁLICO DE FOLIVORA E

OUTROS XENARTHRA
Roberta Veronese do AMARAL

roberta.veronese@gmail.com
Rafael DELCOURT
Luciana Barbosa de CARVALHO
lucbc@acd.ufrj.br
Sergio Alex Kugland de AZEVEDO
sazedo@mn.ufrj.br
A Magnaordem Xenarthra é representada por animais que se apresentam em três

grandes grupos, Cingulata, Folivora e Vermilingua, com representantes atuais e
fósseis. O passado destes animais apresenta grande diversidade e variedade em
hábitos, bastante diferente de alguns dos representantes atuais. Este trabalho
objetiva descrever variações na morfologia do encéfalo e explorar se as mudanças nos
hábitos de alguns destes animais encontram correspondência no endocrânio. Para
descrever de modo quantitativo a variação do endocrânio dos Xenarthra fósseis e
atuais, foram usadas técnicas de reconstrução tridimensional, através do software
Mimics 10.01. A morfometria geométrica 2D foi aplicada, baseada em marcos
anatômicos dos tipos I e II, obtidos através do software TpsDig 2.32. Foram avaliadas
separadamente as vistas dorsal, ventral e lateral das imagens geradas, centralizadas,
do endocrânio reconstruído de Xenarthra atuais e fósseis. Foram realizadas análises
de Procrustes generalizada (GPA) nas coordenadas dos marcos anatômicos, Análises
de Componentes Principais (PCA) e Análises de Variáveis Canônicas (CVA) através do
MorphoJ 1.06. Para vista dorsal PC1 demonstrou 32,8% da variação e PC2 19,9%, já
em vista lateral PC1 respondeu por 19,8% estando bem próximo de PC2, com 19,3%
da variação da forma. Finalmente em vista dorsal PC1 exibiu 34,2% da variação e
PC2 19,6%. De modo geral o PCA demonstrou componentes de variação relacionados
à forma dos bulbos olfatórios e tamanho do neocórtex mostrando muitas vezes uma
divisão relacionada a cingulados e pilosas no eixo PC1. PC2 pareceu mostrar
disposição diferente entre Folivoras arborícolas/semi e terrícolas. Nothrotherium
maquinense nos gráficos de PCAs se posiciona próximo aos táxons reconhecidamente
arbóreos/semi-arbóreos. As CVAs demonstraram tendências de separação por
ordens e por hábitos posicionando os considerados arbóreos e semi-arbóreos ao
longo dos eixos. A morfologia encefálica pareceu mostrar um padrão relacionado aos
bulbos olfatórios, largos em cingulados e em táxons fossoriais e terrícolas. A forma
média da morfológia evidenciou encéfalos e bulbos olfatórios de vermilíngues
tendendo ao alongamento ântero-posterior, Folivoras com bulbos olfatórios curtos e
encéfalos largos e cingulados com grandes bulbos olfativos. As características da
forma parecem se relacionar em ordens e hábitos. [INT/MCTI; CAPES].
67
PRIMEIRO REGISTRO DE RESTOS DE MEGAFAUNA QUATERNÁRIA PARA O

QUADRILÁTERO FERRÍFERO

Luciano VILABOIM
palaiosvilaboim@hotmail.com
Alex HUBBE
alexhubbe@yhaoo.com
Tomás CORREA
tomas.correa@vale.com
Ingrid FERNANDES
Alexandre LIPARINI
alexandreliparini@yhaoo.com.br
Até o momento, fósseis de vertebrados na região do Quadrilátero Ferrífero (QF; Minas

Gerais, Brasil) eram restritos a icnofósseis, representados por ranhuras produzidas
por Xenathra extintos. Essas estruturas estão associadas às formações ferríferas e se
localizam no topo das montanhas. Neste trabalho é reportado e descrito o primeiro
fóssil corpóreo de vertebrado para essa estrutura geológica. O material foi encontrado
na Gruta do Rodeio, localizada no Parque Estatual Serra do Ouro Branco (Miguel
Burnier, distrito de Ouro Preto, MG). A caverna, que se formou nos dolomitos
ferruginosos da Formação Gandarela, ainda recebe material externo, fato que
contribui para seus depósitos sedimentares serem constantemente retrabalhados por
águas pluviais. Neste contexto é que o osso foi descoberto. Ele se encontrava
parcialmente pisoteado, sobre o piso, e próximo a uma área erodida. Trata-se de um
úmero incompleto, cuja morfologia preservada permite atribuí-lo à preguiça-terrícola
Nothrotherium maquinense Lund, 1839. As seguintes características, que estão
claramente visíveis, suportam essa atribuição: a diáfise cilíndrica, se torna triangular
à medida que as cristas peitoral e deltoide formam o processo delto peitoral
juntamente com a crista braquiocelfálica. Mais lateralizado, esse processo é formado
principalmente pela crista peitoral, que é desenvolvida e mediolateralmente o osso
apresenta a passagem do forame entepicondilar. A diáfise se quebrou um pouco
antes da junção da crista deltoide com a crista supinadora em sua porção latero
caudal. Apesar da gruta ser visitada desde a década de 1930, e ter sido alvo de
escavações arqueológicas e de trabalhos técnicos realizados ao longo desses anos, o
material somente foi descoberto, e formalmente divulgado, quando ela foi visitada por
especialistas em paleontologia de caverna. Desta forma, o novo achado reforça a
necessidade da atuação de profissionais, comprovadamente qualificados, em
trabalhos paleontológicos no QF, principalmente direcionados para a elaboração de
diagnósticos paleontológicos que integram os relatórios de impactos ambientais.
Escavações serão feitas na cavidade, a fim de localizar outros ossos dessa preguiça e
de outros animais. O úmero está depositado na coleção de paleontologia do Museu de
Ciências Naturais da Pontifícia Universidade Católica de Minas Gerais.
68
OCORRÊNCIA DE Catonyx cuvieri (XENARTHRA; SCELIDOTHERIIDAE) EM NOVA

CAVIDADE FOSSILÍFERA, NORTE DE MINAS GERAIS, BRASIL
Alexandre LIPARINI
Matheus Simões SANTOS
matheus_simoes@id.uff.br
Nildo Brito RIBEIRO
Regiane Farias de Castro SALES
regianefcastro@gmail.com
Nicolas Misailidis STRÍKIS
strikis@gmail.com
Hamilton dos Reis SALES
hamilton.sales@ifnmg.edu.br
Luciano VILABOIM
vilaboim@puminas.br
A região do norte de Minas Gerais, no Médio São Francisco, é extremamente rica em

cavernas desenvolvidas nos calcários do grupo Bambuí. No povoado do Vale do
Gaim, município de Cônego Marinho, próximo a Januária, afloram maciços calcários
com uma alta concentração de cavernas, ainda sem registro no Cadastro Nacional de
Informações Espeleológicas (CANIE). Nesse contexto, na Caverna do Formigueiro,
foram encontrados fósseis de um úmero e uma falange, parcialmente incrustados,
soltos sobre o piso da caverna em uma área de desnível a poucos metros de sua
entrada. As peças se encontram tombadas no Laboratório de Evolução e
Paleontologia do Instituto Federal do Norte de Minas Gerais, Campus Januária, com
o acrônimo LABEP. O úmero (LABEP 0001) apresenta diáfise cilíndrica e robusta,
bordas medial e lateral retas e extremidade distal alargada e achatada no eixo
anteroposterior. Epífise distal não preservada, podendo se tratar de um indivíduo
jovem. Cabeça articular esférica com 8,5 cm de diâmetro. Crista peitoral evidente a
partir da metade distal da diáfise arqueando lateralmente, confluindo com a base da
crista deltoide. Crista deltoide lateralmente projetada, proeminente em vista cranial,
na metade distal da diáfise. Forame e barra entepicondilar ausentes, sendo esta
característica diagnóstica de Catonyx cuvieri, entre os Scelidotheriinae. LABEP 0001
tem 34 cm de comprimento, 8,5 cm e 5,5 cm nas larguras médio-lateral e
anteroposterior respectivamente, na parte média da diáfise, e circunferência de 23
cm. A falange (LABEP 0002) se trata de uma falange proximal esquerda do dedo III,
do membro posterior. Apresenta 5,5 cm de comprimento anteroposterior e em sua
base proximal, 6 cm largura médio-lateral e 6,5 cm de altura dorso-plantar. Em
comparação com elementos equivalentes de outras preguiças terrícolas de tamanhos
similares, LABEP 0002 apresenta um sulco mediano de articulação com o metatarso
III mais fundo do que o observado em Valgipes bucklandi. Além disso, apresenta uma
concavidade bem evidente no centro de sua face dorsal, características de Catonyx
cuvieri. A partir deste achado direcionaremos novas expedições à região, além de
trabalhos de educação patrimonial com a população local.
69
SURVEY OF SMALL AND MEDIUM-SIZED MAMMALS FROM THE PLEISTOCENE

OF THE ATLANTIC FOREST
Rodrigo Ventura GERMANO

rodrigogermano16@gmail.com
Taissa Rodrigues Marques da SILVA
taissa.rodrigues@gmail.com
Even though the Atlantic Forest bears striking modern diversity and endemism, there
are no studies published on the fossil mammals of this biome. Here we used the
Paleobiology Database (PBDB) to search for paleontological data on small and
medium-sized mammals from the Pleistocene of the Atlantic Forest. First, we
downloaded records from the PBDB, searching for occurrences using the keywords
Mammalia, Pleistocene, and South America. We then filtered the generated Excel
table by excluding unwanted information, such as the families with megamammals
and countries outside of the Pleistocene range of the Atlantic Forest, leaving just
Brazil, Argentina, and Paraguay. With that sorted data, pivot tables were used to
combine information on the number of occurrences by order, family, and genus. We
then selected the coordinates and collection name data and used Google Earth to
import and open these coordinates, enabling us to save a file with location points.
These data were posteriorly analyzed with QGis to produce an image with collection
points across the selected countries. Finally, we used an image edition software to
overlap the coordinates on top of a published map of the potential Atlantic Forest
extension during the Pleistocene. As results, all the coordinates in Argentina and
Paraguay were outside the coverage of the Atlantic Forest, and in Brazil thirteen
points between the northeast and south regions were within its potential area.
Rodentia was the order with more records, followed by Artiodactyla, Cingulata, and
Chiroptera. The family with the largest number of specimens reported at the PBDB is
the Dasypodidae, followed by Cricetidae and Caviidae. Currently in the Atlantic
Forest the orders with most species are Chiroptera and Rodentia. Therefore, this
survey gathers all data, from the PBDB, on mammals that possibly inhabited the
Atlantic Forest during the Pleistocene. [FAPES]
70
TAXONOMIC REVIEW OF THE GENUS Asmithwoodwardia (MAMMALIA:

PANPERISSODACTYLA LITOPTERNA) OF THE PALEOGENE OF SOUTH AMERICA
Tábata Zanesco FERREIRA

tabatazf@yahoo.com.br
Javier N. GELFO
jgelfo@fcnym.unlp.edu.ar
The genus Asmithwoodwardia includes the species A. subtrigona (Eocene, Patagonia,

Argentina) and A. scotti (Paleogene, Rio de Janeiro, Brazil). This genus was originally
assigned to the Phenacodontidae, later reassigned to the Hyopsodontidae and,
currently, to the Didolodontidae or Protolipternidae, depending on different authors’
views. One of the reasons for so many reassignments is that the holotype of A.
subtrigona, an isolated molar, has been interpreted both as an M2 and an M3 by
different authors, leading to different taxonomic interpretations. The monophyly of
the genus has also been questioned. Therefore, the present work’s objectives are: (1)
identifying the dental locus of the holotype of A. subtrigona using morphometric
methods, (2) reviewing the genus Asmithwoodwardia. We studied 31 specimens of
the genus housed at the Museu de Ciências da Terra (Rio de Janeiro, Brasil), the
Laboratorio de Investigaciones en Evolución y Biodiversidad (Esquel, Argentina), the
Museo de La Plata (La Plata, Argentina) and the Museo Argentino de Ciencias
Naturales “Bernadino Rivadavia” (Buenos Aires, Argentina). To identify the tooth’s
locus, we used linear and geometric morphometrics, and also morphological analysis.
Other M1-3 of bunodont ungulates were used for comparison. For the linear
morphometric analysis, we used the angle between the tooth’s distal face and the
alignment of the paracone and metacone; for the geometric morphometric, we used
17 landmarks. On the morphological analysis, we observed the development and
placement of the metacone. The holotypes of A. subtrigona and A. scotti were coded
and included in a phylogenetic matrix of 87 characters and 26 taxa to test their
relationships with other bunodont ungulates. Every analysis suggested a greater
similarity of the holotype of A. subtrigona with M3s, corroborating, by more objective
methods, previous interpretations of it as being an M3. The phylogenetic analysis
agreed with previous interpretations of the polyphyletic nature of this genus.
Currently, A. scotti represents a taxon present at lower latitudes, with a simpler
structure (narrower M3 lacking a hypocone), while A. subtrigona represents a more
advanced form, present at the higher latitudes of Patagonia. [CNPq; FAPERJ]
71
ESTUDO SISTEMÁTICO DO MATERIAL DE TOXODONTIDAE (MAMMALIA,

NOTOUNGULATA) DO QUATERNÁRIO DO ESTADO DA BAHIA
Carolina Saldanha SCHERER

carolinass@ufrb.edu.br
Denise Fonseca dos SANTOS
denisefonsecads@gmail.com
A família Toxodontidae, da ordem Notoungulata, é nativa da America do Sul, a qual

viveu desde o Oligoceno Inferior até o Pleistoceno, com ampla distribuição no
Pleistoceno do Nordeste do Brasil. Tendo em vista esta abundante distribuição e o
breve conhecimento e entendimento acerca desta família, o presente trabalho possui
como objetivo a identificação de espécimes da família Toxodontidae provenientes de
depósitos fossilíferos do estado da Bahia. Os materiais estudados advêm de dois
municípios, sendo o de Santaluz, onde foram encontrados e retirados de três
depósitos fossilíferos do tipo tanque denominados Lagoa Escura (11º59’56”S,
39º24’49”O), Varzea Funda (11º18’69”S, 39º42’81”O) e Vargem Grande
(11º16’55.96”s, 39º34’30.12”O); e o município de Ourolândia, cujo material é
proveniente da Toca dos Ossos (41º55’51.25” S, 41º3’27.10” O”). Todos os materiais
estudados estão tombados e armazenados na Coleção de Paleontologia de
Vertebrados do Museu de Zoologia e Paleontologia da Universidade Federal do
Recôncavo da Bahia (UFRB-PV). Devido a insuficiência de material de comparação e
similaridade entre as espécies, não foi possível a realização da identificação a nível
específico da maior parte do material. No total, o material estudado trata-se de 19
espécimes, os quais dividem-se em material dentário e pós-cranianos, sendo este
último a maioria. Para o material dentário, foram identificados 6 espécimes, sendo 1
incisivo direito (UFRB-PV412), 1 molar direito (UFRB-PV4107) e 2 pré-molares
direitos (UFRB-PV714 e UFRB-PV1112), que se assemelham à Toxodon platensis;
além de 1 molar esquerdo (UFRB-PV413) e 2 pré-molares direitos (UFRB-PV384 e
706), que foram identificados apenas como Toxodontinae. Para o pós crânio, foram
identificados como Toxodontidae: 1 fêmur esquerdo (UFRB-PV4292), 1 fragmento de
fêmur esquerdo (UFRB-PV235), 2 metatarsais (UFRB-PV4208, 4223 e UFRB-
PV4260), 3 cuneiformes (UFRB-PV4240, 3593 e 3598), 3 metacarpais (UFRB-PV705,
4266 e UFRB-PV4270), 2 fragmentos de metatarsais (UFRB-PV4215 e 3640). No
estado da Bahia, foi confirmada simpatria entre as espécies de Toxodon platensis e
Trigodonops lopesi Ourolândia e Santana. A diferença crucial entre essas espécies é
dada através da dentição. Ademais, alguns pesquisadores consideram a espécie
Trigodonops lopesi sinônimo para Piauhytherium capivarae. Porém, ainda permanece
a dificuldade de identificação do material pós-craniano devido a fragmentação e
similaridade entre os táxons.
72
NOVOS ACHADOS PALEONTOLOGÓGICOS PARA O CARSTE DE ARCOS-PAINS-

DORESOPOLIS, MINAS GERAIS
Luciano VILABOIM
palaiosvilaboim@hotmail.com
Agenor O. NATIVO
agenornativo@hotmail.com
Jader C. OLIVEIRA
jader.caetano.oliveira@gmail.com
Ingrid FERNANDES
Alexandre LIPARINI
No carste de Arcos-Pains-Doresópolis são relatadas mais de cinco mil cavernas, e de

maneira oposta aos achados arqueológicos, o registro paleontológico é limitado a
menos de uma dezena. Isso pode estar diretamente relacionado aos
empreendimentos minerários da região, que por estratégia, não autorizam a
divulgação desse tipo de achado. A Toca do Angá (Córrego Fundo, MG) ficou
nacionalmente conhecida devido a um achado paleontológico, feito por um morador
local, enquanto pescava em 1997. De lá foram retirados vários ossos de
Notiomastondon platensis Ameghino, 1906. Em janeiro de 2021, boatos de novos
ossos vinham sendo relatados para a caverna. Um desses ossos, objeto do presente
trabalho, foi entregue no Laboratório de Paleontologia do Museu de Ciências Naturais
da PUC MINAS. Trata-se de um fragmento proximal de úmero direito,
preliminarmente atribuído a uma preguiça-terrícola da Família Megalonychidae. Um
pouco mais de um terço da porção proximal foi encontrado. Possui morfologia muito
semelhante, mas tamanho diferente comparado aos Megalonychidae conhecidos do
Quaternário brasileiro. A diáfise se inicia cilíndrica e se comprime crânio-
caudalmente ao meio. A cabeça circular apresenta as tuberosidades maior e menor
semelhantes em proporção e morfologia próximo a Australonyx aquae. Separadas por
um sulco bem definido onde é notável como a borda medial da tuberosidade maior e
contígua com a crista peitoral e o início da crista deltoíde está muito próximo da
borda lateral da tuberosidade maior, assim como em A. aquae. Está em andamento
os tramites burocráticos para a coleta de outros fósseis reportados para a Toca do
Angá. Vértebras gigantes foram vistas às margens do rio subterrâneo que inunda a
caverna, porém, em uma visita à caverna, tal material não foi visto. Assim, tal
resgate é de suma importância, visto que a caverna é de fácil acesso, apresentando
uso intenso por parte da população local e turistas, o que acarretará em perdas
irreparáveis ao patrimônio aqui relatado.
73
NEW OCCURRENCE OF Notiomastodon platensis (MAMMALIA, PROBOSCIDEA) IN

AGUALINDA, NORTE DE SANTANDER, QUATERNARY OF COLOMBIA
Jaime Y. SUÁREZ-IBARRA
jysuarezibarra@gmail.com
Lidiane de ASEVEDO
Lucas de Melo FRANÇA
lucasmfranca@hotmail.com
Michael ZIEGLER
mziegler@shh.mpg.de
Saddy MOLINA-ESCALANTE
juliana13roa@gmail.com
Fossil remains of proboscidean Notiomastodon platensis and xenarthran

Eremotherium laurillardi reported in the state of Norte de Santander, Colombia, along
with the diverse geomorphological characteristics of the north Andes Mountain
ranges, make the NE portion of Colombia a significant place to study the
paleoecology of extinct megamammals. Gomphothere findings in Norte de Santander
permit taxonomic assignment to species-level, overcoming a problem that South
American Quaternary gomphotheres face: having few diagnostic elements for such a
purpose, since only skulls and tusks allow the species determination. Here, we report
a new occurrence of the South American gomphothere N. platensis in the
municipality of Agualinda, Norte de Santander, NE Colombia. Our taxonomic
classification is based on the morphological description of a recently recovered
gomphothere tusk. The specimen was found in September 2021 in Quaternary fluvial
sedimentary deposits, and posteriorly housed at the Archeological and
Paleontological Museum of Agualinda. The upper tusk (largely covered by a
carbonatic crust of varied grain sizes and fragmented into three segments due to the
extraction by locals), presents a curved tip, a robust proximal portion exhibiting a
semi-oval cross-section, a length of 186 cm, and two ratios (length vs. width) at the
posterior and anterior regions of 1.24 and 1.26, respectively. During the initial
excavation of the piece, the most posterior part was broken on a longitudinal plane,
exposing the pulp canal. The disposition and whitish color of the dentine layers
suggest a good quality preservation. The tusk lacks torsion, a feature that identifies
the species Cuvieronius hyodon. Given the diagnostic morphological features and
dimensions described above, it is possible to assign the classification of an adult
specimen of N. platensis. Our reported new occurrence is important because it
contributes to the biogeographical record of proboscideans in northern South
America. Despite Colombia is the gateway for gomphothere taxa during the Great
American Biotic Interchange, currently, there are no reported records of C. hyodon
based on diagnostic remains, making the need to improve the developing fossil record
of proboscideans in Colombia (e.g., C. hyodon, N. platensis) imperative.
74

VII Congresso Latinoamericano de Paleontologia de Vertebrados programa e resumos

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VII Congresso Latinoamericano de Paleontologia de Vertebrados programa e resumos

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